You are on page 1of 397

Mycosphere 11(1): 1060–1456 (2020) www.mycosphere.

org ISSN 2077 7019


Article
Doi 10.5943/mycosphere/11/1/8

Outline of Fungi and fungus-like taxa


Wijayawardene NN1, Hyde KD2, Al-Ani LKT3,4, Tedersoo L5, Haelewaters
D6,7,8,9, Rajeshkumar KC10,11, Zhao RL12,13, Aptroot A14, Leontyev DV15, Saxena
RK16, Tokarev YS17, Dai DQ1,*, Letcher PM 18, Stephenson SL19, Ertz D20,21,
Lumbsch HT22, Kukwa M23, Issi IV17, Madrid H24, Phillips AJL25, Selbmann
L26,27, Pfliegler WP28, Horváth E29, Bensch K30, Kirk PM31, Kolaříková K32,
Raja HA33, Radek R34, Papp V35, Dima B36, Ma J37, Malosso E38, Takamatsu
S39,40, Rambold G41, Gannibal PB42, Triebel D43, Gautam AK44, Avasthi S45,
Suetrong S46,47, Timdal E48, Fryar SC49, Delgado G50, Réblová M51, Doilom
M52,71,72,73, Dolatabadi S53, Pawłowska JZ54, Humber RA55, Kodsueb R56,
Sánchez-Castro I57, Goto BT58, Silva DKA59, de Souza FA60, Oehl F61, da Silva
GA62, Silva IR62, Błaszkowski J63, Jobim K64, Maia LC62, Barbosa FR65, Fiuza
PO66, Divakar PK67, Shenoy BD68, Castañeda-Ruiz RF69, Somrithipol S47,
Lateef AA70, Karunarathna SC71,72,73, Tibpromma S71,72,73, Mortimer PE71,72,73,
Wanasinghe DN71,72,73, Phookamsak R2,71,72,73,74, Xu J71,72,73,74, Wang Y75, Tian
F75, Alvarado P76, Li DW77, Kušan I78, Matočec N78, Mešić A78, Tkalčec Z78,
Maharachchikumbura SSN79, Papizadeh M 80, Heredia G81, Wartchow F82,
Bakhshi M83, Boehm E84, Youssef N85, Hustad VP86, Lawrey JD87, Santiago
ALCMA88, Bezerra JDP89, Souza-Motta CM89, Firmino AL90, Tian Q2,
Houbraken J91, Hongsanan S92, Tanaka K93, Dissanayake AJ79, Monteiro JS94,
Grossart HP95,96, Suija A97, Weerakoon G98, Etayo J99, Tsurykau A100,101,
Vázquez V102,103, Mungai P104, Damm U105, Li QR106, Zhang H107, Boonmee S2,
Lu YZ108,109, Becerra AG110, Kendrick B111, Brearley FQ112, Motiejūnaitė J113,
Sharma B11, Khare R11, Gaikwad S11, Wijesundara DSA114, Tang LZ1,*, He
MQ12,13, Flakus A115, Rodriguez-Flakus P116, Zhurbenko MP117, McKenzie
EHC118, Stadler M119,120, Bhat DJ121, Liu JK79, Raza M12, Jeewon R122,
Nassonova ES123, Prieto M124, Jayalal RGU125, Erdoğdu M126, Yurkov A127,
Schnittler M128, Shchepin ON129, Novozhilov YK129, Silva-Filho AGS130,
Gentekaki E2, Liu P131, Cavender JC132, Kang Y133, Mohammad S134, Zhang
LF135, Xu RF135, Li YM135, Dayarathne MC75, Ekanayaka AH2, Wen TC136,137,
Deng CY138, Pereira OL139, Navathe S140, Hawksworth DL141,142, Fan XL143,
Dissanayake LS137, Kuhnert E144, Grossart HP145, 146, Thines M147, 148
1
Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and
Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, P.R. China.
2
Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand
3
Department of Plant Protection, College of Agriculture, University of Baghdad, Baghdad, Iraq
Submitted 5 December 2019, Accepted 17 March 2020, Published 18 March 2020
Corresponding Authors: Dong-Qin Dai – e-mail – cicidaidongqin@gmail.com,
Kevin D. Hyde – e-mail – kdhyde3@gmail.com,
Li-Zhou Tang – e-mail – tanglizhou@163.com 1060
4
School of Biology Science, Universiti Sains Malaysia, Minden Malaysia
5
Natural History Museum, University of Tartu, 14a Ravila, 50411 Tartu, Estonia
6
Faculty of Science, University of South Bohemia, Branišovská 31, 370 05 České Budějovice, Czech Republic
7
Department of Botany and Plant Pathology, Purdue University, 915 W. State Street, West Lafayette, Indiana 47907,
USA
8
Herbario UCH, Universidad Autónoma de Chiriquí, Apartado Postal 0427, David, Panama
9
Department of Biology, Research Group Mycology, Ghent University, K.L. Ledeganckstraat 35, 9000 Ghent,
Belgium
10
National Fungal Culture Collection of India (NFCCI), Biodiversity and Palaeobiology (Fungi) Group, Agharkar
Research Institute, Pune, Maharashtra 411 004, India
11
Biodiversity and Palaeobiology (Lichens) Group, Agharkar Research Institute, Pune, Maharashtra 411004, India
12
State Key Laboratory of Mycology, Institute of Microbiology Chinese Academy of Sciences, Chaoyang District,
Beijing 100101, P.R. China
13
College of Life Sciences, University of Chinese Academy of Sciences, Huairou District, Beijing 100408, P.R. China
14
Laboratório de Botânica / Liquenologia, Instituto de Biociências, Universidade Federal de Mato Grosso do Sul,
Avenida Costa e Silva s/n, Bairro Universitário, CEP 79070-900, Campo Grande, Mato Grosso do Sul, Brazil
15
Department of Botany, H.S. Skovoroda Kharkiv National Pedagogical University, Valentynivs'ka 2, Kharkiv 61168
Ukraine
16
Birbal Sahni Institute of Palaeosciences, 53 University Road, Lucknow-226007, India
17
Laboratory of Microbiological Control, All-Russian Institute of Plant Protection, Shosse Podbelskogo 3, Pushkin,
St. Petersburg, 196608, Russia
18
Department of Biological Sciences, The University of Alabama, Tuscaloosa, Alabama 35487, USA
19
Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas 72701, USA
20
Botanic Garden Meise, Department of Research, Nieuwelaan 38, BE-1860 Meise, Belgium
21
Fédération Wallonie-Bruxelles, Direction générale de l’Enseignement non obligatoire et de la Recherche
scientifique, Rue A. Lavallée 1, BE-1080 Bruxelles, Belgium
22
Science & Education, The Field Museum, 1400 S. Lake Shore Drive, Chicago, IL 60605, USA
23
Department of Plant Taxonomy and Nature Conservation, Faculty of Biology, University of Gdańsk, Wita Stwosza
59, PL-80-308 Gdańsk, Poland
24
Centro de Genómica y Bioinformática, Facultad de Ciencias, Universidad Mayor, Camino La Pirámide 5750,
Huechuraba, Santiago, Chile
25
Universidade de Lisboa, Faculdade de Ciências, Biosystems and Integrative Sciences Institute (BioISI), Campo
Grande, 1749-016 Lisbon, Portugal
26
Department of Ecological and Biological Sciences (DEB) University of Tuscia, Viterbo, Italy
27
Italian Antarctic National Museum (MNA), Mycological Section, Genoa, Italy
28
Department of Molecular Biotechnology and Microbiology, University of Debrecen, Debrecen, Egyetem tér 1.,
4032, Hungary
29
Department of Genetics and Applied Microbiology, University of Debrecen, Debrecen, Egyetem tér 1., 4032
Hungary
30
SNSB IT-Center, Botanische Staatssammlung München, Menzinger Straße 67, 80638 München, Germany /
Westerdijk Fungal Biodiversity Institute, Uppsalalaan 8, 3534 CT Utrecht, The Netherlands
31
Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3DS, UK
32
Institute of Botany, Czech Academy of Sciences Průhonice 252 43, Czech Republic
33
Department of Chemistry and Biochemistry, University of North Carolina at Greensboro, Greensboro, NC 27402,
USA
34
Evolutionary Biology, Institute for Biology/Zoology, Freie Universität Berlin, 14195 Berlin, Germany
35
Department of Botany, Szent István University, Ménesi st. 44, 1118 Budapest, Hungary
36
Institute of Biology, Eötvös Loránd University, Pázmány Péter sétány 1/c, 1117 Budapest, Hungary
37
College of Agronomy, Jiangxi Agricultural University, Nanchang, 330045, Jiangxi, P.R. China
38
Laboratório de Hifomicetos de Folhedo, Departamento de Micologia, Centro de Biociências, Universidade Federal
de Pernambuco, 50740-600, Brazil
39
Graduate School of Bioresources, Mie University, 1577 Kurima-Machiya, Tsu, Mie 514-8507, Japan
40
University of Southern Queensland, Centre of Crop Protection, Toowoomba, Queensland, 4350, Australia
41
Department of Mykologie, Universität of Bayreuth, Universitätsstraße 30, 95440 Bayreuth, Germany
42
All-Russian Institute of Plant Protection, Saint Petersburg, Russia
43
Staatliche Naturwissenschaftliche Sammlungen Bayerns, IT Center, Menzinger Straße 67, 80638 München,
Germany
44
School of Agriculture, Faculty of Agriculture, Abhilashi University, Mandi, Himachal Pradesh, India
45
School of Studies in Botany, Jiwaji University, Gwalior, Madhya Pradesh, India
46
National Biobank of Thailand (NBT), National Science and Technology Development Agency (NSTDA), 113
Thailand Science Park, Thanon Phahonyothin, Tambon Khlong Nueng, Amphoe Khlong Luang, Pathum Thani 12120,
Thailand

1061
47
BIOTEC, National Science and Technology Development Agency (NSTDA), 113 Thailand Science Park, Thanon
Phahonyothin, Tambon Khlong Nueng, Amphoe Khlong Luang, Pathum Thani 12120, Thailand
48
Natural History Museum, University of Oslo, Blindern, P.O. Box 1172, 0318 Oslo, Norway
49
College of Science and Engineering, Flinders University, G.P.O. Box 2100, Adelaide 5001 SA, Australia
50
EMLab P&K Houston, 10900 Brittmoore Park Drive Suite G Houston, Texas 77041, USA
51
Institute of Botany, Academy of Sciences, Průhonice 252 43, Czech Republic
52
Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand
53
Faculty of Engineering, Sabzevar University of New Technology, Sabzevar, Iran
54
Department of Molecular Phylogenetics and Evolution, Faculty of Biology, Biological and Chemical Research
Center, University of Warsaw, ul. Żwirki i Wigury 101, 02-089 Warsaw, Poland
55
USDA-ARS Emerging Pests and Pathogens Research, Robert W. Holley Center for Agriculture and Health, Tower
Road, Ithaca, NY 14853-2901, USA
56
Microbiology Program, Faculty of Science and Technology, Pibulsongkram Rajabhat University, Phitsanulok
65000, Thailand
57
Departamento de Microbiología, Campus de Fuentenueva, Universidad de Granada, 18071, Granada, Spain
58
Departamento de Botânica e Zoologia, Universidade Federal do Rio Grande do Norte, Campus Universitário,
59072–970, Natal, RN, Brazil
59
Universidade Federal da Paraíba, Programa de Pós-Graduação em Ecologia e Monitoramento Ambiental, Centro
de Ciências Aplicadas e Educação, Campus IV, Litoral Norte, Rio Tinto, PB 58297-000, Brazil
60
Embrapa Milho & Sorgo, BR-35702098 Sete Lagoas, MG, Brazil
61
Agroscope, Competence Div Plants & Plant Prod, Ecotoxicol, Schloss 1, CH-8820 Wadenswil, Switzerland
62
Universidade Federal de Pernambuco, Programa de Pós-Graduação em Biologia de Fungos, Av. da Engenharia,
s/n, Cidade Universitária, CEP 50740-600, Recife, PE, Brazil
63
Department of Ecology, Protection and Shaping of Environment, West Pomeranian University of Technology,
Słowackiego 17, PL–71434 Szczecin, Poland
64
Departamento de Botânica e Zoologia, Universidade Federal do Rio Grande do Norte, Campus Universitário,
59072–970, Natal, RN, Brazil
65
Acervo Biológico da Amazonia Meridional, Universidade Federal de Mato Grosso, Sinop 78557267, Brazil
66
Universidade Federal do Rio Grande do Norte (UFRN), Programa de Pós-graduação em Sistemática e Evolução,
Centro de Biociências, Campus Universitário, Av. Senador Salgado Filho, 3000, Lagoa Nova, Natal-RN, 59078-970,
Brazil
67
Departamento de Farmacología, Farmacognosia y Botánica, Facultad de Farmacia, Universidad Complutense de
Madrid, Madrid 28040, Spain
68
CSIR-National Institute of Oceanography Regional Centre, 176, Lawson’s Bay Colony, Visakhapatnam – 530017,
Andhra Pradesh, India
69
Instituto de Investigaciones Fundamentales en AgriculturaTropical, ‘Alejandro de Humboldt’ (INIFAT), Santiago
deLas Vegas, Habana, Cuba
70
Department of Plant Biology, Faculty of Life Sciences, University of Ilorin, Nigeria
71
Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy
of Science, Kunming 650201, Yunnan, P.R. China
72
World Agroforestry Centre, East and Central Asia, Kunming 650201, Yunnan, P.R. China
73
Centre for Mountain Futures (CMF), Kunming Institute of Botany, Kunming, Yunnan, 650201, P.R. China
74
Institute of Animal Science, State Key Laboratory of Animal Nutrition, Chinese Academy of Agricultural Sciences,
Beijing, 100193, P.R. China
75
Department of Plant Pathology, Agriculture College, Guizhou University, Guiyang 550025, P.R. China
76
ALVALAB, Dr. Fernando Bongera st., Severo Ochoa Bldg. S1.04, Oviedo, 33006, Spain
77
The Connecticut Agricultural Experiment Station, Valley Laboratory, 153 Cook Hill Road, Windsor, CT 06095,
USA
78
Ruđer Bošković Institute, Bijenička 54, HR-10000 Zagreb, Croatia
79
School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu
611731, P.R. China
80
Department of Microbiology, Pasteur Institute of Iran, Tehran, Iran
81
Laboratory of Microfungi, Instituto de Ecologı ´a A. C., Xalapa, Ver., Mexico
82
Universidade Federal da Paraíba, Departamento de Sistemática e Ecologia, CEP 58051-970, João Pessoa, PB,
Brazil
83
Department of Botany, Iranian Research Institute of Plant Protection, P.O. Box 19395-1454, Agricultural
Research, Education and Extension Organization (AREEO), Tehran, Iran
84
42 Longacre Drive Livingston, NJ, 07039, USA
85
Oklahoma State University, Department of Microbiology and Molecular Genetics, Stillwater OK, USA
86
Department of Natural Sciences, Northwest Missouri State University, Maryville, Missouri, 64468, USA
87
Department of Environmental Science and Policy, George Mason University, 4400 University Drive, Fairfax, VA
22030-4444, USA

1062
88
Post Graduate Program in Biology of Fungi, Department of Mycology, Federal University of Pernambuco, Av.
Nelson Chaves, s/ n, 50670-420 Recife, PE, Brazil
89
Departamento de Micologia Prof. Chaves Batista, Universidade Federal de Pernambuco, Av. Prof. Moraes Rego,
s/n, Centro de Biociências, Cidade Universitária, CEP: 50670-901, Recife, PE, Brazil
90
Instituto de Ciências Agrárias, Universidade Federal de Uberlândia, Monte Carmelo 38500-000, Brazil
91
Westerdijk Fungal Biodiversity Institute, Uppsalalaan 8, 3584 CT Utrecht, the Netherlands
92
Shenzhen Key Laboratory of Laser Engineering, College of Optoelectronic Engineering, Shenzhen University,
Shenzhen, P.R. China
93
Faculty of Agriculture and Life Science, Hirosaki University, 3 Bunkyo-cho, Hirosaki, Aomori 036-8561, Japan
94
Coordination of Botany, Museu Paraense Emílio Goeldi, 66077830, Belém, PA, Brazil
95
Leibnitz Institute of Freshwater Ecology and Inland Fisheries (IGB), Department Experimental Limnology, Alte
Fischerhuette 2, D-16775 Stechlin, Germany
96
Institute for Biochemistry and Biology, Potsdam University, Maulbeerallee 2, 14469 Potsdam, Germany
97
Institute of Ecology and Earth Sciences, University of Tartu, 40 Lai st., EE-51005 Tartu, Estonia
98
Algae, Fungi and Plants Division, Department of Life Sciences, The Natural History Museum, Cromwell Road,
London, SW7 5BD, UK
99
Department of Biology, IES Zizur, Ronda S. Cristo´bal 196, 31180 Zizur Mayor, Navarra, Spain
100
F. Skorina Gomel State University, Department of Biology, Sovetskaja Str. 104, Gomel 246019, Belarus
101
Samara National Research University, Institute of Natural Sciences, Department of Ecology, Botany and Nature
Protection, Moskovskoye shosse 34, Samara 443086, Russia
102
Department of Ecology, Faculty of Sciences, University of Málaga, Boulevard Louis Pasteur s/n, 29071-Málaga,
Spain
103
Department of Research and Development, Coccosphere Environmental Analysis, Málaga, Spain
104
Ecological Monitoring Unit, Biodiversity Research and Monitoring Division, Kenya Wildlife Service, P.O. Box
40241, Nairobi 00100, Kenya
105
Senckenberg Museum of Natural History Görlitz, PF 300 154, 02806 Görlitz, Germany
106
Department of Pharmacology of Materia Medica, School of Pharmaceutical Sciences, Guizhou Medical
University, University Town, Guian New District, Guizhou 550025, P.R. China
107
Yunnan Institute of Food Safety, Kunming University of Science and Technology, Kunming 650500, P.R. China
108
Engineering and Research Center for Southwest Bio-Pharmaceutical Resources of National Education Ministry of
China, Guizhou University, Guiyang 550025, Guizhou, P.R. China
109
School of Pharmaceutical Engineering, Guizhou Institute of Technology, Guiyang, Guizhou, 550003, P.R. China
110
Laboratorio de Micología, Instituto Multidisciplinario de Biología Vegetal (IMBIV), CONICET, Universidad
Nacional de Córdoba, Casilla de Correo 495, 5000 Córdoba, Argentina.
111
Mycologue, 8727 Lochside Drive, Sidney, BC V8L 1M8, Canada
112
Department of Natural Sciences, Manchester Metropolitan University, Chester Street, Manchester, M1 5GD, UK.
113
Laboratory of Mycology, Nature Research Centre, Žaliųjų ežerų Str. 49, 08406 Vilnius, Lithuania
114
National Institute of Fundamental Studies, Hantane Road, Kandy, Sri Lanka
115
Department of Lichenology, W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz 46, PL–31–512
Krakow, Poland
116
Laboratory of Molecular Analyses, W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz 46, PL–31–
512 Krakow, Poland
117
Laboratory of the Systematics and Geography of Fungi, Komarov Botanical Institute, Russian Academy of
Sciences, Professor Popov 2, St.-Petersburg, 197376, Russia
118
Manaaki Whenua-Landcare Research, Private Bag No 92170, Auckland, New Zealand
119
Department of Microbial Drugs, Helmholtz-Zentrum für Infektionsforschung GmbH, Inhoffenstrasse 7, 38124
Braunschweig, Germany
120
German Centre for Infection Research (DZIF), partner site Hannover-Braunschweig, 38124 Braunschweig,
Germany
121
No. 128/1-J, Azad Co-Op Housing Society, Curca, P.O. Goa Velha-403108, India
122
Dept Health Sciences, Faculty of Science, University of Mauritius, Reduit, Mauritius
123
Laboratory of Cytology of Unicellular Organisms, Institute of Cytology of the Russian Academy of Sciences,
Tikhoretsky ave. 4, St. Petersburg, 194064, Russia
124
Departamento de Biología y Geología, Física y Química Inorgánica. Universidad Rey Juan Carlos. C/Tulipan sn,
28933, Móstoles, Madrid, Spain
125
Department of Natural Resources, Sabaragamuwa University of Sri Lanka, Belihuloya, 70140, Sri Lanka
126
Department of Landscape Architects, Faculty of Agriculture, Kırşehir Ahi Evran University, Kırşehir, Turkey
127
Leibniz Institute DSMZ-German Collection of Microorganisms and Cell Cultures GmbH, Inhoffenstrasse 7 B,
38124 Braunschweig, Germany
128
Institute of Botany and Landscape Ecology, Ernst Moritz Arndt University Greifswald, Soldmannstr. 15,
Greifswald 17487, Germany
129
The Komarov Botanical Institute of the Russian Academy of Sciences, Laboratory of Systematics and Geography
of Fungi, Prof. Popov Street 2, 197376 St. Petersburg, Russia

1063
130
Universidade Federal do Rio Grande do Norte, Programa de Pós Graduação em Sistemática e Evolução, CEP
59064-741, Natal, RN, Brazil
131
Engineering Research Center of Chinese Ministry of Education for Edible and Medicinal Fungi, Jilin Agricultural
University, Changchun, 130118, P.R. China
132
Department of Environmental and Plant Biology, Ohio University, Athens, Ohio 45701
133
Key Laboratory of Environmental Pollution Monitoring and Disease Control, Ministry of Education of Guizhou &
Guizhou Talent Base for Microbiology and Human Health, Key Laboratory of Medical Microbiology and
Parasitology of Education Department of Guizhou, School of Basic Medical Sciences, Guizhou Medical University,
Guiyang, P.R. China
134
Department of Biotechnology, Iranian Research Organization for Science and Technology (IROST), Tehran, Iran
135
College of Biological Resource and Food Engineering, Qujing Normal University, Qujing, Yunnan 655011, P.R.
China
136
State Key Laboratory Breeding Base of Green Pesticide and Agricultural Bioengineering, Key Laboratory of
Green Pesticide and Agricultural Bioengineering, Ministry of Education, Guizhou University, Guiyang, 550025, P.R.
China
137
The Engineering Research Center of Southwest Bio–Pharmaceutical Resource, Ministry of Education, Guizhou
University, Guiyang, 550025, P.R. China
138
Guizhou institute of biology, Guizhou academy of science, Guiyang, 550009, P.R. China
139
Universidade Federal de Viçosa, Departamento de Fitopatologia, Viçosa – Minas Gerais, Brazil
140
Agharkar Research Institute, G. G. Agarkar Road, Pune 411004, India
141
Department of Life Sciences, The Natural History Museum, London SW7 5BD, United Kingdom, and Comparative
Plant and Fungal Biology, Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3DS, UK
142
Jilin Agricultural University, Changchun, Jilin Province, 130118, P.R. China
143
The Key Laboratory for Silviculture and Conservation of Ministry of Education, Beijing Forestry University,
Beijing 100083, P.R. China
144
Leibniz University Hannover, Institute for Organic Chemistry and MWZ, Hannover 30167, Germany
145
Leibnitz Institute of Freshwater Ecology and Inland Fisheries (IGB), Dept. Experimental Limnology, Alte
Fischerhuette 2, D-16775 Stechlin, Germany
146
Institute for Biochemistry and Biology, Potsdam University, Maulbeerallee 2, 14469 Potsdam, Germany
147
Goethe University, Department for Biological Sciences, Institute of Ecology, Evolution and Diversity, Max-von-
Laue-Str. 13, D-60486 Frankfurt am Main, Germany
148
Senckenberg Biodiversity and Climate Research Centre, Senckenberganlage 25, D-60325 Frankfurt am Main,
Germany

Wijayawardene NN, Hyde KD, Al-Ani LKT, Tedersoo L, Haelewaters D, Rajeshkumar KC, Zhao
RL, Aptroot A, Leontyev DV, Saxena RK, Tokarev YS, Dai DQ, Letcher PM, Stephenson SL,
Ertz D, Lumbsch HT, Kukwa M, Issi IV, Madrid H, Phillips AJL, Selbmann L, Pfliegler WP,
Horváth E, Bensch K, Kirk PM, Kolaříková K, Raja HA, Radek R, Papp V, Dima B, Ma J,
Malosso E, Takamatsu S, Rambold G, Gannibal PB, Triebel D, Gautam AK, Avasthi S, Suetrong
S, Timdal E, Fryar SC, Delgado G, Réblová M, Doilom M, Dolatabadi S, Pawłowska J, Humber
RA, Kodsueb R, Sánchez-Castro I, Goto BT, Silva DKA, de Souza FA, Oehl F, da Silva GA,
Silva IR, Błaszkowski J, Jobim K, Maia LC, Barbosa FR, Fiuza PO, Divakar PK, Shenoy BD,
Castañeda-Ruiz RF, Somrithipol S, Lateef AA, Karunarathna SC, Tibpromma S, Mortimer PE,
Wanasinghe DN, Phookamsak R, Xu J, Wang Y, Tian F, Alvarado P, Li DW, Kušan I, Matočec
N, Maharachchikumbura SSN, Papizadeh M, Heredia G, Wartchow F, Bakhshi M, Boehm E,
Youssef N, Hustad VP, Lawrey JD, Santiago ALCMA, Bezerra JDP, Souza-Motta CM, Firmino
AL, Tian Q, Houbraken J, Hongsanan S, Tanaka K, Dissanayake AJ, Monteiro JS, Grossart HP,
Suija A, Weerakoon G, Etayo J, Tsurykau A, Vázquez V, Mungai P, Damm U, Li QR, Zhang H,
Boonmee S, Lu YZ, Becerra AG, Kendrick B, Brearley FQ, Motiejūnaitė J, Sharma B, Khare R,
Gaikwad S, Wijesundara DSA, Tang LZ, He MQ, Flakus A, Rodriguez-Flakus P, Zhurbenko MP,
McKenzie EHC, Stadler M, Bhat DJ, Liu JK, Raza M, Jeewon R, Nassonova ES, Prieto M,
Jayalal RGU, Erdoğdu M, Yurkov A, Schnittler M, Shchepin ON, Novozhilov YK, Silva-Filho
AGS, Liu P, Cavender JC, Kang Y, Mohammad S, Zhang LF, Xu RF, Li YM, Dayarathne MC,
Ekanayaka AH, Wen TC, Deng CY, Pereira OL, Navathe S, Hawksworth DL, Fan XL,
Dissanayake LS, Kuhnert E, Grossart HP, Thines M 2020 – Outline of Fungi and fungus-like
taxa. Mycosphere 11(1), 1060–1456, Doi 10.5943/mycosphere/11/1/8

1064
Abstract
This article provides an outline of the classification of the kingdom Fungi (including fossil
fungi. i.e. dispersed spores, mycelia, sporophores, mycorrhizas). We treat 19 phyla of fungi. These
are Aphelidiomycota, Ascomycota, Basidiobolomycota, Basidiomycota, Blastocladiomycota,
Calcarisporiellomycota, Caulochytriomycota, Chytridiomycota, Entomophthoromycota,
Entorrhizomycota, Glomeromycota, Kickxellomycota, Monoblepharomycota, Mortierellomycota,
Mucoromycota, Neocallimastigomycota, Olpidiomycota, Rozellomycota and Zoopagomycota. The
placement of all fungal genera is provided at the class-, order- and family-level. The described
number of species per genus is also given. Notes are provided of taxa for which recent changes or
disagreements have been presented. Fungus-like taxa that were traditionally treated as fungi are
also incorporated in this outline (i.e. Eumycetozoa, Dictyosteliomycetes, Ceratiomyxomycetes and
Myxomycetes). Four new taxa are introduced: Amblyosporida ord. nov. Neopereziida ord. nov.
and Ovavesiculida ord. nov. in Rozellomycota, and Protosporangiaceae fam. nov. in
Dictyosteliomycetes. Two different classifications (in outline section and in discussion) are
provided for Glomeromycota and Leotiomycetes based on recent studies. The phylogenetic
reconstruction of a four-gene dataset (18S and 28S rRNA, RPB1, RPB2) of 433 taxa is presented,
including all currently described orders of fungi.

Keywords – Four new taxa – Ascomycota – Amblyosporida ord. nov. – Basal clades –
Basidiomycota – Classification – Emendation – Microsporidia – Neopereziida ord. nov. –
Ovavesiculida ord. nov. – Protosporangiaceae fam. nov. – Redonographaceae stat nov.

Introduction
Classification of the kingdom Fungi has been updated continuously, with the frequent
inclusion of data from DNA sequences in recent studies. Re-collecting historic taxa and neo- or
epitypifying them by using both fresh material and cultures is also an increasingly common
practice among mycologists, although yet not easily accomplished in some groups. Utilization of
environmental sequences for recognizing taxa that are not observed directly and naming them
with only a sequence as a holotype is a controversial topic that remains to be addressed
(Hongsanan et al. 2018, Lücking & Hawksworth 2018, Lücking et al. 2018, Thines et al. 2018,
Zamora et al. 2018).
Tedersoo et al. (2018) proposed a novel classification for the kingdom Fungi that was based
on phylogenies and the divergence time of particular taxa. Using these criteria, they accepted 18
phyla: Aphelidiomycota, Ascomycota, Basidiobolomycota, Basidiomycota, Blastocladiomycota,
Calcarisporiellomycota, Caulochytriomycota, Chytridiomycota, Entomophthoromycota,
Glomeromycota, Kickxellomycota, Monoblepharomycota, Mortierellomycota, Mucoromycota,
Neocallimastigomycota, Olpidiomycota, Rozellomycota and Zoopagomycota. This study was,
however, based on only 111 taxa and it remains to be seen how widely it will be accepted and
stand up as more taxa are analyzed. Outlines for the Ascomycota (and notes for genera in the
Ascomycota) and the basal clades of fungi (Wijayawardene et al. 2017, 2018a, b) have been
published, with the participation of experts in particular groups. Jaklitsch et al. (2016a) provided a
synopsis of accepted Ascomycota families with descriptions and lists of included genera (and their
synonyms), and Begerow et al. (2018) prepared a parallel treatment for the families of
Basidiomycota and Entorrhizomycota, including brief diagnoses and indications of ecology and
distributions for all genera (though without listing synonyms of genera and with some genera that
are still debated). A separate outline, with notes and divergence times of Basidiomycota was also
published by He et al. (2019).

Classification of basal clades


The higher level classification of basal clades has been subjected to drastic changes as in
Tedersoo et al. (2018), who took up Rozellomycota to include Cryptomycota and Microsporidia
and also accepted Aphelidiomycota in a fungal clade as did Letcher & Powell (2019) in a synopsis

1065
of that group. Moreover, classes and orders of respective phyla were also provided in Tedersoo et
al. (2018). Wijayawardene et al. (2018b) provided a detailed classification system (from phyla to
genera) for basal clades of Fungi, agreeing with Tedersoo et al. (2018). Hence, Wijayawardene et
al. (2018b) accepted 16 phyla viz. Aphelidiomycota, Basidiobolomycota, Blastocladiomycota,
Calcarisporiellomycota, Caulochytriomycota, Chytridiomycota, Entomophthoromycota,
Glomeromycota, Kickxellomycota, Monoblepharomycota, Mortierellomycota, Mucoromycota,
Neocallimastigomycota, Olpidiomycota, Rozellomycota and Zoopagomycota.

Classification of Glomeromycota
Classification of Glomeromycota (which includes arbuscular mycorrhizal fungi [AMF]) has
been a subject of debate. We provide two different classifications (phyla to genera) which are
commonly used by taxonomists (see outline and discussion). In the outline section, we provide the
classification which is supported by Tedersoo et al. (2018) and our analyses (Fig. 1). The
classification provided in Wijayawardene et al. (2018b) is included in the discussion.

Placement of the Rozellomycota in the tree of life


The position of Microsporidia in the Eukaryotic Tree of Life has been a subject of
discussion. Primarily identified as yeast-like fungi in Schizomycetes (Nägeli 1857), they were
further recognized as protists, while drastic reconsiderations of taxonomy of unicellular
eukaryotes resulted in placing to Sporozoa (Balbiani 1882), and particularly Cnidosporidia (Labbé
1899); Sarcodina in Protozoa (Cavalier-Smith 1981); Archezoa (Cavalier-Smith 1983) and
Protista (Puytorac et al. 1987); as well as to different classes of Fungi (Keeling et al. 2000, Gill &
Fast 2006, James et al. 2006, 2013). The mycological community has widely accepted the
affiliation of Microsporidia with the early diverging clades of Fungi. The Microsporidia,
Cryptomycota and Aphelidea, have also been considered to represent a monophyletic lineage with
shared ecological and structural features, defined as superphylum Opisthosporidia belonging to
supergroup Opisthokontha and separated from Fungi (Karpov et al. 2014). In another system,
however, it was proposed that Cryptomycota (also known as Rozellida, Rozellomycota, or
Rozellosporidia) and Microsporidia join the phylum Rozellomycota, while Aphelidea were
considered as a separate, though related phylum and all these groups were considered basal
lineages of the kingdom Fungi (Tedersoo et al. 2018). The taking up of the name Rozellomycota
in such a broad sense appears premature, especially as the structure and biological features of a
larger part of these organisms are unclear as they are known only from environmental sequences.
The borders between Fungi and Protista are therefore unstable and final delimitation of taxa is
problematic due to poor coverage of molecular data for the representatives of the most basal
groups. However, whatever the conclusion is on placement, it has been decided that the
nomenclature of the names in Microsporidia will continue to follow the International Code of
Zoological Nomenclature even if they are treated as fungi (Turland et al. 2018).

Classification of Ascomycota
Periodic outlines of the Ascomycota have been issued since 1982, with notes in the journal
Systema Ascomycetum which was devoted to this project, and later by Lumbsch & Huhndorf
(2010) who accepted three subphyla: Pezizomycotina with eleven classes, the Saccharomycotina
with one class, and Taphrinomycotina with four classes. The taxonomy of the phylum has been
rapidly updated over the last few years (Hyde et al. 2013, 2017, 2020, Jaklitsch et al. 2016a,
Ekanayaka et al. 2017, Hongsanan et al. 2017, Liu et al. 2017).
Recently, two studies were published on the classification of Leotiomycetes. These are
Ekanayaka et al. (2019) based on a five-locus phylogeny, Johnston et al. (2019) based on
genomic-scale and 15-gene phylogenies. We provide two outlines; i) based on Johnston et al.

1066
(2019), Karakehian et al (2019) and Quijada et al. (2020); and ii) based on Ekanayaka et al.
(2019). These classifications are placed in the general outline and discussion, respectively.
Moreover, the concept of One fungus-One name, which ended the use of different names for
morphs of the same fungus in July 2011, has resulted in several name changes in pleomorphic
genera. Wijayawardene et al. (2018a) provided an updated outline of Ascomycota with three
subphyla - Pezizomycotina (including the 13 classes Arthoniomycetes, Coniocybomycetes,
Dothideomycetes, Eurotiomycetes, Geoglossomycetes, Laboulbeniomycetes, Lecanoromycetes,
Leotiomycetes, Lichinomycetes, Orbiliomycetes, Pezizomycetes, Sordariomycetes, Xylonomycetes
and Xylobotryomycetes), Saccharomycotina (with only class Saccharomycetes) and
Taphrinomycotina (with five classes Archaeorhizomycetes, Neolectomycetes,
Pneumocystidomycetes, Schizosaccharomycetes and Taphrinomycetes). These taxa along with a
summary of other taxonomic ranks are summarized in Table 1.

Classification of Basidiomycota
Basidiomycota constitute a major phylum of the kingdom Fungi and is second in numbers of
described species, to Ascomycota (Wijayawardene et al. 2017, 2018a). Since the last edition of
Ainsworth & Bisby’s Dictionary of the Fungi (Kirk et al. 2008), numerous sequenced-based
studies have enabled the introduction of a vast array of new taxa, which has greatly enriched the
known diversity of Basidiomycota. It has also become clear that several basidiome forms can be
found in the same order, family, or even genus (Hibbett et al. 2007). At the same time, related new
taxonomic categories have been proposed. For example, in phylogenetic studies of
basidiomycetous yeasts, three new classes Malasseziomycetes, Monilielliomycetes, and
Spiculogloeomycetes, were introduced and three new orders, 16 new families, and 47 new genera
were also introduced (Nasr et al. 2014, Wang et al. 2014a, 2015a, b, Liu et al. 2015, Riess et al.
2016). On the other hand, many new changes have also occurred in the Agaricomycotina.
Approximately 60 new genera have been recognized for agarics, 40 for boletes, and 50 for bracket
fungi (Desjardin et al. 2009, Hjortstam & Ryvarden 2010, Petersen & Hughes 2010, Cui et al.
2011, Vellinga et al. 2011, Vizzini et al. 2011, Hao et al. 2014, Hofstetter et al. 2014, Smith et al.
2015, Castellano et al. 2016, Henkel et al. 2016, Wu et al. 2016, Buyck et al. 2017, Orihara &
Smith 2017). Attention has already been drawn to the valuable syntheses of accepted genera, with
diagnosis, provided by Begerow et al. (2018).
The phylogeny and divergence time ranges for higher level Basidiomycota, with the phylum
originating ca. 530 Mya, the subphyla 406–490 Mya, most classes 245–393 Mya and orders 120–
290 Mya were inferred by Zhao et al. (2017). The outline includes 1928 genera with 1263
synonyms within Basidiomycota (He et al. 2019). The latest version of Ainsworth & Bisby’s
Dictionary of the Fungi (Kirk et al. 2008), contains three subphyla, 16 classes, 52 orders, 177
families, 1589 genera and 31515 species in Basidiomycota. The updated outline of Basidiomycota
has updated the numbers in Kirk et al. (2008) to four subphyla, 18 classes, 68 orders, 241 families,
1928 genera and 41270 species in Basidiomycota (He et al. 2019). Agaricomycotina embrace most
of the species in Basidiomycota (30788 species) which includes three classes, 29 orders, 150
families and 1514 genera. Pucciniomycotina was estimated to comprise 8653 species including 10
classes, 22 orders, 49 families and 270 genera. Ustilaginomycetes with 1185 species is the largest
group in Ustilaginomycotina. Malasseziomycetes and Moniliellomycetes, are the only two new
classes recognized in the Basidiomycota since 2008, include 32 estimated species.
Wallemiomycotina is a recently recognized subphylum (Zhao et al. 2017) with 12 species
estimated in a single class, two orders and two families. Another early diverging group in the
Agaricomycotina has recently identified by a phylogenomic study, the Bartheletiomycetes,
including just a single species associated with Gingko biloba as a ‘living fossil’ (Mishra et al.
2018). Accepted taxa of Basidiomycota are summarized in Table 1.

1067
Table 1 Phyla, classes, orders and families of kingdom Fungi. The number of accepted genera in
each family is indicated in brackets after the family name.

Phylum Class* Order* Family*


Aphelidiomycota Aphelidiomycetes Aphelidiales Aphelidiaceae (4)
Ascomycota Archaeorhizomycetes Archaeorhizomycetales Archaeorhizomycetaceae (1)
Arthoniomycetes Arthoniales Andreiomycetaceae (1)
Arthoniaceae (23)
Chrysotrichaceae (3)
Lecanographaceae (7)
Opegraphaceae (15)
Roccellaceae (41)
Roccellographaceae (3)
Arthoniales genera
incertae sedis (23)
Lichenostigmatales Phaeococcomycetaceae (3)
Candelariomycetes Candelariales Candelariaceae (4)
Pycnoraceae (1)
Coniocybomycetes Coniocybales Coniocybaceae (2)
Dothideomycetes Abrothallales Abrothallaceae (2)
Acrospermales Acrospermaceae (3)
Acrospermales genus
incertae sedis (1)
Asterinales Asterinaceae (19)
Asterotexaceae (1)
Hemigraphaceae (1)
Lembosiaceae (1)
Melaspileellaceae (1)
Neobueliellaceae (1)
Stictographaceae (5)
Asterinales genera
incertae sedis (8)
Botryosphaeriales Aplosporellaceae (2)
Botryosphaeriaceae (22)
Melanopsaceae (1)
Phyllostictaceae (2)
Planistromellaceae (4)
Saccharataceae (4)
Botryosphaeriales
genera incertae sedis (8)
Capnodiales Aeminiaceae (1)
Antennulariellaceae (4)
Capnodiaceae (9)
Cladosporiaceae (8)
Cystocoleaceae (1)
Dissoconiaceae (5)
Euantennariaceae (7)
Extremaceae (8)
Johansoniaceae (2)
Metacapnodiaceae (3)
Mycosphaerellaceae (111)
Neodevriesiaceae (2)
Phaeothecaceae (1)
Phaeothecoidiellaceae (8)
Piedraiaceae (1)
Racodiaceae (1)
Schizothyriaceae (10)

1068
Table 1 Continued.

Phylum Class* Order* Family*


Teratosphaeriaceae (60)
Xenodevriesiaceae (1)
Capnodiales genera
incertae sedis (14)
Catinellales Catinellaceae (1)
Cladoriellales Cladoriellaceae (1)
Collemopsidiales Xanthopyreniaceae (6)
Dothideales Dothideaceae (13)
Neocelosporiaceae (3)
Saccotheciaceae (7)
Zalariaceae (1)
Dothideales genera
incertae sedis (6)
Dyfrolomycetales Pleurotremataceae (3)
Eremithallales Melaspileaceae (2)
Eremomycetales Eremomycetaceae (2)
Eremomycetales genus
incertae sedis (1)
Gloniales Gloniaceae (3)
Hysteriales Hysteriaceae (13)
Hysteriales genus
incertae sedis (1)
Jahnulales Aliquandostipitaceae (7)
Manglicolaceae (1)
Kirschsteiniotheliales Kirschsteiniotheliaceae (1)
Kirschsteiniotheliales
genera incertae sedis (2)
Lembosinales Lembosinaceae (1)
Lichenotheliales Lichenotheliaceae (2)
Microthyriales Microthyriaceae (11)
Microthyriales genera
incertae sedis (6)
Minutisphaerales Acrogenosporaceae (1)
Minutisphaeraceae (1)
Monoblastiales Monoblastiaceae (6)
Murramarangomycetales Murramarangomycetaceae
(1)
Muyocopronales Muyocopronaceae (9)
Myriangiales Elsinoaceae (2)
Myriangiaceae (11)
Myriangiales genus
incertae sedis (1)
Mytilinidiales Mytilinidiaceae (9)
Natipusillales Natipusillaceae (1)
Parmulariales Parmulariaceae (35)
Patellariales Patellariaceae (21)
Phaeotrichales Phaeotrichaceae (3)
Pleosporales Acrocalymmaceae (1)
Aigialaceae (6)
Amniculicolaceae (6)
Amorosiaceae (4)
Anteagloniaceae (2)
Aquasubmersaceae (1)
Arthopyreniaceae (2)

1069
Table 1 Continued.

Phylum Class* Order* Family*


Ascocylindricaceae (1)
Astrosphaeriellaceae (7)
Bambusicolaceae (3)
Biatriosporaceae (1)
Camarosporiaceae (2)
Camarosporidiellaceae (1)
Caryosporaceae (1)
Coniothyriaceae (5)
Corynesporascaceae (2)
Cryptocoryneaceae (1)
Cucurbitariaceae (13)
Cyclothyriellaceae (2)
Dacampiaceae (6)
Delitschiaceae (3)
Diademaceae (2)
Dictyosporiaceae (15)
Didymellaceae (33)
Didymosphaeriaceae (32)
Dothidotthiaceae (7)
Fuscostagonosporaceae (1)
Fusculinaceae (2)
Halojulellaceae (1)
Halotthiaceae (6)
Hermatomycetaceae (1)
Hypsostromataceae (1)
Latoruaceae (4)
Lentimurisporaceae (2)
Lentitheciaceae (13)
Leptosphaeriaceae (13)
Libertasomycetaceae (2)
Ligninsphaeriaceae (2)
Lindgomycetaceae (7)
Lizoniaceae (1)
Longipedicellataceae (3)
Longiostiolaceae (1)
Lophiostomataceae (28)
Lophiotremataceae (7)
Macrodiplodiopsidaceae (2)
Massariaceae (3)
Massarinaceae (8)
Melanommataceae (35)
Morosphaeriaceae (6)
Mycoporaceae (1)
Neocamarosporiaceae (2)
Neohendersoniaceae (5)
Neomassariaceae (1)
Neomassarinaceae (2)
Neophaeosphaeriaceae (1)
Neopyrenochaetaceae (1)
Nigrogranaceae (1)
Occultibambusaceae (5)
Ohleriaceae (1)
Parabambusicolaceae (9)
Paradictyoarthriniaceae (2)

1070
Table 1 Continued.

Phylum Class* Order* Family*


Paralophiostomataceae (1)
Parapyrenochaetaceae (2)
Periconiaceae (4)
Phaeoseptaceae (2)
Phaeosphaeriaceae (82)
Pleomassariaceae (7)
Pleomonodictydaceae (2)
Pleosporaceae (23)
Pseudoastrosphaeriellaceae
(3)
Pseudoberkleasmiaceae (1)
Pseudocoleodictyosporaceae
(2)
Pseudolophiotremataceae (2)
Pseudomassarinaceae (1)
Pseudopyrenochaetaceae (1)
Pyrenochaetopsidaceae (3)
Roussoellaceae (12)
Salsugineaceae (2)
Shiraiaceae (3)
Sporormiaceae (9)
Striatiguttulaceae (2)
Sulcatisporaceae (6)
Teichosporaceae (13)
Testudinaceae (9)
Tetraplosphaeriaceae (8)
Thyridariaceae (7)
Torulaceae (6)
Trematosphaeriaceae (6)
Tzeananiaceae (1)
Wicklowiaceae (1)
Zopfiaceae (6)
Pleosporales genera
incertae sedis (48)
Stigmatodiscales Stigmatodiscaceae (1)
Strigulales Strigulaceae (13)
Tenuitholiascaceae (1)
Superstratomycetales Superstratomycetaceae (1)
Trypetheliales Polycoccaceae (2)
Trypetheliaceae (19)
Tubeufiales Bezerromycetaceae (3)
Tubeufiaceae (46)
Wiesneriomycetaceae (6)
Valsariales Valsariaceae (3)
Venturiales Sympoventuriaceae (9)
Venturiaceae (14)
Venturiales genera
incertae sedis (3)
Zeloasperisporiales Zeloasperisporiaceae (1)
Incertae sedis* Alinaceae (1)
Argynnaceae (2)
Ascoporiaceae (1)
Aulographaceae (4)
Balladynaceae (3)

1071
Table 1 Continued.

Phylum Class* Order* Family*


Cleistosphaeraceae (1)
Coccoideaceae (3)
Cookellaceae (2)
Dimeriaceae (1)
Dubujianaceae (1)
Dysrhynchisceae (1)
Endosporiaceae (1)
Englerulaceae (8)
Eremomycetaceae (3)
Eriomycetaceae (5)
Homortomycetaceae (1)
Hyalomeliolinaceae (1)
Leptopeltidaceae (4)
Macrovalsariaceae (1)
Meliolinaceae (2)
Mesnieraceae (3)
Naetrocymbaceae (5)
Nematotheciaceae (3)
Neoparodiaceae (1)
Palawaniaceae (1)
Paranectriellaceae (2)
Parodiellaceae (1)
Perisporiopsidaceae (5)
Phaeodimeriellaceae (1)
Pododimeriaceae (2)
Polyclypeolinaceae (1)
Polystomellaceae (4)
Protoscyphaceae (1)
Pseudoperisporiaceae (4)
Pseudorobillardaceae (1)
Pyrenidiaceae (1)
Seynesiopeltidaceae (1)
Stomatogeneceae (1)
Thyrinulaceae (3)
Toroaceae (1)
Trichopeltinaceae (7)
Trichothyriaceae (4)
Vizellaceae (3)
Dothideomycetes
genera incertae sedis
(278)
Eurotiomycetes Arachnomycetales Arachnomycetaceae (2)
Chaetothyriales Chaetothyriaceae (20)
Coccodiniaceae (3)
Cyphellophoraceae (2)
Epibryaceae (1)
Herpotrichiellaceae (16)
Lyrommataceae (1)
Microtheliopsidaceae (1)
Paracladophialophoraceae
(1)
Pyrenotrichaceae (2)
Trichomeriaceae (8)

1072
Table 1 Continued.

Phylum Class* Order* Family*


Chaetothyriales genera
incertae sedis (10)
Coryneliales Coryneliaceae (7)
Eremascaceae (1)
Eurotiales Aspergillaceae (15)
Elaphomycetaceae (2)
Thermoascaceae (2)
Trichocomaceae (7)
Mycocaliciales Mycocaliciaceae (7)
Onygenales Ajellomycetaceae (6)
Arthrodermataceae (10)
Ascosphaeraceae (3)
Gymnoascaceae (10)
Nannizziopsidaceae (1)
Onygenaceae (31)
Spiromastigaceae (4)
Onygenales genera
incertae sedis (3)
Phaeomoniellales Celotheliaceae (10)
Pyrenulales Pyrenulaceae (12)
Pyrenulales genera
incertae sedis (2)
Sclerococcales Dactylosporaceae (5)
Verrucariales Adelococcaceae (3)
Sarcopyreniaceae (1)
Verrucariaceae (52)
Verrucariales genera
incertae sedis (5)
Incertae sedis Rhynchostomataceae (2)
Eurotiomycetes
genus incertae sedis
(1)
Geoglossomycetes Geoglossales Geoglossaceae (7)
Geoglossomycetes
genera incertae
sedis (2)
Laboulbeniomycetes Herpomycetales Herpomycetaceae (1)
Laboulbeniales Ceratomycetaceae (12)
Euceratomycetaceae (5)
Laboulbeniaceae (125)
Teratomyceteae (1)
Laboulbeniales genera
incertae sedis (3)
Pyxidiophorales Pyxidiophoraceae (3)
Laboulbeniomycetes
genus incertae sedis
(1)
Lecanoromycetes Acarosporales Acarosporaceae (11)
Eigleraceae (1)
Baeomycetales Arctomiaceae (4)
Arthrorhaphidaceae (1)
Baeomycetaceae (5)
Cameroniaceae (1)
Hymeneliaceae (3)
Protothelenellaceae (3)

1073
Table 1 Continued.

Phylum Class* Order* Family*


Trapeliaceae (12)
Xylographaceae (4)
Caliciales Caliciaceae (36)
Physciaceae (18)
Graphidales Diploschistaceae(35)
Fissurinaceae (6)
Gomphillaceae (26)
Graphidaceae (31)
Redonographaceae (2)
Thelotremataceae (7)
Gyalectales Coenogoniaceae (1)
Gyalectaceae (3)
Phlyctidaceae (2)
Sagiolechiaceae (2)
Trichotheliaceae (7)
Lecanorales Bruceomycetaceae (2)
Catillariaceae (5)
Cladoniaceae (22)
Gypsoplacaceae (1)
Haematommataceae (1)
Lecanoraceae (28)
Malmideaceae (7)
Megalariaceae (2)
Parmeliaceae (71)
Pilocarpaceae (32)
Psilolechiaceae (1)
Psoraceae (6)
Ramalinaceae (37)
Ramboldiaceae (1)
Scoliciosporaceae (1)
Sphaerophoraceae (6)
Tephromelataceae (4)
Lecanorales genera
incertae sedis (14)
Lecideales Lecideaceae (29)
Lopadiaceae (1)
Leprocaulales Leprocaulaceae (3)
Micropeltidales Micropeltidaceae (12)
Ostropales Odontotremataceae (10)
Phaneromycetaceae (1)
Spirographaceae (1)
Stictidaceae (28)
Ostropales genera
incertae sedis (6)
Peltigerales Coccocarpiaceae (3)
Collemataceae (8)
Koerberiaceae (3)
Massalongiaceae (3)
Pannariaceae (27)
Peltigeraceae (15)
Placynthiaceae (3)
Vahliellaceae (1)
Pertusariales Agyriaceae (2)
Coccotremataceae (3)

1074
Table 1 Continued.

Phylum Class* Order* Family*


Icmadophilaceae (7)
Megasporaceae (6)
Microcaliciaceae (1)
Ochrolechiaceae (1)
Pertusariaceae (3)
Varicellariaceae (1)
Variolariaceae (1)
Rhizocarpales Rhizocarpaceae (4)
Sarrameanales Sarrameanaceae (2)
Schaereriales Schaereriaceae (1)
Sporastatiales Sporastatiaceae (2)
Teloschistales Brigantiaeaceae (2)
Megalosporaceae (3)
Teloschistaceae (63)
Teloschistales genus
incertae sedis (1)
Thelenellales Thelenellaceae (3)
Turquoiseomycetales Turquoiseomycetaceae (1)
Umbilicariales Elixiaceae (2)
Fuscideaceae (4)
Ophioparmaceae (3)
Ropalosporaceae (1)
Umbilicariaceae (3)
Incertae sedis Epigloeaceae (1)
Lecanoromycetes
genera incertae
sedis (15)
Leotiomycetes Chaetomellales Chaetomellaceae (4)
Cyttariales Cyttariaceae (1)
Helotiales Amicodiscaceae (1)
Arachnopezizaceae (5)
Ascocorticiaceae (1)
Ascodichaenaceae (2)
Bloxamiaceae (1)
Bryoglossaceae (3)
Calloriaceae (14)
Cenangiaceae (11)
Chlorociboriaceae (1)
Chlorospleniaceae (1)
Chrysodiscaceae (1)
Cordieritidaceae (18)
Dermateaceae (14)
Discinellaceae (12)
Drepanopezizaceae (8)
Erysiphaceae (20)
Gelatinodiscaceae (9)
Godroniaceae (5)
Helotiaceae (33)
Heterosphaeriaceae (1)
Hyaloscyphaceae (37)
Lachnaceae (18)
Leptodontidiaceae (1)
Loramycetaceae (2)
Mitrulaceae (1)

1075
Table 1 Continued.

Phylum Class* Order* Family*


Mollisiaceae (18)
Myxotrichaceae (3)
Neolauriomycetaceae (3)
Pezizellaceae (23)
Ploettnerulaceae (12)
Rutstroemiaceae (7)
Sclerotiniaceae (31)
Vibrisseaceae (5)
Helotiales genera
incertae sedis (144)
Lahmiales Lahmiaceae (1)
Lauriomycetales Lauriomycetaceae (1)
Leotiales Cochlearomycetaceae (2)
Leotiaceae (4)
Mniaeciaceae (2)
Tympanidaceae (7)
Leotiales genera
incertae sedis (3)
Lichinodiales Lichinodiaceae (1)
Marthamycetales Marthamycetaceae (9)
Medeolariales Medeolariaceae (1)
Micraspidales Micraspidaceae (1)
Phacidiales Helicogoniaceae (7)
Phacidiaceae (9)
Phacidiales genus
incertae sedis (1)
Rhytismatales Cudoniaceae (2)
Rhytismataceae (52)
Triblidiaceae (2)
Rhytismatales genera
incertae sedis (9)
Thelebolales Pseudeurotiaceae (8)
Thelebolaceae (10)
Thelebolales genera
incertae sedis (3)
Leotiomycetes
genera incertae
sedis (20)
Lichinomycetes Lichinales Gloeoheppiaceae (3)
Lichinaceae (43)
Peltulaceae (1)
Neolectomycetes Neolectales Neolectaceae (1)
Orbiliomycetes Orbiliales Orbiliaceae (12)
Orbiliales genus
incertae sedis (1)
Orbiliomycetes
genus incertae sedis
(1)
Pezizomycetes Pezizales Ascobolaceae (5)
Ascodesmidaceae (10)
Caloscyphaceae (1)
Chorioactidaceae (6)
Discinaceae (5)
Glaziellaceae (1)

1076
Table 1 Continued.

Phylum Class* Order* Family*


Helvellaceae (5)
Kallistoskyphaceae (1)
Karstenellaceae (1)
Morchellaceae (7)
Pezizaceae (38)
Pseudombrophilaceae (4)
Pulvinulaceae (3)
Pyronemataceae (61)
Rhizinaceae (3)
Sarcoscyphaceae (12)
Sarcosomataceae (9)
Strobiloscyphaceae (1)
Tarzettaceae (6)
Tuberaceae (7)
Pezizales genera incertae
sedis (18)
Pneumocystomycetes Pneumocystidales Pneumocystidaceae (1)
Saccharomycetes Saccharomycetales Alloascoideaceae (1)
Ascoideaceae (1)
Cephaloascaceae (1)
Debaryomycetaceae (13)
Dipodascaceae (5)
Lipomycetaceae (5)
Metschnikowiaceae (3)
Phaffomycetaceae (5)
Pichiaceae (10)
Saccharomycetaceae (16)
Saccharomycodaceae (2)
Saccharomycopsidaceae (2)
Trichomonascaceae (9)
Trigonopsidaceae (3)
Saccharomycetales
genera incertae sedis (21)
Schizosaccharomycetes Schizosaccharomycetales Schizosaccharomycetaceae
(1)
Sordariomycetes Amphisphaeriales Amphisphaeriaceae (3)
Apiosporaceae (5)
Beltraniaceae (9)
Clypeophysalosporaceae
(4)
Cylindriaceae (1)
Hansfordiaceae (1)
Hyponectriaceae (17)
Iodosphaeriaceae (1)
Melogrammataceae (1)
Phlogicylindriaceae (3)
Pseudomassariaceae (4)
Pseudotruncatellaceae (1)
Sporocadaceae (33)
Vialaeaceae (1)
Amphisphaeriales genus
incertae sedis (1)
Amplistromatales Amplistromataceae (3)

1077
Table 1 Continued.

Phylum Class* Order* Family*


Catabotryaceae (1)
Annulatascales Annulatascaceae (10)
Annulatascales genus
incertae sedis (1)
Atractosporales Atractosporaceae (2)
Conlariaceae (2)
Pseudoproboscisporaceae (2)
Boliniales Boliniaceae (9)
Calosphaeriales Calosphaeriaceae (4)
Jobellisiaceae (1)
Pleurostomataceae (1)
Calosphaeriales genera
incertae sedis (4)
Cephalothecales Cephalothecaceae (5)
Chaetosphaeriales Chaetosphaeriaceae (44)
Helminthosphaeriaceae (7)
Leptosporellaceae (1)
Leptosporellaceae (1)
Linocarpaceae (2)
Chaetosphaeriales
genera incertae sedis
(7)
Coniochaetales Coniochaetaceae (2)
Cordanaceae (1)
Coniochaetales genera
incertae sedis (2)
Conioscyphales Conioscyphaceae (1)
Coronophorales Bertiaceae (2)
Ceratostomataceae (14)
Chaetosphaerellaceae (3)
Coronophoraceae (1)
Nitschkiaceae (14)
Scortechiniaceae (10)
Coronophorales genera
incertae sedis (2)
Delonicicolales Delonicicolaceae (2)
Diaporthales Apiosporopsidaceae (1)
Apoharknessiaceae (2)
Asterosporiaceae (1)
Auratiopycnidiellaceae (1)
Coryneaceae (1)
Cryphonectriaceae (28)
Cytosporaceae (6)
Diaporthaceae (15)
Diaporthosporellaceae (1)
Diaporthostomataceae (1)
Dwiroopaceae (1)
Erythrogloeaceae (4)
Gnomoniaceae (36)
Harknessiaceae (2)
Juglanconidaceae (1)
Lamproconiaceae (2)

1078
Table 1 Continued.

Phylum Class* Order* Family*


Macrohilaceae (1)
Melanconidaceae (1)
Melanconiellaceae (7)
Neomelanconiellaceae (1)
Phaeoappendicosporaceae
(2)
Prosopidicolaceae (1)
Pseudomelanconidaceae (2)
Pseudoplagiostomataceae (1)
Schizoparmaceae (1)
Stilbosporaceae (4)
Sydowiellaceae (16)
Synnemasporellaceae (1)
Tubakiaceae (8)
Diaporthales genera
incertae sedis (36)
Distoseptisporales Distoseptisporaceae (1)
Falcocladiales Falcocladiaceae (1)
Fuscosporellales Fuscosporellaceae (6)
Glomerellales Australiascaceae (1)
Glomerellaceae (1)
Malaysiascaceae (1)
Plectosphaerellaceae (24)
Reticulascaceae (4)
Glomerellales genus
incertae sedis (1)
Hypocreales Bionectriaceae (47)
Calcarisporiaceae (1)
Clavicipitaceae (42)
Cocoonihabitaceae (1)
Cordycipitaceae (17)
Flammocladiellaceae (1)
Hypocreaceae (17)
Myrotheciomycetaceae (4)
Nectriaceae (69)
Niessliaceae (21)
Ophiocordycipitaceae (10)
Sarocladiaceae (2)
Stachybotryaceae (39)
Tilachlidiaceae (3)
Hypocreales genera
incertae sedis (29)
Jobellisiales Jobellisiaceae (1)
Koralionastetales Koralionastetaceae (2)
Lulworthiales Lulworthiaceae (15)
Magnaporthales Ceratosphaeriaceae (1)
Magnaporthaceae (22)
Ophioceraceae (1)
Pseudohalonectriaceae (1)
Pyriculariaceae (11)
Meliolales Armatellaceae (1)
Meliolaceae (8)
Microascales Ceratocystidaceae (11)

1079
Table 1 Continued.

Phylum Class* Order* Family*


Chadefaudiellaceae (2)
Gondwanamycetaceae (2)
Graphiaceae (1)
Halosphaeriaceae (66)
Microascaceae (23)
Triadelphiaceae (2)
Microascales genera
incertae sedis (6)
Myrmecridiales Myrmecridiaceae (2)
Xenodactylariaceae (1)
Ophiostomatales Kathistaceae (2)
Ophiostomataceae (13)
Pararamichloridiales Pararamichloridiaceae (1)
Parasympodiellales Parasympodiellaceae (1)
Phomatosporales Phomatosporaceae (3)
Phyllachorales Phaeochoraceae (4)
Phaeochorellaceae (1)
Phyllachoraceae (54)
Telimenaceae (1)
Phyllachorales genus
incertae sedis (1)
Pisorisporiales Pisorisporiaceae (2)
Pleurotheciales Pleurotheciaceae (11)
Pseudodactylariales Pseudodactylariaceae (1)
Savoryellales Savoryellaceae (4)
Sordariales Chaetomiaceae (37)
Lasiosphaeriaceae (32)
Podosporaceae (3)
Sordariaceae (7)
Sordariales genera
incertae sedis (22)
Spathulosporales Hispidicarpomycetaceae (1)
Spathulosporaceae (2)
Sporidesmiales Sporidesmiaceae (1)
Tirisporellales Tirisporellaceae (3)
Togniniales Togniniaceae (2)
Torpedosporales Etheirophoraceae (2)
Juncigenaceae (5)
Torpedosporaceae (1)
Tracyllalales Tracyllaceae (1)
Vermiculariopsiellales Vermiculariopsiellaceae (1)
Xenospadicoidales Xenospadicoidaceae (5)
Xylariales Anungitiomycetaceae (1)
Barrmaeliaceae (2)
Castanediellaceae (1)
Clypeosphaeriaceae (6)
Coniocessiaceae (2)
Diatrypaceae (20)
Fasciatisporaceae (1)
Graphostromataceae (5)
Hypoxylaceae (19)
Induratiaceae (2)
Leptosilliaceae (1)
Lopadostomataceae (4)

1080
Table 1 Continued.

Phylum Class* Order* Family*


Microdochiaceae (3)
Myelospermataceae (1)
Nothodactylariaceae (1)
Oxydothidaceae (1)
Polystigmataceae (1)
Pseudosporidesmiaceae (1)
Requienellaceae (4)
Xyladictyochaetaceae (1)
Xylariaceae (32)
Zygosporiaceae (1)
Xylariales genera
incertae sedis (56)
Incertae sedis Acrodictyaceae (1)
Barbatosphaeriaceae (3)
Batistiaceae (1)
Cainiaceae (6)
Junewangiaceae (2)
Lautosporaceae (1)
Obryzaceae (1)
Papulosaceae (4)
Rhamphoriaceae (4)
Thyridiaceae (2)
Trichosphaeriaceae (10)
Woswasiaceae (3)
Sordariomycetes
genera incertae
sedis (131)
Taphrinomycetes Taphrinales Protomycetaceae (6)
Taphrinaceae (1)
Xylobotryomycetes Xylobotryales Cirrosporiaceae (1)
Xylobotryaceae (1)
Xylonomycetes Symbiotaphrinales Symbiotaphrinaceae (1)
Xylonales Xylonaceae (2)
Incertae sedis Thelocarpales Thelocarpaceae (2)
Vezdaeales Vezdaeaceae (1)
Incertae sedis Incertae sedis Aphanopsidaceae (2)
Diporothecaceae (1)
Eoterfeziaceae (2)
Harpidiaceae (2)
Mucomassariaceae (1)
Saccardiaceae (7)
Seuratiaceae (2)
Strangosporaceae (1)
Ascomycota genera
incertae sedis
(1485)
Basidiobolomycota Basidiobolomycetes Basidiobolales Basidiobolaceae (2)
Basidiomycota Agaricomycetes Agaricales Agaricaceae (59)
Amanitaceae (5)
Biannulariaceae (7)
Bolbitiaceae (15)
Broomeiaceae (1)
Chromocyphellaceae (1)
Clavariaceae (10)

1081
Table 1 Continued.

Phylum Class* Order* Family*


Cortinariaceae (5)
Crassisporiaceae (2)
Crepidotaceae (6)
Cyphellaceae (16)
Cystostereaceae (7)
Entolomataceae (7)
Hemigasteraceae (1)
Hydnangiaceae (4)
Hygrophoraceae (26)
Hymenogastraceae (10)
Inocybaceae (3)
Limnoperdaceae (1)
Lycoperdaceae (7)
Lyophyllaceae (18)
Macrocystidiaceae (1)
Marasmiaceae (10)
Mycenaceae (16)
Mythicomycetaceae (2)
Niaceae (9)
Omphalotaceae (14)
Physalacriaceae (28)
Pleurotaceae (5)
Pluteaceae (3)
Porotheleaceae (2)
Psathyrellaceae (13)
Pseudoclitocybaceae (7)
Pterulaceae (13)
Schizophyllaceae (3)
Stephanosporaceae (5)
Strophariaceae (11)
Tricholomataceae (10)
Tubariaceae (7)
Typhulaceae (4)
Agaricales genera
incertae sedis (134)
Amylocorticiales Amylocorticiaceae (11)
Atheliales Atheliaceae (20)
Auriculariales Auriculariaceae (12)
Hyaloriaceae (3)
Auriculariales genera
incertae sedis (31)
Boletales Boletaceae (92)
Boletinellaceae (2)
Calostomataceae (1)
Coniophoraceae (5)
Diplocystidiaceae (4)
Gasterellaceae (1)
Gomphidiaceae (4)
Gyroporaceae (1)
Hygrophoropsidaceae (2)
Paxillaceae (10)
Protogastraceae (1)
Rhizopogonaceae (3)
Sclerodermataceae (5)

1082
Table 1 Continued.

Phylum Class* Order* Family*


Serpulaceae (3)
Suillaceae (2)
Tapinellaceae (3)
Boletales genera
incertae sedis (4)
Cantharellales Aphelariaceae (3)
Botryobasidiaceae (5)
Ceratobasidiaceae (6)
Hydnaceae (21)
Oliveoniaceae (1)
Tulasnellaceae (2)
Bartheletiomycetes Bartheletiales Bartheletiaceae (1)
Cantharellales genera
incertae sedis (8)
Corticiales Corticiaceae (12)
Dendrominiaceae (1)
Punctulariaceae (3)
Vuilleminiaceae (3)
Corticiales genera
incertae sedis (7)
Geastrales Geastraceae (7)
Sclerogastraceae (1)
Geastrales genus
incertae sedis (1)
Gloeophyllales Gloeophyllaceae (12)
Gloeophyllales genus
incertae sedis (1)
Gomphales Clavariadelphaceae (2)
Gomphaceae (14)
Lentariaceae (3)
Hymenochaetales Hymenochaetaceae (40)
Neoantrodiellaceae (1)
Nigrofomitaceae (1)
Oxyporaceae (1)
Rickenellaceae (8)
Schizoporaceae (13)
Hymenochaetales
genera incertae sedis
(15)
Hysterangiales Gallaceaceae (3)
Hysterangiaceae (4)
Mesophelliaceae (8)
Phallogastraceae (2)
Trappeaceae (3)
Jaapiales Jaapiaceae (1)
Lepidostromatales Lepidostromataceae (3)
Phallales Claustulaceae (5)
Gastrosporiaceae (1)
Phallaceae (26)
Phallales genera
incertae sedis (2)
Polyporales Cerrenaceae (4)
Dacryobolaceae (7)
Fomitopsidaceae (25)

1083
Table 1 Continued.

Phylum Class* Order* Family*


Fragiliporiaceae (1)
Gelatoporiaceae (4)
Grifolaceae (2)
Hyphodermataceae (1)
Incrustoporiaceae (5)
Irpicaceae (14)
Ischnodermataceae (1)
Laetiporaceae (3)
Meripilaceae (3)
Meruliaceae (22)
Panaceae (2)
Phanerochaetaceae (18)
Podoscyphaceae (3)
Polyporaceae (85)
Sparassidaceae (3)
Steccherinaceae (22)
Polyporales genera
incertae sedis (67)
Russulales Albatrellaceae (8)
Auriscalpiaceae (6)
Bondarzewiaceae (9)
Echinodontiaceae (3)
Hericiaceae (6)
Hybogasteraceae (1)
Peniophoraceae (16)
Russulaceae (7)
Stereaceae (22)
Xenasmataceae (3)
Russulales genera
incertae sedis (15)
Sebacinales Sebacinaceae (8)
Serendipitaceae (1)
Stereopsidales Stereopsidaceae (1)
Thelephorales Bankeraceae (5)
Thelephoraceae (9)
Thelephorales genus
incertae sedis (1)
Trechisporales Hydnodontaceae (13)
Tremellodendropsidales Tremellodendropsidaceae
(1)
Agaricomycetes
genera incertae
sedis (40)
Agaricostilbomycete Agaricostilbales Agaricostilbaceae (3)
s
Chionosphaeraceae (5)
Kondoaceae (2)
Ruineniaceae (1)
Agaricostilbales genera
incertae sedis (2)
Atractiellomycetes Atractiellales Atractogloeaceae (1)
Hoehnelomycetaceae (2)
Phleogenaceae (7)
Classiculomycetes Classiculales Classiculaceae (2)

1084
Table 1 Continued.

Phylum Class* Order* Family*


Cryptomycocolacomycetes Cryptomycocolacales Cryptomycocolacaceae (2)
Cystobasidiomycetes Buckleyzymales Buckleyzymaceae (1)
Cystobasidiales Cystobasidiaceae (3)
Erythrobasidiales Erythrobasidiaceae (2)
Erythrobasidiales
genera incertae sedis
(3)
Naohideales Naohideaceae (1)
Sakaguchiales Sakaguchiaceae (1)
Incertae sedis Microsporomycetaceae (1)
Incertae sedid Symmetrosporaceae(1)
Cystobasidiomycetes genus
incertae sedis (1)
Dacrymycetes Dacrymycetales Cerinomycetaceae (1)
Dacrymycetaceae (10)
Unilacrymales Unilacrymaceae (1)
Exobasidiomycetes Ceraceosorales Ceraceosoraceae (1)
Doassansiales Doassansiaceae (11)
Melaniellaceae (1)
Rhamphosporaceae (1)
Entylomatales Entylomataceae (2)
Exobasidiales Brachybasidiaceae (5)
Cryptobasidiaceae (6)
Exobasidiaceae (4)
Graphiolaceae (2)
Laurobasidiaceae (1)
Georgefischeriales Eballistraceae (1)
Georgefischeriaceae (2)
Gjaerumiaceae (1)
Tilletiariaceae (3)
Golubeviales Golubeviaceae (1)
Microstromatales Microstromataceae (1)
Quambalariaceae (1)
Volvocisporiaceae (1)
Microstromatales
genera incertae sedis
(4)
Robbauerales Robbaueraceae (1)
Tilletiales Erratomycetaceae (1)
Tilletiaceae (6)
Malasseziomycetes Malasseziales Malasseziaceae (1)
Microbotryomycetes Heterogastridiales Heterogastridiaceae (3)
Kriegeriales Camptobasidiaceae (2)
Kriegeriaceae (4)
Leucosporidiales Leucosporidiaceae (1)
Microbotryales Microbotryaceae (4)
Ustilentylomataceae (4)
Sporidiobolales Sporidiobolaceae (3)
Incertae sedis Chrysozymaceae (4)
Colacogloeaceae (1)
Microbotryomycetes
genera incertae sedis (15)
Mixiomycetes Mixiales Mixiaceae (1)
Moniliellomycetes Moniliellales Moniliellaceae (1)

1085
Table 1 Continued.

Phylum Class* Order* Family*


Pucciniomycetes Helicobasidiales Helicobasidiaceae (2)
Pachnocybales Pachnocybaceae (1)
Platygloeales Eocronartiaceae (5)
Platygloeaceae (4)
Pucciniales Chaconiaceae (9)
Coleosporiaceae (5)
Cronartiaceae (3)
Melampsoraceae (1)
Mikronegeriaceae (3)
Phakopsoraceae (15)
Phragmidiaceae (13)
Pileolariaceae (4)
Pucciniaceae (21)
Pucciniastraceae (10)
Pucciniosiraceae (10)
Raveneliaceae (24)
Sphaerophragmiaceae (2)
Uncolaceae (2)
Uropyxidaceae (16)
Pucciniales genera
incertae sedis (24)
Septobasidiales Septobasidiaceae (6)
Spiculogloeomycetes Spiculogloeales Spiculogloeaceae (2)
Tremellomycetes Cystofilobasidiales Cystofilobasidiaceae (1)
Mrakiaceae (7)
Filobasidiales Filobasidiaceae (5)
Piskurozymaceae (2)
Holtermanniales Holtermanniaceae (2)
Tremellales Bulleraceae (4)
Bulleribasidiaceae (6)
Carcinomycetaceae (1)
Cryptococcaceae (2)
Cuniculitremaceae (3)
Naemateliaceae (2)
Phaeotremellaceae (2)
Phragmoxenidiaceae (1)
Rhynchogastremaceae (3)
Sirobasidiaceae (1)
Tremellaceae (3)
Trimorphomycetaceae (4)
Tremellales genera
incertae sedis (8)
Trichosporonales Tetragoniomycetaceae (3)
Trichosporonaceae (8)
Tremellomycetes
genera incertae sedis
(3)
Tritirachiomycetes Tritirachiales Tritirachiaceae (2)
Ustilaginomycetes Uleiellales Uleiellaceae (1)

1086
Table 1 Continued.

Phylum Class* Order* Family*


Urocystidales Doassansiopsidaceae (1)
Fereydouniaceae (1)
Floromycetaceae (2)
Glomosporiaceae (1)
Mycosyringaceae (1)
Urocystidaceae (7)
Ustilaginales Anthracoideaceae (19)
Cintractiellaceae (1)
Clintamraceae (1)
Geminaginaceae (1)
Melanotaeniaceae (3)
Pericladiaceae (1)
Ustilaginaceae (6)
Websdaneaceae (2)
Ustilaginales genera
incertae sedis (20)
Violaceomycetales Violaceomycetaceae (1)
Ustilaginomycetes
genus incertae sedis
(1)
Wallemiomycetes Geminibasidiales Geminibasidiaceae (2)
Wallemiales Wallemiaceae (1)
Wallemiomycetes
genus incertae sedis
(1)
Basidiomycota
genera incertae
sedis (11)
Blastocladiomycot Blastocladiomycetes Blastocladiales Blastocladiaceae (3)
a
Catenariaceae (2)
Paraphysodermataceae (1)
Sorochytriaceae (1)
Blastocladiales genus
incertae sedis (1)
Callimastigales Callimastigaceae (1)
Coelomomycetaceae (2)
Catenomycetales Catenomycetaceae (1)
Blastocladiomycetes
genus incertae sedis
(1)
Physodermatomycet Physodermatales Physodermataceae (1)
es
Calcarisporiellom Calcarisporiellomyc Calcarisporiellales Calcarisporiellaceae (2)
ycota etes
Caulochytriomyco Caulochytriomycete Caulochytriales Caulochytriaceae (1)
ta s
Chytridiomycota Chytridiomycetes Chytridiales Asterophlyctaceae (2)
Chytridiaceae (6)
Chytriomycetaceae (11)
Phlyctochytriaceae (1)
Phlyctorhizaceae (1)
Pseudorhizidiaceae (1)
Scherffeliomycetaceae (1)
Zygorhizidiaceae (1)

1087
Table 1 Continued.

Phylum Class* Order* Family*


Chytridiales genus
incertae sedis (1)
Nephridiophagales Nephridiophagaceae (4)
Polyphagales Polyphagaceae (1)
Saccopodiales Saccopodiaceae (1)
Incertae sedis Amoebochytriaceae (1)
Sparrowiaceae (1)
Sphaeromonadaceae (1)
Tetrachytriaceae (1)
Thalassochytriaceae (1)
Chytridiomycetes
genera incertae sedis
(39)
Cladochytriomycetes Cladochytriales Catenochytridiaceae (1)

Cladochytriaceae (1)
Endochytriaceae (2)
Nowakowskiellaceae (1)
Septochytriaceae (1)
Cladochytriales
genera incertae sedis
(3)
Lobulomycetes Lobulomycetales Alogomycetaceae (1)
Lobulomycetaceae (4)
Lobulomycetales
genus incertae sedis
(1)
Mesochytriomycetes Gromochytriales Gromochytriaceae (1)
Mesochytriales Mesochytriaceae (1)
Polychytriomycetes Polychytriales Arkayaceae (1)
Polychytriaceae (4)
Rhizophydiomycetes Rhizophydiales Alphamycetaceae (3)
Angulomycetaceae (1)
Aquamycetaceae (1)
Batrachochytriaceae (1)
Collimycetaceae (1)
Coralloidiomycetaceae (1)
Dinomycetaceae (1)
Globomycetaceae (2)
Gorgonomycetaceae (1)
Halomycetaceae (4)
Kappamycetaceae (1)
Operculomycetaceae (1)
Pateramycetaceae (1)
Protrudomycetaceae (1)
Rhizophydiaceae (1)
Staurastromycetaceae (1)
Terramycetaceae (2)
Uebelmesseromycetaceae (1)
Rhizophydiales genus
incertae sedis (1)
Rhizophlyctidomycetes Rhizophlyctidales Arizonaphlyctidaceae (1)
Borealophlyctidaceae (1)

1088
Table 1 Continued.

Phylum Class* Order* Family*


Rhizophlyctidaceae (1)
Sonoraphlyctidaceae (1)
Spizellomycetes Spizellomycetales Powellomycetaceae (4)
Spizellomycetaceae (8)
Synchytriomycetes Synchytriales Synchytriaceae (4)
Synchytriales genus
incertae sedis (1)
Chytridiomycota
genera incertae sedis
(3)
Entomophthoromycot Entomophthoromycetes Entomophthorales Ancylistaceae (3)
a
Completoriaceae (1)
Entomophthoraceae (11)
Meristacraceae (1)
Neozygitomycetes Neozygitales Neozygitaceae (4)
Entorrhizomycota Entorrhizomycetes Entorrhizales Entorrhizaceae (1)
Talbotiomycetales Talbotiomycetaceae (1)
Glomeromycota Archaeosporomycetes Archaeosporales Ambisporaceae (1)
Archaeosporaceae (3)
Geosiphonaceae (1)
Glomeromycetes Diversisporales Acaulosporaceae (1)
Diversisporaceae (7)
Pacisporaceae (1)
Sacculosporaceae (1)
Gigasporales Dentiscutataceae (3)
Gigasporaceae (1)
Intraornatosporaceae
(2)
Racocetraceae (2)
Scutellosporaceae (3)
Glomerales Entrophosporaceae (3)
Glomeraceae (17)
Paraglomeromycetes Paraglomerales Paraglomeraceae (2)
Pervetustaceae (1)
Kickxellomycota Asellariomycetes Asellarialles Asellariaceae (1)
Asellariales genus
incertae sedis (1)
Barbatosporomycetes Barbatosporales Barbatosporaceae (1)
Dimargaritomycetes Dimargaritales Dimargaritaceae (3)
Dimargaritales
genus incertae sedis
(1)
Harpellomycetes Harpellales Harpellaceae (6)
Legeriomycetaceae (38)
Harpellales genus
incertae sedis (1)
Kickxellomycetes Kickxellales Kickxellaceae (11)
Ramicandelaberomycetes Ramicandelaberales Ramicandelaberaceae
(1)
Monoblepharomycota Hyaloraphidiomycetes Hyaloraphidiales Hyaloraphidiaceae (1)
Monoblepharidomycetes Monoblepharidales Gonapodyaceae (2)
Harpochytriaceae (1)
Monoblepharidaceae (1)

1089
Table 1 Continued.

Phylum Class* Order* Family*


Oedogoniomycetaceae
(1)
Telasphaerulaceae (1)
Sanchytriomycetes Sanchytriales Sanchytriaceae (2)
Mortierellomycota Mortierellomycetes Mortierellales Mortierellaceae (6)
Mucoromycota Endogonomycetes Endogonales Densosporaceae (1)
Endogonaceae (5)
Mucoromycetes Mucorales Backusellaceae (1)
Choanephoraceae (4)
Cunninghamellaceae (6)
Lentamycetaceae (1)
Lichtheimiaceae (9)
Mucoraceae (20)
Mycocladaceae (1)
Mycotyphaceae (1)
Phycomycetaceae (2)
Pilobolaceae (2)
Radiomycetaceae (1)
Rhizopodaceae (3)
Saksenaeaceae (2)
Syncephalastraceae (2)
Umbelopsidomycetes Umbelopsidales Umbelopsidaceae (1)
Mucoromycota genus
incertae sedis (1)
Neocallimastigomycota Neocallimastigomycetes Neocallimastigales Neocallimastigaceae
(11)
Olpidiomycota Olpidiomycetes Olpidiales Olpidiaceae (4)
Rozellomycota Rudimicrosporea Metchnikovellida Amphiacanthidae (1)
Metchnikovellidae (4)
Microsporidea Amblyosporida Amblyosporidae (17)
Caudosporidae (10)
Gurleyidae (13)
Amblyosporida
genera incertae
sedis (5)
Neopereziida Berwaldiidae (2)
Neopereziidae (6)
Tubulinosematidae (3)
Neopereziida
genera incertae
sedis (2)
Ovavesiculida Ovavesiculidae (3)
Ovavesiculida
genus incertae
sedis (1)
Glugeida Facilisporidae (1)
Glugeidae (8)
Myosporidae (1)
Pereziidae (4)
Pleistophoridae (7)
Spragueidae (7)
Thelohaniidae (15)
Unikaryonidae (4)

1090
Table 1 Continued.

Phylum Class* Order* Family*


Glugeida genus
incertae sedis (1)
Nosematida Encephalitozoonidae (2)
Enterocytozoonidae (6)
Heterovesiculidae (1)
Mrazekiidae (8)
Nosematidae (2)
Ordosporidae (1)
Nosematida genera
incertae sedis (15)
Incertae sedis Abelsporidae (1)
Areosporiidae (1)
Burenellidae (3)
Cougourdellidae (1)
Cylindrosporidae (1)
Duboscqiidae (5)
Golbergiidae (3)
Microfilidae (1)
Neonosemoidiidae (1)
Pleistosporidiidae (1)
Pseudopleistophoridae
(2)
Striatosporidae (1)
Telomyxidae (1)
Toxoglugeidae (2)
Tuzetiidae (4)
Incertae sedis Chytridiopsidea Buxtehudidae (2)
Chytridiopsidae (5)
Hesseidae (1)
Rozellomycota genera
incertae sedis (5)
Zoopagomycota Zoopagomycetes Zoopagales Cochlonemataceae (7)
Helicocephalidaceae (4)
Piptocephalidaceae (3)
Sigmoideomycetaceae
(4)
Zoopagaceae (7)
Zoopagales genus
incertae sedis (1)
*Orders/families could be listed under different subclasses in this outline. In this table, we do not indicate
auxiliary (intermediate) taxonomic ranks.

Fossil Fungi
Fossil fungi are reported in the form of dispersed spores, mycelia, sporophores,
mycorrhizae, and are commonly observed in macerated residues prepared for palynological
studies. Although fungal remains are encountered in the sediments of all ages, their frequency
increases remarkably in the Tertiary Period. This clearly suggests that their proliferation is linked
with diversification of angiosperms. Being fragmentary in nature, fossil fungi lack characteristic
features that are diagnostic of extant taxa, hampering their classification with extant fungi. These
are, therefore, described on the basis of morphological characters only. For example, spore taxa
are based on their shape, size, symmetry, number and nature of apertures, septa and spore wall
characters. On the other hand, fossil fungi (other than spores) can be assigned to their extant
counterpart (up to order/family level).

1091
Artificial classification systems for fungal spores have been proposed by Van der Hammen
(1954), Clarke (1965) and Elsik (1968). Pirozynski & Weresub (1979) suggested the use of the
‘Saccardoan System’ for classifying fungal spore types. Kendrick & Nag Raj (1979) modified the
Saccardoan System to eliminate some of its inconsistencies. This scheme is based on shape and
number of cells and accordingly fungal spores are recognized under Amerosporae,
Didymosporae, Phragmosporae, Dictyosporae, Helicosporae, Staurosporae and Scolecosporae.
This system is followed here.
Fungal sporophores of various kinds commonly occur on the surface of leaves, stems and
flowers of vascular plants and have been extensively recorded over the world (Cookson 1947,
Dilcher 1965, Elsik 1968, Kalgutkar & Jansonius 2000, van Geel & Aptroot 2006). Some are
catathecia provided with radiating rows of mycelial cells giving an appearance of tissues arranged
in a radial fashion. Ascomata contain asci that are surrounded by or enclosed within protective
tissues and may be globose, flask-shaped or saucer-shaped open bodies. These may or may not
possess an ostiolar opening. Fossil sporophores are also placed under artificial genera. Several
workers have attempted to classify and formally describe the fossil structures (Edwards 1922,
Rosendahl 1943, Cookson 1947, Rao 1959, Dilcher 1965, Venkatachala & Kar 1969, Jain &
Gupta 1970, Elsik 1978, Pirozynski 1978). Fossil sporophores are classified on the basis of
dehiscence mode (through irregular or regular cracking pattern or by a central pore or ostiole).
Other characteristic features considered for their classification are shape and margin of the
sporophores, presence or absence of pores in individual cells and nature of the central part of the
sporophores.

Slime molds
The terms ‘slime molds’ or ‘mycetozoans’ have traditionally been used to describe motile,
unicellular terrestrial predatory phagotrophs, which are capable of forming minute to relatively
large spore-producing structures, referred to as fruiting bodies. The slime molds in which
unicellular units aggregate to form first a pseudoplasmodium and then a sporocarp are referred to
as cellular slime molds. Those organisms in which the cells remain solitary but undergo a
dramatic increase in size and the number of nuclei to form a plasmodium and then a sporocarp are
referred to as plasmodial slime molds (Martin & Alexopoulos 1969). Both groups are
polyphyletic. They represent two peculiar life strategies, which have appeared several times in
different groups of Eukaryotes (Shadwick et al. 2009, Brown & Silberman 2013). A strategy
similar to what is found in cellular slime molds also occurs in the prokaryotic myxobacteria,
which are sometimes predatory but never phagotrophic (Keane & Berlemann 2016). The major
groups of slime molds are listed in Table 2.
The slime molds were once considered as fungi, due to the presence of spore-producing
structures in their life cycle. However, as it was pointed out long ago by de Bary (1887), these
similarities relate only the dispersal biology of these groups. Slime molds do not have a fungal
form of life, being predominately phagotrophic, demonstrating active motility and lacking a cell
wall during their trophic stages. Consequently, the slime molds cannot be assigned to the kingdom
Fungi in a taxonomic sense, but their nomenclature remains governed by the International Code
of Nomenclature for algae, fungi, and plants. However, an exception is Fonticula alba (Brown et
al. 2009) which belongs to the true Fungi (Nucletmycea sensu Adl et al. 2019), being a solitary
slime mold that can be considered as a fungus in the taxonomic sense.

Eumycetozoa
Among the groups of slime molds, the Eumycetozoa (the “true” slime molds) are the most
diverse and most complex in terms of morphology. All available phylogenies support the
placement of these organisms in the supergroup Amoebozoa (Shadwick et al. 2009, Tice et al.
2016, Kang et al. 2017) (Table 2).
The name Eumycetozoa was initially proposed for three groups of slime molds – the
myxomycetes, dictyostelids and protostelids (Olive & Stoianovitch 1975). However, the latter

1092
taxon appears to be polyphyletic and includes spore-forming members of several different
branches of the Amoebozoa, including the Protosporangiida, Protostelida sensu stricto,
Fractovitellida, Cavosteliida, Centramoebia and Flabellinea (see Table 2). This has led to the
conclusion that if all the protostelids with all their non-fruiting relatives are included in the
Eumycetozoa, the latter becomes nearly synonymous with the Amoebozoa (Shadwick et al. 2009).
To preserve this widely used name, Kang et al. (2017) proposed including in the Eumycetozoa
only one group of protostelids, the Protosporangiida, which forms a monophyletic unit with
myxomycetes and dictyostelids.
In terms of botanical nomenclature, the Protosporangiida may be considered as the class
Ceratiomyxomycota (Leontyev et al. 2019). Therefore, in the classification given below, we
recognize three classes within the Eumycetozoa. These are the Dictyosteliomycetes,
Ceratiomyxomycetes and Myxomycetes, as outlined by Leontyev et al. (2019).

Table 2 Position of the cellular (C) and plasmodial (P) slime mold taxa in the classification of
Eukaryotes, according to Adl et al. (2019)

Supergroups of Eukarya Group of slime molds


Amoebozoa Evosea Eumycetozoa Dictyosteliomycetes (C)
Ceratiomyxomycetes (P)
= Protosporangiida
Myxomycetes (P)
Protosteliida Protostelium (P)
Fractovitellida Ceratiomyxella (P)
Nematostelium (P)
Schizoplasmodium (P)
Soliformovum (P)
Cavosteliida Cavostelium (P)
Schizoplasmodiopsis (P)
Tychosporium (P)
Tubulinea Euamoebida Copromyxa (C)
Discosea Centramoebia Endostelium (P)
Luapelamoeba (P)
Protosteliopsis (P)
Flabellinea Vannella (P)
Discoba Heterolobosea Tetramitia Acrasidae (C)
Obazoa Opisthokonta Nucletmycea Fonticulida (C)
Sar Alveolata Ciliata Sorogena (C)
Rhizaria Guttulinopsida Guttulinopsis (C)
Stramenopiles Sagenista Sorodiplophrys (C)

Dictyosteliomycetes
The dictyostelid cellular slime molds (also called dictyostelids) are common to sometimes
abundant inhabitants of forest soil and leaf litter (Romeralo et al. 2013), grassland soil (Rollins et
al. 2010), canopy soil (Stephenson & Landolt 1998, 2011), the soil of agricultural fields
(Stephenson & Rajguru 2010) and animal dung (Stephenson & Landolt 1992), where they feed
primarily on bacteria (Singh 1947, Cavender & Raper 1965a, b). Raper & Smith (1939) and
Sanders et al. (2017) reported that dictyostelids can feed on pathogenic bacteria, including biofilm
enmeshed bacteria produced by human and plant pathogens. Interestingly, migratory birds have
been demonstrated to serve as vectors for dictyostelids (Suthers 1985), which greatly increases
their potential for long-distance dispersal.
The first species of dictyostelid, Dictyostelium mucoroides, was isolated from horse dung
and rabbit dung by Brefeld (1869). Later, a number of additional species and three additional
genera (Acytostelium, Coenonia and Polysphondylium) were described, although one of these
(Coenonia) has not been isolated since it was first described by van Tieghem in 1884. It is
possible that this organism is not a dictyostelid. Traditionally, these four genera (if Coenonia is

1093
retained), two families, and one order were classified on the basis of the morphology of the
sorophore and the pattern of branching (Raper 1984, Hagiwara 1989). This type of traditional
morphology-based classification was used by everyone working with dictyostelids until a
phylogenetic analysis based on 18S rRNA and α-tubulin gene markers indicated that the group
needed a complete revision (Schaap et al. 2006, Romeralo et al. 2011, 2012), and the traditional
genera did not hold together. More recently, a new classification of the class was proposed by
Sheikh et al. (2018), using a single 18S rRNA gene marker. This new classification provided
additional insight into the phylogeny of dictyostelids, with 12 genera, four families, and two
orders currently being recognized (Table 3).

Ceratiomyxomycetes
This class unites protosteloid eumycetozoans, in which individual sporocarps may arise
separately on a substrate (Protosporangium and Clastostelium), or form on a common layer of
extracellular slime, which may be smooth, poroid or dissected into variously branched pillars
(Ceratiomyxa) (Shadwick et al. 2009). This group was initially described under the zoological
name Protosporangiida (Kang et al. 2017). The botanical name Ceratiomyxomycetes was
proposed by Hawksworth et al. (1983) as nom. inval. (ICN, Art. 39.1), and later validated by
Leontyev et al. (2019).

Myxomycetes
The myxomycetes (or myxogastrids, Myxogastrea) differ from the rest of the slime molds
by their capability to form large fruiting bodies with a complicated structure, which may contain
millions of spores. The traditional classification, first proposed by Massee (1892) and later
developed by Martin & Alexopoulos (1969), recognized within the myxomycetes four or five
orders (Echinosteliales, Liceales, Physarales, Stemonitales, and Trichiales) based on a number of
criteria, including the presence or absence of a capillitium and lime deposits in the fruiting bodies.
This classification received worldwide recognition and was applied even in the most recent
monographs (e.g., Poulain et al. 2011). However, the results obtained from studies of the
molecular phylogeny of myxomycetes have shown that this classification does not properly reflect
evolutionary relationships within the group (Fiore-Donno et al. 2012, 2013). Based on a
comprehensive review of all published phylogenies of myxomycete subgroups and the full-length
18S rDNA phylogeny of the entire group, a new classification of the class was recently proposed
(Leontyev et al. 2019). In this classification, myxomycetes are divided into 13 families, nine
orders, four superorders and two subclasses, the Lucisporomycetidae and the Columellomycetidae
(Table 3). An additional order for the group is proposed below.
For a very long time, scientists studying myxomycetes have had a consensus about the use
of botanical nomenclature (in its mycological version) for the myxomycetes. This nomenclature
has been used in practically all published monographs of the group, from the late 19th century
(Lister 1894) to the most recent efforts (Poulain et al. 2011). This agrees with the statement in
Preamble 8 of the International Code of Nomenclature for algae, fungi, and plants, that the ‘slime
molds’ are among the organisms for which the Code is applied (Turland et al. 2018). In contrast,
the International Code of Zoological Nomenclature does not mention slime molds,
eumycetozoans or myxomycetes (Ride et al. 1999). As indicated by Ronikier & Halamski (2018),
a transfer of myxomycetes to zoological nomenclature would cause nomenclatural chaos due to
the existence of numerous homonyms and the difference between nomenclatural starting points of
the two Codes. Such a transfer is as well not required by theoretical reasons, since myxomycetes
are neither plants or fungi, nor animals, thus none of the two existing codes can reflect their
proper position in the contemporary classification of living organisms. In order to preserve
nomenclatural stability we use herein botanical names for members of the Eumycetozoa,
corresponding to the rules of ICN. The botanical name for the family Protosporangiaceae, which
had not yet been proposed, is published below according to the requirements of ICN.

1094
Oomycota
The Oomycota are a phylum of the kingdom Straminipila which evolved fungal
characterstics – such as an osmotrophic mode of nutrition and hyphal growth – convergently to
the fungal groups of the Mycota (Beakes & Thines 2017). Thus, they are traditionally studied by
mycologists and also covered by ICN. Since the last comprehensive monographic treatment by
Dick (2001) their classification underwent significant revision, with the latest classification before
the current article being that of Beakes & Thines (2017). Based on more recent discoveries,
especially regarding the early diverging oomycete lineages, this classification is updated here.

Aims of the study


The main aim of this study is to compile all outlines of fungi and fungus-like groups,
updated with recent findings and published data. We believe that this type of compilation will be
important for scientists to have a better understanding of the limitations and the definitions of the
fungal clades. For example, the classification of basal clades of fungi is debatable without a broad
agreement (e.g. Humber 2016 vs. Spatafora et al. 2016 on Entomophthoromycota and
Glomeromycota; Karpov et al. 2014, 2017 vs Bass et al. 2018 vs Tedersoo et al. 2018 vs Adl et al.
2019 on aphelids, rozellids and microsporidia). Moreover, recent proposals of classification in
Tedersoo et al. (2018) (such as elevating lower ranks to higher ranks and demoting higher ranks to
lower ranks) might also cause disagreement and thus, need to be discussed.
Fossil fungi, which is another important area of fungal taxonomy is also included in this
study. We also include fungus-like organisms to emphasize the reasons why they are excluded
from the fungal clade. We will launch a new web page, outlineoffungi.org, which will provide an
outline down to the level of genus for true Fungi, fossil fungi, and fungus-like organisms. This
data will be important for many scientific disciplines such as genomics, medicine, plant pathology,
novel compound discovery and biotechnology (Hyde et al. 2019).
It must always be borne in mind that the classifications being proposed now are based on
only perhaps 3-8 % of the fungal species present on Earth today (Hawksworth & Lücking 2017).
Any system proposed is therefore likely to be unstable and subject to change in the light of newly
discovered Fungi or fungus-like organisms. For example, sequencing of the type species of a
genus of Gyalectaceae for the first time supported the treatment of four genera as synonyms of
Gyalecta (Lücking et al. 2019), while a recent re-analysis of Dothideomycetes following the
discovery of the new genus Tenuitholiascus (which forms foliicolous lichens in China) found that
five currently accepted orders formed a single well-supported clade (Jiang et al. 2020). The
discovery of novel fungi and the sequencing of hitherto unsequenced genera can be expected to
continually yield unexpected results which prompt a re-evaluation of which taxa merit recognition
at particular ranks.
This outline is therefore not to be treated as a definitive, but a statement of the current
situation as a basis for further discussion and in some cases future consensus. In particular, now
the IBC permits lists of names to be proposed for protected status, we hope that it will be of value
in working towards a protected list of generic names for fungi, updated from that of Kirk et al.
(2013), which can be reviewed and in due course approved through the provisions of the Code.

Materials & methods

True Fungi
To list genera and other higher taxonomic ranks into a single outline, we used Kirk et al.
(2008, 2013) Lumbsch & Huhndorf (2010), Humber (2012, 2016), Wijayawardene et al. (2012,
2017a, b, 2018a, b), Hyde et al. (2013, 2020), Benny et al. (2016), Jaklitsch et al. (2016a),
Spatafora et al. (2016), Desirò et al. (2017), Lücking et al. (2017), Begerow et al. (2018),
Kraichak et al. (2018a), Tedersoo et al. (2018), Haelewaters et al. (2019b), Species Fungorum
(2019), Mapook et al. (2020) and Catalogue of Life (http://www.catalogueoflife.org/). Index
Fungorum (2019), LIAS names (http://liasnames.lias.net/) and MycoBank

1095
(http://www.mycobank.org/) were consulted concerning supplementary information on synonyms.
We generally followed He et al. (2019) for Basidiomycota classification.

Table 3 Classes, subclasses, orders and families of the Eumycetozoa with number of genera (in
brackets)
Class Subclass Order Family
Dictyosteliomycetes Acytosteliales Acytosteliaceae (3)
Cavenderiaceae (1)
Dictyosteliales Dictyosteliaceae (2)
Raperosteliaceae (4)
Incertae sedis (1)
Incertae sedis (1)
Ceratiomyxomycetes Ceratiomyxales Ceratiomyxaceae (1)
Protosporangiidae (2)
Myxomycetes Lucisporomycetidae Cribrariales Cribrariaceae (3)
Reticulariales Reticulariaceae (6)
Liceales Liceaceae (2)
Trichiales Dianemataceae (4)
Trichiaceae (8)
Incertae sedis (4)
Columellomycetidae Echinosteliopsidales Echinosteliopsidaceae
(1)
Echinosteliales Echinosteliaceae (3)
Clastodermatales Clastodermataceae (1)
Meridermatales Meridermataceae (1)
Stemonitidales Amaurochaetaceae (7)
Stemonitidaceae (3)
Physarales Lamprodermataceae (5)
Didymiaceae (4)
Physaraceae (9)
Incertae sedis (5)

The subdivision of Rozellomycota at the order and family levels is redefined according to
the phylogenetic relationships of the respective type genera representatives. The list of genera is
updated in accordance with the acknowledged checklists (Becnel et al. 2014, Cali et al. 2017,
Sokolova et al. 2018) and recent studies. In particular, genus Kabatana is suppressed as it was
shown to be the synonym of Inodosporus (Stentiford et al. 2018). Genera allocation to families
and higher rank taxa is modified after Wijayawardene et al. (2018a, b) using molecular
phylogenetic data when available. Polyphyletic higher rank taxa are suppressed. The major clades
of Microsporidia tree of life established by Vossbrinck and co-authors (2014) are redefined as the
order-rank taxa using previously published or novel names depending upon availability of
information on the type taxa.
For the classification of Leotiomycetes, Johnston et al. (2019) and Quijada et al. (2020) are
followed in the outline because the phylogenies in these papers are based on 15 loci. Ekanayaka et
al. (2019) provided an alternative classification based on less genes, but more taxa and is included
in the discussion to encourage positive dialogue.

Fossil fungi
For the sake of clarity and convenience, the fossil fungal genera are split here into three
parts, e.g. 1. Fossil fungal spores (according to Saccardoan System); 2. Fossil fungal sporophores,
mycelia and other fungal remains; and 3. Modern fungal genera to which fossil species have been
assigned. The genera are listed in three separate tables after the outline of fungi.
The data presented here have been obtained from the literature on fossil fungi published
during last seven decades or so, briefly mentioned below. In order to include all records of fossil
fungal remains from the Indian Tertiary sediments, published till 2005, three catalogues were

1096
published (Lakhanpal et al. 1976, Saxena 1991, 2006). Besides, a monographic study was carried
out by Saxena & Tripathi (2011) with the objective to synthesize the available information on
Indian fossil fungi. This incorporates description of 152 genera and 388 species, including 15
new species and 12 new combinations, with comments wherever required. Kalgutkar & Jansonius
(2000) published a synopsis of fossil fungi and tried to streamline taxonomic status of many
fossil fungal genera and species. They described about 950 validly published species, attributed
to approximately 300 genera. They proposed twelve new genera and about 350 new
combinations. Transfers of species to more appropriate genera resulted in 31 junior homonyms,
for which they provided new names. They also validated one genus and several species. In
addition to the above monographic studies, data have been gathered from scores of publications,
containing information on fossil fungi from all parts of the globe, published in various journals
and conference proceedings.

Fungus-like organisms
The classification systems used for the Dictyosteliomycetes and Myxomycetes as presented
herein are based on the critical revisions of Sheikh et al. (2018) and Leontyev et al. (2019),
respectively. In each paper, the taxonomy of the particular group was strongly revised on the basis
of original 18S rDNA phylogenies and analyses of morphological synapomorphies.
The classification of the Oomycota follows the outline presented by Beakes & Thines
(2017), with some modifications in accordance to recent studies (Bennett et al. 2019, Buaya et al.
2017, 2019, Buaya & Thines 2020).
In this classification, we have included all genera of the Eumycetozoa accepted as valid in
the nomenclatural database of Lado (2005–2019), although some of the smaller myxomycete
genera will probably be incorporated into larger ones on the basis of phylogenetic data (Leontyev
et al. 2019). This is likely to be the case for such genera as Arcyodes, Badhamia, Collaria,
Colloderma, Cornuvia, Elaeomyxa, Metatrichia, Diacheopsis, Listerella, Oligonema and
Semimorula.
A resurrection of the forgotten order Echinosteliopsidales is proposed herein, based on the
18S rDNA phylogeny of Columellomycetidae, which includes Echinosteliopsis oligospora
together with a number of environmental sequences, obtained in three different studies (Shchepin
et al. 2019).
All authors listed contributed information and comments to this work, but the inclusion of
their names does not imply that all necessarily support all details of the outline presented. Notes
are provided for recently introduced genera as well as changes in classification (marked with an
asterisk in the outline). The authors of each note are indicated in brackets, after the notes.

Phylogenetic analyses
To build a reference phylogeny, we utilized information from four genes, 18S rRNA, 28S
rRNA, RPB1 and RPB2. We used the initial set of 18S and 28S reference sequences from 1)
James et al. (2006) and supplemented by Tedersoo et al. (2017); 2) and at least one representative
sequence of each recognized order or order-level taxon, or orphan taxon as based on Tedersoo et
al. (2018) and classification of this study. Order and family representatives were selected based on
their type status, presence of all four genes, and length of the genes. The nucleotide sequences of
all genes were aligned separately using MAFFT 7 (Katoh & Standley 2013), followed by manual
checking and editing where necessary. We took advantage of the protein alignment to remove
codon-switching indels and make decision about the gaps and removal of introns. We checked for
severe conflicts in the phylogenies of all genes and replaced or removed 10 sequences that were
obviously obtained from contaminant or misidentified taxa. The initial alignment included 441
terminal taxa, which we reduced to 433 taxa to exclude putative contaminants and taxa with ultra-
long branches such as Oedogonomyces spp.

1097
Figure 1 – Maximum likelihood phylogeny of the kingdom Fungi based on LSU, SSU, RPB1 and
RPB2 combined sequence data. Numbers above branches indicate bootstrap support. Accession
numbers of terminal taxa are indicated in appendant table.

1098
Figure 1 – Continued.

1099
Figure 1 – Continued.

1100
Figure 1 – Continued.

1101
Figure 1 – Continued.

Results

Taxonomy
Helicobolomyces cinnabarinula (Müll. Arg.) Wijayaw. & Ertz, comb. nov.
Bas. Arthonia cinnabarinula Müll. Arg., Flora, Regensburg 64: 234 (1881).
Syn. nov. Helicobolomyces lichenicola Matzer, in Grube, Matzer & Hafellner, Lichenologist 27:
28 (1995).
IF Registration Identifier: 555393

Description: Grube et al. (1995).

Redonographaceae (Lücking, Tehler & Lumbsch) Lumbsch, stat nov.


Bas.: Redonographoideae Lücking, Tehler & Lumbsch, Am. J. Bot. 100: 846 (2013)
IF Registration Identifier: 555399

Description: Lücking et al. (2013)

Amblyosporida Tokarev & Issi, ord. nov.


IF Registration Identifier: 555592
Monophyletic group represented by Amblyosporidae (type genus Amblyospora) and related taxa,
based on SSU rRNA gene phylogeny, corresponding to Clade 1 (Vossbrinck et al. 2014). Life
cycles are diverse, either monomorphic (one type of sporogony) or dimorphic (two types of
sporogony within the same or different hosts). Additional sporogonial sequences may also be
present. Parasites of aquatic insects and crustaceans with rare exceptions of terrestrial insects
(Multilamina in termites).

Order type: Amblyosporidae Weiser emend. Tokarev & Issi

Neopereziida Tokarev & Issi, ord. nov.


IF Registration Identifier: 555594
Monophyletic group represented by Neopereziidae (type genus Neoperezia) and related taxa,
based on SSU rRNA gene phylogeny, corresponding to Clade 3 (Vossbrinck et al. 2014). Life
cycles are diverse, either monomorphic (one type of sporogony) or dimorphic (two types of
sporogony within the same or different hosts). Additional sporogonial sequences may also be
present. Parasites of bryozoans, insects, crustaceans and human (Anncaliia, Tubulinosema).
Order type: Neopereziidae Voronin emend. Issi, Tokarev, Seliverstova & Voronin

1102
Ovavesiculida Tokarev & Issi, ord. nov.
IF Registration Identifier: 555610
Monophyletic group represented by Ovavesiculidae (type genus Ovavesicula) and related taxa,
based on SSU rRNA gene phylogeny, corresponding to Clade 2 (Vossbrinck et al. 2014). Life
cycles are diverse, either monomorphic (one type of sporogony) or dimorphic (two types of
sporogony within the same or different hosts). Parasites of aquatic and terrestrial insects.

Order type: Ovavesiculidae Tokarev & Issi

Protosporangiaceae Leontyev, Stephenson, Schnittler, Shchepin, Novozhilov, fam. nov.


MycoBank number: MB 833618
Typus: Protosporangium L.S. Olive & Stoian., J. Protozool. 19(4): 563 (1972)
Sporocarps stalked, arise separately on a substrate with no common structures, formed by
extracellular slime. Stalk long, delicate, strait, flexuous or bent at one or several articulations.
Apical portion of the stalk thin (Protosporangium) or inflated, banana-shaped (Clastostelium).
Spores in clusters from two, four or eight units, spherical, hemispherical or compressed to each
other to form quarter-spheres. Plasmodium colorless.

Outline of fungi

Aphelidiomycota Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,


M. Ryberg & Abarenkov
Aphelidiomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,
M. Ryberg & Abarenkov
Aphelidiales Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May, M.
Ryberg & Abarenkov
Aphelidiaceae Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May, M.
Ryberg & Abarenkov
Amoeboaphelidium Scherff. (5)
Aphelidium Zopf (7)
Paraphelidium Karpov, Moreira, López-García (2)
Pseudaphelidium Schweikert & Schnepf (1)

Ascomycota Caval.-Sm.
Pezizomycotina O.E. Erikss. & Winka
Arthoniomycetes O.E. Erikss. & Winka
Arthoniales Henssen ex D. Hawksw. & O.E. Erikss.
Andreiomycetaceae B.P. Hodk. & Lendemer
Andreiomyces B.P. Hodk. & Lendemer (2)

Arthoniaceae Reichenb. ex Reichenb.


Amazonomyces Bat. (2)
Arthonia Ach. (ca. 50 + c. 300 orphaned)
Arthothelium A. Massal. (10 + ca. 100 orphaned)
Briancoppinsia Diederich, Ertz, Lawrey & van den Boom (1)
Coniangium Fr. (4)
Coniarthonia Grube (12)
Coniocarpon DC. (6)
Crypthonia Frisch & G. Thor (16)
Cryptophaea Van den Broeck & Ertz (1)
Cryptothecia Stirt. (ca. 65)
Eremothecella Syd. & P. Syd. (8)

1103
Glomerulophoron Frisch, Ertz & G. Thor (2)
Herpothallon Tobler (ca. 50)
Inoderma (Ach.) Gray (4)
Leprantha Dufour ex Körb. (1)
Myriostigma Kremp. (7)
Pachnolepia A. Massal. (1)
Reichlingia Diederich & Scheid. (4)
Snippocia Ertz, Kukwa & Sanderson (1)*
Sporodophoron Frisch (4)
Staurospora Grube (1)
Stirtonia A.L. Sm. (ca. 25)
Tylophoron Nyl. ex Stizenb. (8 + 3 orphaned in Sporodochiolichen Aptroot & Sipman)

Chrysotrichaceae Zahlbr.
Chrysothrix Mont. (ca. 18)
Galbinothrix Frisch, G. Thor, K.H. Moon & Y. Ohmura (1)*
Melarthonis Frisch & G. Thor (1)

Lecanographaceae Ertz, Tehler, G. Thor & Frisch


Alyxoria Ach. (12)
Heterocyphelium Vain. (2)
Lecanographa Egea & Torrente (ca. 40)
Mixtoconidium Etayo (2)
Phacographa Hafellner (3)
Plectocarpon Fée (ca. 40)
Zwackhia Körb. (6)

Opegraphaceae Körb. ex Stizenb.


Combea De Not. (2)
Dictyographa Müll. Arg. (2)
Dolichocarpus R. Sant. (2)
Fouragea Trevis. (4)
Ingaderia Darb. (3)
Llimonaea Egea & Torrente (4)
Nyungwea Sérus., Eb. Fisch. & Killmann (3)
Opegrapha Ach. (= Kalaallia Alstrup & D. Hawksw.) (ca. 100 + ca. 200 orphaned)
Paraingaderia Ertz & Tehler (1)
Paralecanographa Ertz & & Tehler (1)
Paraschismatomma Ertz & Tehler (1)
Pentagenella Darb. (5)
Schizopelte Th. Fr. (4)
Sclerophyton Eschw. (ca. 15)
Sparria Ertz & Tehler (2)

Roccellaceae Chevall.
Ancistrosporella G. Thor (3)
Austrographa Sparrius, Elix & A.W. Archer (3)
Austroroccella Tehler & Ertz (1)
Chiodecton Ach. (ca. 22)
Cresponea Egea & Torrente (21)
Crocellina Tehler & Ertz (1)
Dendrographa Darb. (7)

1104
Dichosporidium Pat. (8)
Dirina Fr. (13)
Diromma Ertz & Tehler (1)
Enterodictyon Müll. Arg. (2)*
Enterographa Fée (ca. 30 and 25 orphaned)
Erythrodecton G. Thor (3)
Follmanniella Peine & Werner (1)
Gorgadesia Tav. (1)
Graphidastra (Redinger) G. Thor (4)
Gyrographa Ertz & Tehler (3)
Gyronactis Ertz & Tehler (2)
Halographis Kohlm. & Volkm.-Kohlm. (1)
Haplodina Zahlbr. (3)
Isalonactis Ertz, Tehler, Eb. Fisch., Killmann, Razafindr. & Sérus. (1)
Lecanactis Körb. (ca. 30)
Mazosia A. Massal. (27)
Neosergipea M. Cáceres, Ertz & Aptroot (3)
Ocellomma Ertz & Tehler (1)
Protoroccella Follmann ex Follmann (2)
Pseudolecanactis Zahlbr. (1)
Pseudoschismatomma Ertz & Tehler (1)
Psoronactis Ertz & Tehler (1)
Pulvinodecton Henssen & G. Thor (2)
Roccella DC. (32)
Roccellina Darb. (29 + 5 orphaned in Sigridea)
Sagenidiopsis R.W. Rogers & Hafellner (4)
Schismatomma Flot. & Körb. ex A. Massal. (10)
Sigridea Tehler (6)
Simonyella J. Steiner (1)
Sipmania Egea & Torrente (1)
Streimannia G. Thor (1)
Syncesia Taylor (ca. 25)
Tania Egea, Torrente & Sipman (2)
Vigneronia Ertz (3)

Roccellographaceae Ertz & Tehler


Dimidiographa Ertz & Tehler (3)
Fulvophyton Ertz & Tehler (6)
Roccellographa J. Steiner (4)

Arthoniales genera incertae sedis


Angiactis Aptroot & Sparrius (3)
Arthophacopsis Hafellner (1)
Bactrospora A. Massal. (35)
Bryostigma Poelt & Döbbeler (3)
Catarraphia A. Massal. (1)
Felipes Frisch & G. Thor (1)
Glyphopsis Aptroot (1)
Gossypiothallon Aptroot (1)
Helminthocarpon Fee (3)
Hormosphaeria Lév. (1)
Minksia Müll. Arg. (2)

1105
Nipholepis Syd. (1)
Paradoxomyces Matzer (1)
Perigrapha Hafellner (5)
Phacothecium Trevis. (1)
Phoebus R.C. Harris & Ladd (1)
Sporostigma Grube (1)
Synarthonia Müll. Arg. (5)
Synarthothelium Sparrius (2)
Tarbertia Dennis (1)
Trichophyma Rehm (2)
Tylophorella Vain. (1)
Wegea Aptroot & Tibell (1)

Lichenostigmatales Ertz, Diederich & Lawrey


Phaeococcomycetaceae McGinnis & Schell
Etayoa Diederich & Ertz (1)
Lichenostigma Hafellner (= Phaeosporobolus D. Hawksw. & Hafellner) (5 and 26 orphaned
species)
Phaeococcomyces de Hoog (5)

Candelariomycetes Voglmayr & Jaklitsch


Candelariomycetidae Timdal & M. Westb.
Candelariales Miądl., Lutzoni & Lumbsch
Candelariaceae Hakul.
Candelaria A. Massal. (7)
Candelariella Müll. Arg. (ca. 50)
Candelina Poelt (3)
Placomaronea Räsänen (6)

Pycnoraceae Bendiksby & Timdal


Pycnora Hafellner (3)

Coniocybomycetes M. Prieto & Wedin


Coniocybales M. Prieto & Wedin
Coniocybaceae Rchb.
Chaenotheca (Th. Fr.) Th. Fr. (ca. 25)
Sclerophora Chevall. (6)

Dothideomycetes sensu O.E. Erikss & Winka


Dothideomycetidae P.M. Kirk, P.F. Cannon, J.C. David & Stalpers ex C.L. Schoch, Spatafora,
Crous & Shoemaker
Capnodiales Woron.
Aeminiaceae J. Trovão, I. Tiago & A. Portugal
Aeminium J. Trovão, I. Tiago & A. Portugal (1)

Antennulariellaceae Woron.
Achaetobotrys Bat. & Cif. (1)
Antennulariella Woron. (1)
Eumela Syd. (4)

Capnodiaceae (Sacc.) Höhn. ex Theiss.


Capnodium Mont. (83)

1106
Chaetocapnodium Hongsanan & K.D. Hyde (1)
Conidiocarpus Woron. (= Phragmocapnias Theiss. & Syd.) (ca. 10)
Fumiglobus D.R. Reynolds & G.S. Gilbert (10)
Leptoxyphium Speg. (19)
Limaciniaseta D.R. Reynolds (1)
Readerielliopsis Crous & Decock (2)
Scoriadopsis Mend. (1)
Scorias Fr. (11)

Cladosporiaceae Chalm. & R.G. Archibald


Acroconidiella J.C. Lindq. & Alippi (5)
Cladosporium Link (237 accepted species, 631 legitimate names at species level)
Davidiellomyces Crous (2)
Graphiopsis Trail (11)
Neocladosporium J.D.P. Bezerra, Sandoval-Denis, C.M. Souza-Motta & Crous (1)
Rachicladosporium Crous, U. Braun & C.F. Hill (14)
Toxicocladosporium Crous & U. Braun (14)
Verrucocladosporium K. Schub., Aptroot & Crous (2)

Cystocoleaceae Locq. ex Lücking, B.P. Hodk. & S.D. Leav.


Cystocoleus A. Massal. (1)

Dissoconiaceae Crous & de Hoog


Dissoconium de Hoog, Oorschot & Hijwegen (5)
Globoramichloridium Y. Marín & Crous (1)
Pseudoveronaea Crous & Batzer (2)
Ramichloridium Stahel ex de Hoog (35)
Uwebraunia Crous & M.J. Wingf. (7)

Euantennariaceae Hughes & Corlett


Capnokyma S. Hughes (2)
Euantennaria Speg. (9)
Hormisciomyces Bat. & Nascim. (3)
Plokamidomyces Bat., C.A.A. Costa & Cif. (1)
Rasutoria M.E. Barr (2)
Strigopodia Bat. (2)
Trichothallus F. Stevens (2)

Extremaceae Quaedvl. & Crous (= Paradevriesiaceae Crous)*


Castanedospora G. Delgado & A.N. Mill. (1)*
Extremus Quaedvl. & Crous (2)
Paradevriesia Crous (3)
Petrophila de Hoog & Quaedvl. (1)*
Pseudoramichloridium Cheew. & Crous (3)
Saxophila Selbmann & de Hoog (1)*
Staninwardia B. Sutton (2)
Vermiconidia Egidi & Onofri (= Vermiconia Egidi & Onofri) (4)*

Johansoniaceae Doilom, Phook. & K.D. Hyde


Johansonia Sacc. (13)
Orthobellus Silva & Cavalc. (3)

1107
Metacapnodiaceae Hughes & Corlett
Capnobotrys S. Hughes (10)
Hyphosoma Syd. (6)
Metacapnodium Speg. (14)

Mycosphaerellaceae Lindau
Acervuloseptoria Crous & Jol. Roux (2)
Amycosphaerella Quaedvl. & Crous (2)
Annellosympodiella Crous & Assefa (1)
Apseudocercosporella Videira & Crous (1)
Asperisporium Maubl. (24)
Australosphaerella Videira & Crous (1)
Brunneosphaerella Crous (3)
Brunswickiella Videira & Crous (1)
Camptomeriphila Crous & M.J. Wingf. (1)
Caryophylloseptoria Verkley, Quaedvl. & Crous (4)
Catenulocercospora C. Nakash., Videira & Crous (1)
Cercoramularia Videira, H.D. Shin, C. Nakash. & Crous (1)
Cercospora Fresen. (ca. 1125)
Cercosporella Sacc. (ca. 100)
Cercosporidium Earle (ca. 10)
Chuppomyces Videira & Crous (1)
Clarohilum Videira & Crous (1)
Clypeosphaerella Guatim., R.W. Barreto & Crous (3)
Collapsimycopappus A. Hashim., Y. Harada & Kaz. Tanaka (1)
Collarispora Videira & Crous (1)
Coremiopassalora U. Braun, C. Nakash., Videira & Crous (2)
Cytostagonospora Bubák (5)
Deightonomyces Videira & Crous (1)
Devonomyces Videira & Crous (1)
Dictyosporina L.M. Abreu, R.F. Castañeda & O.L. Pereira (1)
Distocercospora N. Pons & B. Sutton (4)
Distocercosporaster Videira, H.D. Shin, C. Nakash. & Crous (1)
Distomycovellosiella U. Braun, C. Nakash., Videira & Crous (1)
Dothistroma Hulbary (5)
Epicoleosporium Videira & Crous (1)
Exopassalora Videira & Crous (1)
Exosporium Link (123)
Exutisphaerella Videira & Crous (1)
Filiella Videira & Crous (1)
Fulvia Cif. (2)*
Fusoidiella Videira & Crous (2)
Graminopassalora U. Braun, C. Nakash., Videira & Crous (1)
Hyalocercosporidium Videira & Crous (1)
Hyalozasmidium U. Braun, C. Nakash., Videira & Crous (2)
Janetia M.B. Ellis (22)
Lecanosticta Syd. (8)
Madagascaromyces U. Braun, C. Nakash., Videira & Crous (1)
Microcyclosporella J. Frank, Schroers & Crous (1)
Micronematomyces U. Braun, C. Nakash., Videira & Crous (2)
Miuraea Hara (1)
Mycodiella Crous (3)

1108
Mycosphaerelloides Videira & Crous (1)
Mycovellosiella Rangel (ca. 34)
Neoceratosperma Crous & Cheew. (6)
Neocercospora Bakhshi, Arzanlou, Babai-ahari & Crous (1)
Neocercosporidium Videira & Crous (1)
Neodeightoniella Crous & W.J. Swart (1)
Neomycosphaerella Crous (1)
Neopenidiella Quaedvl. & Crous (1)
Neophloeospora U. Braun, C. Nakash., Videira & Crous (1)
Neopseudocercospora Crous (2)
Neopseudocercosporella Videira & Crous (2)
Neoramichloridium Phook., Thambug. & K.D. Hyde (1)
Neoseptoria Quaedvl., Verkley & Crous (1)
Nothopassalora U. Braun, C. Nakash., Videira & Crous (1)
Nothopericoniella Videira & Crous (1)
Nothophaeocryptopus Videira, C. Nakash. & Crous (1)
Pachyramichloridium Videira & Crous (1)
Pallidocercospora Crous (9)
Pantospora Cif. (1)
Paracercospora Deighton (5)
Paracercosporidium Videira & Crous (2)
Paramycosphaerella Crous & Jol. Roux (17)
Paramycovellosiella Videira, H.D. Shin & Crous (1)
Parapallidocercospora Videira, Crous, U. Braun & C. Nakash. (2)
Passalora Fr. (ca. 250)
Phaeocercospora Crous (2)
Phaeophleospora Rangel (31)
Phaeoramularia Munt.-Cvetk. (ca. 10)
Phloeospora Wallr. (141)
Piricauda Bubák (31)*
Pleopassalora Videira & Crous (2)
Pleuropassalora U. Braun, C. Nakash., Videira & Crous (1)
Pluripassalora Videira & Crous (1)
Plurivorosphaerella O. Hassan & T.H. Chang (1)
Polyphialoseptoria Quaedvl., R.W. Barreto, Verkley & Crous (2)
Polythrincium Kunze (5)
Protostegia Cooke (2)
Pseudocercospora Speg. (ca. 1000)
Pseudocercosporella Deighton (127)
Pseudopericoniella Videira & Crous (1)
Pseudophaeophleospora C. Nakash., Videira & Crous (2)
Pseudozasmidium Videira & Crous (4)
Ragnhildiana Solheim (18)
Ramularia Unger (100<)
Ramulariopsis Speg. (4)
Ramulispora Miura (18)
Rhachisphaerella U. Braun, C. Nakash., Videira & Crous (1)
Rosisphaerella Videira & Crous (1)
Ruptoseptoria Quaedvl., Verkley & Crous (1)
Scolecostigmina U. Braun (23)
Septoria Sacc. (= Septocyta Petr. fide Quaedvlieg et al. 2013) (200<)
Sonderhenia H.J. Swart & J. Walker (2)

1109
Sphaerulina Sacc. (65)
Stromatoseptoria Quaedvl., Verkley & Crous (1)
Sultanimyces Videira & Crous (1)
Trochophora R.T. Moore (2)
Uwemyces Hern.-Restr., Sarria & Crous (1)
Virosphaerella Videira & Crous (3)
Xenomycosphaerella Quaedvl. & Crous (3)
Xenopassalora Crous (1)
Xenoramularia Videira, H.D. Shin & Crous (3)
Xenosonderhenia Crous (2)
Xenosonderhenioides Videira & Crous (1)
Zasmidium Fr. (=Periconiella Sacc. fide Quaedvlieg et al. 2013) (ca. 150)
Zymoseptoria Quaedvl. & Crous (8)

Neodevriesiaceae Quaedvl. & Crous


Neodevriesia Quaedvl. & Crous (21)
Tripospermum Speg. (27)

Phaeothecaceae Darveaux
Phaeotheca Sigler, Tsuneda & J.W. Carmich. (4)

Phaeothecoidiellaceae K.D. Hyde & Hongsanan (= Nowamycetaceae Crous)


Chaetothyrina Theiss. (6)
Exopassalora Videira & Crous (1)
Houjia G.Y. Sun & Crous (2)
Nowamyces Crous (2)
Phaeothecoidiella Batzer & Crous (2)
Rivilata Kohlm., Volkm.-Kohlm. & O.E. Erikss. (1)
Sporidesmajora Batzer & Crous (1)
Translucidithyrium X.Y. Zeng & K.D. Hyde (1)

Piedraiaceae Viégas ex Cif., Bat. & S. Camposa


Piedraia Fons. & Leao (2)

Racodiaceae Link
Racodium Fr. (5)

Schizothyriaceae Höhn. ex Trotter, Sacc., D. Sacc. & Traverso


Amazonotheca Bat. & H. Maia (2)*
Hexagonella F. Stevens & Guba ex F. Stevens (1)
Kerniomyces Toro (1)
Lecideopsella Höhn. (10)
Metathyriella Syd. (3)
Mycerema Bat., J.L. Bezerra & Cavalc. (1)
Myriangiella Zimm. (5)
Plochmopeltis Theiss. (5)
Schizothyrium Desm. (40)
Vonarxella Bat., J.L. Bezerra & Peres (1)

Teratosphaeriaceae Crous & U. Braun


Acidiella Hujslová & M. Kolařík (3)

1110
Acidomyces B.J. Baker, M.A. Lutz, S.C. Dawson, P.L. Bond & Banfield ex Selbmann, de
Hoog & De Leo (2)
Acrodontium de Hoog (17)
Apenidiella Quaedvl. & Crous (1)
Araucasphaeria Crous & M.J. Wingf. (1)
Aulographina Arx & E. Müll. (2)
Austroafricana Quaedvl. & Crous (3)
Austrostigmidium Pérez-Ort. & Garrido-Benavent (1)
Batcheloromyces Marasas, P.S. van Wyk & Knox-Dav. (5)
Baudoinia J.A. Scott & Unter. (5)
Bryochiton Döbbeler & Poelt (5)
Camarosporula Petr. (1)
Capnobotryella Sugiy. (6)
Catenulostroma Crous & U. Braun (7)
Constantinomyces Egidi & Onofri (4)
Davisoniella H.J. Swart (1)
Devriesia Seifert & N.L. Nick. (11)
Elasticomyces Zucconi & Selbmann (1)
Eupenidiella Quaedvl. & Crous (1)
Euteratosphaeria Quaedvl. & Crous (1)
Friedmanniomyces Onofri (2)
Hispidoconidioma Tsuneda & Davey (2)
Hortaea Nishim. & Miyaji (2)*
Hyweljonesia R.G. Shivas, Y.P. Tan, Marney & Abell (2)
Incertomyces Egidi & Zucconi (2)
Lapidomyces de Hoog & Stielow (1)
Leptomelanconium Petr. (7)
Meristemomyces Isola & Onofri (2)
Microcyclospora J. Frank, Schroers & Crous (5)
Monticola Selbmann & Egidi (1)
Myrtapenidiella Quaedvl. & Crous (8)
Neocatenulostroma Quaedvl. & Crous (3)
Neophaeothecoidea Quaedvl. & Crous (1)
Neotrimmatostroma Quaedvl. & Crous (3)
Oleoguttula Selbmann & de Hoog (1)
Pachysacca Syd. (3)
Parapenidiella Crous & Summerell (2)
Parateratosphaeria Quaedvl. & Crous (6)
Penidiella Crous & U. Braun (4)
Penidiellomyces Crous, Attili-Angelis, A.P.M. Duarte, Pagnocca & J.Z. Groenew. (2)
Penidiellopsis Sand.-Den., Gené, Deanna A. Sutton & Guarro (2)
Phaeothecoidea Crous (5)
Placocrea Syd. (1)*
Pseudotaeniolina J.L. Crane & Schokn. (2)
Pseudoteratosphaeria Quaedvl. & Crous (6)
Queenslandipenidiella Quaedvl. & Crous (1)
Readeriella Syd. & P. Syd. (ca. 23)
Recurvomyces Selbmann & de Hoog (1)
Simplicidiella Crous, Attili-Angelis, A.P.M. Duarte, Pagnocca & J.Z. Groenew. (1)
Stenella Syd. (ca. 45)
Suberoteratosphaeria Quaedvl. & Crous (3)
Teratoramularia Videira, H.D. Shin & Crous (4)

1111
Teratosphaeria Syd. & P. Syd. (58)
Teratosphaericola Quaedvl. & Crous (1)
Teratosphaeriopsis Quaedvl. & Crous (1)
Xanthoriicola D. Hawksw. (1)
Xenoconiothyrium Crous & Marinc. (1)
Xenopenidiella Quaedvl. & Crous (7)
Xenophacidiella Crous (1)
Xenoteratosphaeria Quaedvl. & Crous (1)

Xenodevriesiaceae Crous
Xenodevriesia Crous (1)

Capnodiales genera incertae sedis


Arthrocatena Egidi & Selbmann (1)
Catenulomyces Egidi & de Hoog (1)
Eriosporella Höhn. (2)
Hyphoconis Egidi & Quaedvl. (1)
Mucomycosphaerella Quaedvl. & Crous (1)
Mycophycias Kohlm. & Volkm.-Kohlm
Neohortaea Quaedvl. & Crous (1)
Perusta Egidi & Stielow (1)
Plurispermiopsis Pereira-Carv., Inácio & Dianese (1)
Pseudoepicoccum M.B. Ellis (4)
Racoleus R. Sant. & D. Hawksw. (1)
Ramimonilia Stielow & Quaedvl. (1)
Ramopenidiella Crous & R.G. Shivas (1)
Rosaria N. Carter (2)

Dothideales Lindau (= Neocelosporiales Crous)


Dothideaceae Chevall.
Delphinella (Sacc.) Kuntze (7)
Dictyodothis Theiss. & Syd. (8)
Dothidea Fr. (ca. 20)
Dothiora Fr. (50<)
Endoconidioma Tsuneda (2)
Endodothiora Petr. (1)
Kabatina R. Schneid. & Arx (5)
Neocylindroseptoria Thambug. & K.D. Hyde (1)
Phaeocryptopus Naumov (6)
Plowrightia Sacc. (50)
Stylodothis Arx & E. Müll. (2)
Sydowia Bres. (11)
Uleodothis Theiss. & Syd. (4)

Neocelosporiaceae Crous
Celosporium Tsuneda & M.L. Davey (1)
Muellerites L. Holm (1)
Neocelosporium Crous (1)*

Saccotheciaceae Bonord.
Aureobasidium Viala & G. Boyer (23)
Columnosphaeria Munk (4)

1112
Kabatiella Bubák (19)
Pseudoseptoria Speg. (8)
Pseudosydowia Thambug. & K.D. Hyde (1)
Saccothecium Fr. (9)
Selenophoma Maire (ca. 13)

Zalariaceae Visagie, Z. Humphries & Seifert


Zalaria Visagie, Z. Humphries & Seifert (2)

Dothideales genera incertae sedis


Asteromellopsis H.E. Hess & E. Müll. (1)
Botryochora Torrend (1)
Coniozyma Crous (1)
Hormonema Lagerb. & Melin (7)
Pringsheimia Schulzer (17)
Rhizosphaera L. Mangin & Har. (8)

Myriangiales Starbäck
Elsinoaceae Höhn. ex Sacc. & Trotter
Elsinoë Racib. (ca. 40)
Molleriella G. Winter (4)

Myriangiaceae Nyl.
Anhellia Racib. (9)
Ascostratum Syd. & P. Syd. (2)
Butleria Sacc. (1)
Dictyocyclus Sivan., W.H. Hsieh & Chi Y. Chen (1)
Eurytheca De Seynes (3)
Hemimyriangium J. Reid & Piroz (1)
Mendogia Racib. (7)
Micularia Boedijn (2)
Myriangium Mont. & Berk. (ca. 10)
Uleomyces P. Henn. (12)
Zukaliopsis Henn. (2)

Myriangiales genus incertae sedis


Dictyonella Höhn. (7)

Pleosporomycetidae C.L. Schoch, Spatafora, Crous & Shoemaker


Gloniales Jayasiri & K.D. Hyde*
Gloniaceae (Corda) E. Boehm, C.L. Schoch & Spatafora
Cenococcum Moug. & Fr. (5)
Glonium Mühl. (ca. 13)
Purpurepithecium Jayasiri & K.D. Hyde (2)

Hysteriales Lindau
Hysteriaceae Chevall.
Actidiographium Lar. N. Vassiljeva (1)
Gloniella Sacc. (12)
Gloniopsis De Not. (ca. 17)
Hysterium Pers. (14)
Hysterobrevium E. Boehm & C.L. Schoch (6)

1113
Hysterocarina Zogg (1)
Hysterodifractum D.A.C. Almeida, Gusmão & A.N. Mill. (1)
Hysteroglonium Rehm ex Lindau (2)
Oedohysterium E. Boehm & C.L. Schoch (3)
Ostreichnion Duby (4)
Pseudoscypha J. Reid & Piroz. (1)
Psiloglonium Höhn. (ca. 15)
Rhytidhysteron Speg. (21)

Hysteriales genus incertae sedis


Graphyllium Clem. (11)

Mytilinidiales E. Boehm, C.L. Schoch & Spatafora


Mytilinidiaceae Kirschst.
Actidium Fr. (ca. 6)
Lophium Fr. (ca. 6)
Mytilinidion Duby (12)
Ostreola Darker (8)
Peyronelia Cif. & Gonz. Frag. (6 or 7)
Pseudocamaropycnis Crous (1)
Quasiconcha M.E. Barr & M. Blackw. (1)
Septonema Corda (ca. 15)
Zoggium Lar.N. Vassiljeva (1)

Pleosporales Luttrell ex M.E. Barr


Acrocalymmaceae Crous & Trakun.
Acrocalymma Alcorn & J.A.G. Irwin (6)

Aigialaceae Suetrong, Sakay., E.B.G. Jones, Kohlm., Volkm.-Kohlm. & C.L. Schoch
Aigialus S. Schatz & Kohlm. (5)
Ascocratera Kohlm. (1)
Fissuroma Jian K. Liu, Phook., E.B.G. Jones & K.D. Hyde (11)
Neoastrosphaeriella Jian K. Liu, E.B.G. Jones & K.D. Hyde (3)
Posidoniomyces Vohník & Réblová (1)
Rimora Kohlm., Volkm.-Kohlm., Suetrong, Sakay. & E.B.G. Jones (1)

Amniculicolaceae Y. Zhang ter, C.L. Schoch, J. Fourn., Crous & K.D. Hyde
Amniculicola Y. Zhang ter & K.D. Hyde (4)
Fusiformispora Phukhams. & K.D. Hyde (1)
Murispora Y. Zhang ter, J. Fourn. & K.D. Hyde (7)
Neomassariosphaeria Y. Zhang ter, J. Fourn. & K.D. Hyde (1)
Pseudomassariosphaeria Phukhams., Ariyaw., Camporesi & K.D. Hyde (2)
Vargamyces Tóth (1)

Amorosiaceae Thambug. & K.D. Hyde


Alfoldia D.G. Knapp, Imrefi & Kovács (1)
Amorosia Mantle & D. Hawksw. (1)
Amorocoelophoma Jayasiri, E.B.G. Jones & K.D. Hyde (1)
Angustimassarina Thambug., Kaz. Tanaka & K.D. Hyde (11)

Anteagloniaceae K.D. Hyde, Jian K. Liu & A. Mapook


Anteaglonium Mugambi & Huhndorf (7)

1114
Flammeascoma Phook. & K.D. Hyde (2)
Purplema W. Dong, H. Zhang & K.D. Hyde (1)

Aquasubmersaceae A. Hashim. & Kaz. Tanaka


Aquasubmersa K.D. Hyde & Huang Zhang (2)

Arthopyreniaceae W. Watson
Arthopyrenia A. Massal. (= Arthopyreniomyces Cif. & Tomas.) (5 + ca. 100 orphaned)
Mycomicrothelia Keissl. (ca. 10)*

Ascocylindricaceae Abdel-Wahab, Bahkali, E.B.G. Jones, Ariyaw. & K.D. Hyde


Ascocylindrica Abdel-Wahab, Bahkali & E.B.G. Jones (1)

Astrosphaeriellaceae Phook. & K.D. Hyde


Astrosphaeriella Syd. & P. Syd. (ca. 10)
Astrosphaeriellopsis Phook., Jian K. Liu & K.D. Hyde (2)
Javaria Boise (2)
Mycopepon Boise (5)
Pithomyces Berk. & Broome (ca. 40)
Pteridiospora Penz. & Sacc. (8)
Quercicola Jayasiri, E.B.G. Jones & K.D. Hyde (2)
Xenoastrosphaeriella Jayasiri, E.B.G. Jones & K.D. Hyde (1)

Bambusicolaceae D.Q. Dai & K.D. Hyde


Bambusicola D.Q. Dai & K.D. Hyde (10)
Leucaenicola Jayasiri, E.B.G. Jones & K.D. Hyde (2)
Palmiascoma Phook. & K.D. Hyde (1)

Biatriosporaceae K.D. Hyde


Biatriospora K.D. Hyde & Borse (6)

Camarosporiaceae Wanas., Wijayaw., K.D. Hyde & Crous


Camarosporium Schulzer (100+)
Camarosporomyces Crous (1)

Camarosporidiellaceae Wanas., Wijayaw., Crous & K.D. Hyde


Camarosporidiella Wanas., Wijayaw., K.D. Hyde (22)

Caryosporaceae Huang Zhang, K.D. Hyde & Ariyaw.


Caryospora De Not. (19)

Coniothyriaceae W.B. Cooke


Coniothyrium Corda (ca. 50)
Foliophoma Crous (2)
Neoconiothyrium Crous (3)
Ochrocladosporium Crous & U. Braun (3)
Staurosphaeria Rabenh. (= Hazslinszkyomyces Crous & R.K. Schumach.) (12)

Corynesporascaceae Sivan.
Corynespora Güssow (ca. 130)
Corynesporasca Sivan. (1)

1115
Cryptocoryneaceae A. Hashim. & Kaz. Tanaka
Cryptocoryneum Fuckel (ca. 20)

Cucurbitariaceae G. Winter (= Fenestellaceae M.E. Barr)


Allocucurbitaria Valenz.-Lopez, Stchigel, Guarro & Cano (1)
Astragalicola Jaklitsch & Voglmayr (2)*
Cucitella Jaklitsch & Voglmayr (1)*
Cucurbitaria Gray (= Pleurostromella Petr.) (ca. 40)
Fenestella Tul. & C. Tul. (ca. 4)
Neocucurbitaria Wanas., E.B.G. Jones & K.D. Hyde (21)
Paracucurbitaria Valenz.-Lopez, Stchigel, Guarro & Cano (2)
Parafenestella Jaklitsch & Voglmayr (3)*
Protofenestella Jaklitsch & Voglmayr (1)*
Rhytidiella Zalasky (4)
Seltsamia Jaklitsch & Voglmayr (1)*
Syncarpella Theiss. & Syd. (ca. 6)
Synfenestella Jaklitsch & Voglmayr (2)

Cyclothyriellaceae Jaklitsch & Voglmayr


Cyclothyriella Jaklitsch & Voglmayr (1)
Massariosphaeria (E. Müll.) Crivelli (25)*

Dacampiaceae Körb.
Aaosphaeria Aptroot (1)
Dacampia A. Massal. (15)
Eopyrenula R.C. Harris (6)
Leptocucurthis Aptroot (1)
Pseudonitschkia Coppins & S.Y. Kondr. (1)
Weddellomyces D. Hawksw. (12)

Delitschiaceae M.E. Barr


Delitschia Auersw. (ca. 50)
Ohleriella Earle (1)
Semidelitschia Cain & Luck-Allen (3)

Diademaceae Shoemaker & C.E. Babc.


Diadema Shoemaker & C.E. Babc. (8)

Dictyosporiaceae Boonmee & K.D. Hyde


Aquadictyospora Z.L. Luo, K.D. Hyde & H.Y. Su (1)
Aquaticheirospora Kodsueb & W.H. Ho (1)
Cheirosporium L. Cai & K.D. Hyde (2)
Dendryphiella Bubák & Ranoj. (12)
Dictyocheirospora M.J. D'souza, Boonmee & K.D. Hyde (16)
Dictyopalmispora Pinruan, Boonmee & K.D. Hyde (1)
Dictyosporium Corda (59)
Digitodesmium P.M. Kirk (6)
Gregarithecium Kaz. Tanaka & K. Hiray. (1)
Jalapriya M.J. D'souza, Hong Y. Su, Z.L. Luo & K.D. Hyde (3)
Neodendryphiella Iturrieta-González, Dania García & Gené (3)*
Pseudocoleophoma Kaz. Tanaka & K. Hiray. (3)
Pseudoconiothyrium Crous & R.K. Schumach. (1)

1116
Pseudodictyosporium Matsush. (4)
Vikalpa M.J. D'souza, Boonmee, Bhat & K.D. Hyde (4)

Didymellaceae Gruyter, Aveskamp & Verkley (=Microsphaeropsidaceae Qian Chen, L. Cai &
Crous fide Hongsanan et al. 2020)
Allophoma Q. Chen & L. Cai (9)
Anthodidymella Phukhams., Camporesi & K.D. Hyde (3)
Ascochyta Lib. (= Heracleicola Tibpromma, Camporesi & K.D. Hyde) (ca. 400)
Boeremia Aveskamp, Gruyter & Verkley (22)
Briansuttonomyces Crous (1)
Calophoma Q. Chen & L. Cai (8)
Chaetasbolisia Speg. (7)
Cumuliphoma Valenz.-Lopez, Stchigel, Crous, Guarro & Cano (3)
Didymella Sacc. ex D. Sacc. (ca. 100)
Didysimulans Tibpromma, Camporesi & K.D. Hyde (2)
Ectophoma Valenz.-Lopez, Cano, Crous, Guarro & Stchigel (2)
Epicoccum Link (16)
Heterophoma Q. Chen & L. Cai (6)
Juxtiphoma Valenzuela-Lopez, Cano, Crous, Guarro & Stchigel (1)
Leptosphaerulina McAlpine (30)
Macroventuria Aa (2)
Microsphaeropsis Höhn. (37)
Mixtura O.E. Erikss. & J.Z. Yue (1)
Monascostroma Höhn. (ca. 5)
Neoascochyta Q. Chen & L. Cai (12)
Neodidymella Phook., R.H. Perera & K.D. Hyde (1)
Neodidymelliopsis Q. Chen & L. Cai (10)
Neomicrosphaeropsis Thambug., Camporesi & K.D. Hyde (10)
Nothophoma Q. Chen & L. Cai (9)
Paraboeremia Q. Chen & L. Cai (6)
Phoma Sacc. (= Endophoma Tsuneda & M.L. Davey) (100)
Phomatodes Q. Chen & L. Cai (2)
Platychora Petr. (1)
Remotididymella Valenz.-Lopez (2)
Similiphoma Valenz.-Lopez, Crous, Cano, Guarro & Stchigel (1)
Stagonosporopsis Died. (22)
Vacuiphoma Valenz.-Lopez, Cano, Crous, Guarro & Stchigel (2)
Xenodidymella Q. Chen & L. Cai (6)

Didymosphaeriaceae Munk
Alloconiothyrium Verkley & Stielow (1)
Austropleospora R.G. Shivas & L. Morin (1)
Barria Z.Q. Yuan (1)
Bimuria D. Hawksw., Chea & Sheridan (1)
Chromolaenicola Mapook & K.D. Hyde (5)
Curreya Sacc. (2)
Cylindroaseptospora Jayasiri, E.B.G. Jones & K.D. Hyde (2)
Deniquelata Ariyaw. & K.D. Hyde (2)
Didymocrea Kowalski (1)
Didymosphaeria Fuckel (ca. 25)
Julella Fabre (ca. 20)
Kalmusia Niessl (16)

1117
Kalmusibambusa Phook., Tennakoon, Thambug. & K.D. Hyde (1)*
Karstenula Speg. (16)
Laburnicola Wanas., Camporesi, E.B.G. Jones & K.D. Hyde (4)
Letendraea Sacc. (ca. 3)
Lineostroma H.J. Swart (1)
Montagnula Berl. (ca. 30)*
Neokalmusia Ariyaw. & K.D. Hyde (6)
Neptunomyces M. Gonçalves, T. Vicente & A. Alves (1)*
Paracamarosporium Wijayaw. & K.D. Hyde (7)
Paraconiothyrium Verkley (19)
Paramassariosphaeria Wanas., E.B.G. Jones & K.D. Hyde (2)
Paraphaeosphaeria O.E. Erikss. (33)
Phaeodothis Syd. & P. Syd. (5)
Pseudocamarosporium Wijayaw. & K.D. Hyde (14)
Pseudopithomyces Ariyaw. & K.D. Hyde (10)
Spegazzinia Sacc. (11)*
Tremateia Kohlm., Volkm.-Kohlm. & O.E. Erikss. (5)
Verrucoconiothyrium Crous (4)
Vicosamyces Firmino, Machado & Pereira (1)
Xenocamarosporium Crous & M.J. Wingf. (1)

Dothidotthiaceae Crous & A.J.L. Phillips


Belizeana Kohlm. & Volkm. (1)
Dothidotthia Höhn. (= Neodothidotthia Crous) (ca. 10)
Mycocentrospora Deighton (4)
Phaeomycocentrospora Crous, H.D. Shin & U. Braun (1)
Pleiochaeta (Sacc.) S. Hughes (4)
Thyrostroma Höhn. (ca. 45)
Wilsonomyces Adask., J.M. Ogawa & E.E. Butler (1)

Fuscostagonosporaceae Jayasiri, Camporesi & K.D. Hyde


Fuscostagonospora Kaz. Tanaka & K. Hiray. (3)

Fusculinaceae Crous
Fusculina Crous & Summerell (3)
Gordonomyces Crous & Marinc. (1)

Halojulellaceae Suetrong, K.D. Hyde & E.B.G. Jones


Halojulella Suetrong, K.D. Hyde & E.B.G. Jones (1)

Halotthiaceae Ying Zhang, J. Fourn. & K.D. Hyde


Brunneoclavispora Phook. & K.D. Hyde (1)
Halotthia Kohlm. (1)
Mauritiana Poonyth, K.D. Hyde, Aptroot & Peerally (1)
Neolophiostoma S. Boonmee & K.D. Hyde (1)*
Pontoporeia Kohlm. (1)
Sulcosporium Phook. & K.D. Hyde (1)

Hermatomycetaceae Locq.
Hermatomyces Speg. (ca. 20)

1118
Hypsostromataceae Huhndorf
Hypsostroma Huhndorf (2)
Latoruaceae Crous
Latorua Crous (1)
Matsushimamyces Rahul Sharma & Rohit Sharma (2)
Polyschema H.P. Upadhyay (22)
Pseudoasteromassaria M. Matsum. & Kaz. Tanaka (2)

Lentimurisporaceae N.G. Liu, J.K Liu & K.D. Hyde


Bahusandhika Subram. (9)*
Lentimurispora N.G. Liu, Bhat & K.D. Hyde (1)*

Lentitheciaceae Y. Zhang ter, C.L. Schoch, J. Fourn., Crous & K.D. Hyde
Darksidea D.G. Knapp, Kovács, J.Z. Groenew. & Crous (6)
Halobyssothecium Dayar., E.B.G. Jones & K.D. Hyde (1)
Katumotoa Kaz. Tanaka & Y. Harada (1)
Keissleriella Höhn. (ca. 36)
Lentithecium K.D. Hyde, J. Fourn. & Ying Zhang (9)*
Murilentithecium Wanas., Camporesi, E.B.G. Jones & K.D. Hyde (3)
Neoophiosphaerella Kaz. Tanaka & K. Hiray. (1)
Phragmocamarosporium Wijayaw., Yong Wang bis & K.D. Hyde (2)
Pleurophoma Höhn. (ca. 9)
Poaceascoma Phook. & K.D. Hyde (4)
Pseudomurilentithecium Mapook & K.D. Hyde (1)
Setoseptoria Quaedvl., Verkley & Crous (7)
Tingoldiago K. Hiray. & Kaz. Tanaka (1)
Towyspora Wanas., E.B.G. Jones & K.D. Hyde (1)

Leptosphaeriaceae M.E. Barr


Alloleptosphaeria Ariyaw., Wanas. & K.D. Hyde (1)
Alternariaster E.G. Simmons (4)
Chaetoplea (Sacc.) Clem. (ca. 20)
Heterosporicola Crous (2)
Leptosphaeria Ces. & De Not. (151)
Neoleptosphaeria Ariyaw. & K.D. Hyde (2)
Paraleptosphaeria Gruyter, Aveskamp & Verkley (7)
Plenodomus Preuss (19)
Pseudoleptosphaeria Ariyaw. & K.D. Hyde (1)
Querciphoma Crous (2)
Sclerenchymomyces Phukhams. & K.D. Hyde (2)
Sphaerellopsis Cooke (6)
Subplenodomus Gruyter, Aveskamp & Verkley (6)

Libertasomycetaceae Crous
Libertasomyces Crous & Roets (3)
Neoplatysporoides Crous & M.J. Wingf. (1)

Ligninsphaeriaceae K.D. Hyde & Ariyaw. (Nom. inval., Art. 38.1(a) (Melbourne) fide Index
Fungorum 2020)
Ligninsphaeria Jin F. Zhang, Jian K. Liu, K.D. Hyde & Zi Y. Liu (1)
Ligninsphaeriopsis Phukh., Feng & K.D. Hyde (1)

1119
Lindgomycetaceae K. Hiray., Kaz. Tanaka & Shearer
Arundellina Wanas., E.B.G. Jones & K.D. Hyde (1)
Clohesyomyces K.D. Hyde (1)
Hongkongmyces C.C.C. Tsang, J.F.W. Chan, Trend.-Sm., A.H.Y. Ngan, I.W.H. Ling,
S.K.P. Lau & P.C.Y. Woo (3)
Lindgomassariosphaeria W. Dong, H. Zhang & K.D. Hyde (1)
Lindgomyces K. Hiray., Kaz. Tanaka & Shearer (13)
Lolia Abdel-Aziz & Abdel-Wahab (1)
Neolindgomyces Jayasiri, E.B.G. Jones & K.D. Hyde (4)

Lizoniaceae Boonmee & K.D. Hyde


Lizonia (Ces. & De Not.) De Not. (22)

Longipedicellataceae Phukhams., Bhat & K.D. Hyde


Longipedicellata H. Zhang, K.D. Hyde & Jian K. Liu (1)
Pseudoxylomyces Kaz. Tanaka & K. Hiray. (1)
Submerspora W. Dong, H. Zhang & K.D. Hyde (1)

Longiostiolaceae Phukhams., Doilom & K.D. Hyde


Longiostiolum Doilom, Ariyaw. & K.D. Hyde (1)

Lophiostomataceae Sacc.
Alpestrisphaeria Thambug. & K.D. Hyde (2)
Biappendiculispora Thambug., Kaz. Tanaka & K.D. Hyde (1)
Capulatispora Thambug. & K.D. Hyde (1)
Coelodictyosporium Thambug. & K.D. Hyde (3)
Crassiclypeus A. Hashim., K. Hiray. & Kaz. Tanaka (1)*
Decaisnella Fabre (13)
Dimorphiopsis Crous (1)
Flabellascoma A. Hashim., K. Hiray. & Kaz. Tanaka (2)*
Guttulispora Thambug., Qing Tian & K.D. Hyde (1)
Kiskunsagia D.G. Knapp, Imrefi & Kovács (1)
Lentistoma A. Hashim., K. Hiray. & Kaz. Tanaka (1)
Leptoparies A. Hashim., K. Hiray. & Kaz. Tanaka (1)*
Lophiohelichrysum Dayar., Camporesi & K.D. Hyde (1)
Lophiopoacea Ariyaw., Thambug. & K.D. Hyde (2)
Lophiostoma Ces. & De Not. (ca. 100)
Neopaucispora Wanas., Gafforov & K.D. Hyde (1)
Neotrematosphaeria Thambug., Kaz. Tanaka & K.D. Hyde (1)
Neovaginatispora A. Hashim., K. Hiray. & Kaz. Tanaka (1)
Parapaucispora A. Hashim., K. Hiray. & Kaz. Tanaka (1)
Paucispora Thambug., Kaz. Tanaka & K.D. Hyde (3)
Platystomum Trevis. (ca. 20)
Pseudocapulatispora Mapook & K.D. Hyde (inpress)
Pseudolophiostoma Thambug., Kaz. Tanaka & K.D. Hyde (5)
Pseudopaucispora A. Hashim., K. Hiray. & Kaz. Tanaka (1)*
Pseudoplatystomum Thambug. & K.D. Hyde (1)
Quintaria Kohlm. & Volkm.-Kohlm (3)
Sigarispora Thambug. & K.D. Hyde (14)
Vaginatispora K.D. Hyde (8)

1120
Lophiotremataceae K. Hiray. & Kaz.
Atrocalyx A. Hashim. & Kaz. Tanaka (6)
Crassimassarina A. Hashim. & Kaz. Tanaka (1)
Cryptoclypeus A. Hashim. & Kaz. Tanaka (2)
Galeaticarpa A. Hashim. & Kaz. Tanaka (1)
Koordersiella Höhn. (6)
Lophiotrema Sacc. (17)
Pseudocryptoclypeus A. Hashim. & Kaz. Tanaka (1)

Macrodiplodiopsidaceae Voglmayr, Jaklitsch & Crous


Macrodiplodiopsis Petr. (2)
Pseudochaetosphaeronema Punith. (4)

Massariaceae Nitschke
Massaria De Not. (31)
Massarioramusculicola Huanral., Thambug. & K.D. Hyde (1)
Paramassaria Samarak. & K.D. Hyde (1)

Massarinaceae Munk
Byssothecium Fuckel (8)
Helminthosporium Link (= Helminthosporiella Hern.-Restr., G.A. Sarria & Crous) (ca. 416)
Massarina Sacc. (ca. 100)
Pseudodidymosphaeria Thambug. & K.D. Hyde (2)
Pseudosplanchnonema Chethana & K.D. Hyde (1)
Semifissispora H.J. Swart (5)
Stagonospora (Sacc.) Sacc. (220)
Suttonomyces Wijayaw., Camporesi & K.D. Hyde (2)

Melanommataceae G. Winter (= Pseudodidymellaceae A. Hashim. & Kaz. Tanaka)


Alpinaria Jaklitsch & Voglmayr (1)
Aposphaeria Sacc. (189)
Asymmetricospora J. Fröhl. & K.D. Hyde (1)
Bertiella (Sacc.) Sacc. & P. Syd. (2)
Bicrouania Kohlm. & Volkm.-Kohlm. (1)
Byssosphaeria Cooke (27)
Calyptronectria Speg. (3)
Camposporium Harkn. (21)*
Exosporiella P. Karst. (1)
Fusiconidium Jun F. Li, Phook. & K.D. Hyde (2)
Herpotrichia Fuckel (101)
Mamillisphaeria K.D. Hyde, S.W. Wong & E.B.G. Jones (1)
Marjia Wanas., Gafforov & K.D. Hyde (1)
Melanocamarosporioides D. Pem, R. Jeewon, Gafforov & K.D. Hyde (1)
Melanocamarosporium Wijayaw., Camporesi, Bhat & K.D. Hyde (2)
Melanocucurbitaria Wanas., Gafforov & K.D. Hyde (1)
Melanodiplodia Wanas., Gafforov & K.D. Hyde (1)
Melanomma Nitschke ex Fuckel (ca. 30)
Monoseptella Wanas., Gafforov & K.D. Hyde (1)
Muriformistrickeria Q. Tian, Wanas., Camporesi & K.D. Hyde (2)
Navicella Fabre (5)
Neobyssosphaeria Wanas., Jones & K.D. Hyde (1)
Petrakia Syd. & P. Syd. (6)

1121
Phragmocephala E.W. Mason & S. Hughes (10)
Phragmotrichum Kunze (4)
Pleotrichocladium Hern.-Restr., R.F. Castañeda & Gené (1)
Praetumpfia Jaklitsch & Voglmayr (1)
Pseudobyssosphaeria H.B. Jiang & K.D. Hyde (1)
Pseudodidymella C.Z. Wei, Y. Harada & Katum. (2)
Pseudostrickeria Q. Tian, Wanas., Camporesi & K.D. Hyde (3)
Sarimanas M. Matsum., K. Hiray. & Kaz. Tanaka (2)
Seifertia Partr. & Morgan-Jones (2)
Tumularia Descals & Marvanová (2)
Uzbekistanica Wanas., Gafforov & K.D. Hyde (3)
Xenostigmina Crous (2)

Morosphaeriaceae Suetrong, Sakay., E.B.G. Jones & C.L. Schoch


Aquihelicascus W. Dong, H. Zhang & Doilom (3)
Aquilomyces D.G. Knapp, Kovács, J.Z. Groenew. & Crous (2)
Clypeoloculus Kaz. Tanaka & K. Hiray. (4)
Helicascus Kohlm. (12)
Morosphaeria Suetrong, Sakay., E.B.G. Jones & C.L. Schoch (4)
Neohelicascus W. Dong, H. Zhang & Doilom (8)

Mycoporaceae Zahlbr.
Mycoporum Flot. ex Nyl. (ca. 5 + c. 35 orphaned, partly in Mycoporellum Müll. Arg.)

Neocamarosporiaceae Wanas., Wijayaw., Crous & K.D. Hyde


Dimorphosporicola Crous (1)
Neocamarosporium Crous & M.J. Wingf. (15)

Neohendersoniaceae Giraldo & Crous


Brevicollum Kaz. Tanaka (2)
Crassiparies M. Matsum., K. Hiray. & Kaz. Tanaka (1)
Medicopsis Gruyter, Verkley & Crous (2)
Neohendersonia Petr. (4)
Neomedicopsis Crous & Akulov (1)

Neomassariaceae H.A. Ariyaw., Jaklitsch & Voglmayr


Neomassaria Mapook, Camporesi & K.D. Hyde (2)

Neomassarinaceae Mapook & K.D. Hyde


Neomassarina Phook., Jayasiri & K.D. Hyde (2)
Pseudohelminthosporium Phukhams. & K.D. Hyde (1)

Neophaeosphaeriaceae Ariyaw. & K.D. Hyde


Neophaeosphaeria M.P.S. Câmara, M.E. Palm & A.W. Ramaley (6)

Neopyrenochaetaceae Valenz.-Lopez, Crous, Cano, Guarro & Stchigel


Neopyrenochaeta Valenz.-Lopez, Crous, Stchigel, Guarro & Cano (5)

Nigrogranaceae Jaklitsch & Voglmayr


Nigrograna Gruyter, Verkley & Crous (12)

1122
Occultibambusaceae D.Q. Dai & K.D. Hyde
Brunneofusispora S.K. Huang & K.D. Hyde (1)*
Neooccultibambusa Doilom & K.D. Hyde (4)
Occultibambusa D.Q. Dai & K.D. Hyde (7)
Seriascoma Phook., D.Q. Dai & K.D. Hyde (2)
Versicolorisporium Sat. Hatak., Kaz. Tanaka & Y. Harada (1)

Ohleriaceae Jaklitsch & Voglmayr


Ohleria Fuckel (13)

Parabambusicolaceae Kaz. Tanaka & K. Hiray.


Aquastroma Kaz. Tanaka & K. Hiray. (1)
Lonicericola Phook., Jayasiri & K.D. Hyde (1)*
Multilocularia Phook. (1)
Multiseptospora Phook. & K.D. Hyde (2)
Neoaquastroma Wanas., E.B.G. Jones & K.D. Hyde (3)
Parabambusicola Kaz. Tanaka & K. Hiray. (2)
Paramonodictys N.G. Liu, K.D. Hyde & J.K. Liu (1)
Paratrimmatostroma Jayasiri, Phook., D.J. Bhat & K.D. Hyde (1)*
Pseudomonodictys Doilom, Ariyaw., Bhat & K.D. Hyde (1)

Paradictyoarthriniaceae Doilom, Ariyaw., Bhat & K.D. Hyde


Paradictyoarthrinium Matsush. (4)
Xenomassariosphaeria Jayasiri, Wanas. & K.D. Hyde (1)

Paralophiostomataceae V.V. Sarma & M. Niranjan.


Paralophiostoma V.V. Sarma & M. Niranjan. (1)

Parapyrenochaetaceae Valenz.-Lopez, Crous, Stchigel, Guarro & Cano


Parapyrenochaeta Valenz.-Lopez, Crous, Stchigel, Guarro & Cano (2)
Quixadomyces Cantillo & Gusmão (1)

Periconiaceae Nann.
Bambusistroma D.Q. Dai & K.D. Hyde (1)
Flavomyces D.G. Knapp, Kovács, J.Z. Groenew. & Crous (1)
Noosia Crous, R.G. Shivas & McTaggart, Persoonia (1)*
Periconia Tode (46)

Phaeoseptaceae S. Boonmee, Thambug. & K.D. Hyde


Phaeoseptum Ying Zhang, J. Fourn. & K.D. Hyde (2)
Pleopunctum N.G. Liu, K.D. Hyde & J.K. Liu (2)

Phaeosphaeriaceae M.E. Barr


Acericola Wanas., Camporesi, E.B.G. Jones & K.D. Hyde (1)
Allophaeosphaeria Ariyaw., Camporesi & K.D. Hyde (3)
Amarenographium O.E. Erikss. (4)
Amarenomyces O.E. Erikss. (2)*
Ampelomyces Ces. ex Schltdl. (ca. 5)
Aphanostigme Syd. (21)
Arezzomyces Y. Marín & Crous (1)
Banksiophoma Crous (1)
Bhagirathimyces S.M. Singh & S.K. Singh (1)

1123
Bhatiellae Wanas., Camporesi & K.D. Hyde (1)
Bricookea M.E. Barr (1)
Brunneomurispora Phook., Wanas. & K.D. Hyde (1)*
Camarosporioides W.J. Li & K.D. Hyde (1)
Chaetosphaeronema Moesz (12)
Dactylidina Wanas., Camporesi & K.D. Hyde (2)
Dematiopleospora Wanas., Camporesi, E.B.G. Jones & K.D. Hyde (8)
Didymocyrtis Vain. (21)
Dlhawksworthia Wanas., Camporesi & K.D. Hyde (3)
Edenia M.C. González, A.L. Anaya, Glenn, Saucedo & Hanlin (2)
Embarria Wanas., Camporesi & K.D. Hyde (1)
Equiseticola Abdelsalam, Tibpromma, Wanas. & K.D. Hyde (1)
Eudarluca Speg. (8)*
Galiicola Tibpromma, Camporesi & K.D. Hyde (3)
Hydeomyces Maharachch., H.A. Ariyaw., Wanas. & Al-Sadi (2)
Hydeopsis J.F. Zhang, J.K. Liu & Z.Y. Liu (1)
Italica Wanas., Camporesi & K.D. Hyde (2)
Jeremyomyces Crous & R.K. Schumach. (1)
Juncaceicola Tennakoon, Camporesi, Phook. & K.D. Hyde (8)
Kwanghwana A. Karunarathna, C. H. Kuo & K. D. Hyde (1)
Leptospora Rabenh. (15)
Longispora Phukhams. & K.D. Hyde (1)
Loratospora Kohlm. & Volkm.-Kohlm. (2)
Mauginiella Cavara (1)
Melnikia Wijayaw., Goonas., Bhat & K.D. Hyde (1)
Murichromolaenicola Mapook & K.D. Hyde (2)
Muriphaeosphaeria Phukhams., Bulgakov & K.D. Hyde (3)
Neoophiobolus Mapook & K.D. Hyde (1)
Neosetophoma Gruyter, Aveskamp & Verkley (15)
Neosphaerellopsis Crous & Trakun. (10)
Neostagonospora Quaedvl., Verkley & Crous (6)
Neostagonosporella C.L. Yang, X.L. Xu & K.D. Hyde (1)*
Neosulcatispora Crous & M.J. Wingf. (2)
Nodulosphaeria Rabenh. (ca. 52)
Ophiobolopsis Phook., Wanas. & K.D. Hyde (1)*
Ophiobolus Riess (350)
Ophiosimulans Tibpromma, Camporesi & K.D. Hyde (1)
Ophiosphaerella Speg. (10)
Paraleptospora Mapook & K.D. Hyde (2)
Paraloratospora Bundhun, Tennakoon, Phookamsak & K.D. Hyde (2)
Paraophiobolus Phook., Wanas. & K.D. Hyde (2)*
Paraphoma Morgan-Jones & J.F. White (8)
Parastagonospora Quaedvl., Verkley & Crous (ca. 19)
Parastagonosporella M. Bakhshi, Arzanlou & Crous (1)
Phaeopoacea Thambug., Dissan. & K.D. Hyde (3)*
Phaeoseptoriella Crous (1)
Phaeosphaeria I. Miyake (ca. 95)
Phaeosphaeriopsis M.P.S. Câmara, M.E. Palm & A.W. Ramaley (12)
Phaeostagonospora A.W. Ramaley (1)
Piniphoma Crous & R.K. Schumach. (1)
Poaceicola W.J. Li, Camporesi, Bhat & K.D. Hyde (10)
Populocrescentia Wanas., E.B.G. Jones & K.D. Hyde (3)

1124
Pseudoophiobolus Phook., Wanas. & K.D. Hyde (8)*
Pseudoophiosphaerella J.F. Zhang, J.K. Liu & Z.Y. Liu (1)
Pseudophaeosphaeria Jayasiri, Camporesi & K.D. Hyde (1)
Pseudostaurosphaeria Mapook & K.D. Hyde (2)
Sclerostagonospora Höhn. (ca. 15)
Scolicosporium Lib. ex Roum. (13)
Septoriella Oudem. (=Wojnowicia Sacc.) (22)*
Setomelanomma M. Morelet (1)
Setophoma Gruyter, Aveskamp & Verkley (6)
Sulcispora Shoemaker & C.E. Babc. (2)
Tiarospora Sacc. & Marchal (3)
Tintelnotia S.A. Ahmed, Hofmüller, M. Seibold & de Hoog (2)
Vagicola K.W.T. Chethana & K.D. Hyde (1)
Vittaliana Devadatha, Nikita, A. Baghela & V.V. Sarma (1)*
Vrystaatia Quaedvl., W.J. Swart, Verkley & Crous (1)
Wingfieldomyces Y. Marín & Crous (1)
Wojnowiciella Crous, Hern.-Restr. & M.J. Wingf. (9)
Xenophaeosphaeria Crous & M.J. Wingf. (1)
Xenophoma Crous & Trakunyingcharoen (1)
Xenoseptoria Quaedvl., H.D. Shin, Verkley & Crous (1)
Yunnanensis Karun., Phook. & K.D. Hyde (1)*

Pleomassariaceae M.E. Barr


Beverwykella Tubaki (3)
Lichenopyrenis Calat., Sanz & Aptroot (1)
Myxocyclus Riess (1)
Peridiothelia D. Hawksw. (3)
Prosthemium Kunze (ca. 8)
Pseudotrichia Kirschst. (ca. 8)
Splanchnonema Corda (37)

Pleomonodictydaceae Hern.-Restr., J. Mena & Gené


Pleomonodictys Hern.-Restr., J. Mena & Gené (2)
Pleohelicoon Jayasiri, E.B.G. Jones & K.D. Hyde (2)

Pleosporaceae Nitschke
Allonecte Syd. (3)
Alternaria Nees (ca. 360)*
Bipolaris Shoemaker (69)
Clathrospora Rabenh. (20)
Comoclathris Clem. (30)
Curvularia Boedijn (119)
Decorospora Inderb., Kohlm. & Volkm.-Kohlm. (1)
Diademosa Shoemaker & C.E. Babc. (4)*
Dichotomophthora Mehrl. & Fitzp. ex P.N. Rao (6)
Exserohilum K.J. Leonard & Suggs (ca. 30)
Extrawettsteinina M.E. Barr (4)
Gibbago E.G. Simmons (1)
Johnalcornia Y.P. Tan & R.G. Shivas (1)
Paradendryphiella Woudenberg & Crous (2)
Platysporoides (Wehm.) Shoemaker & C.E. Babc. (11)
Pleoseptum A.W. Ramaley & M.E. Barr (1)

1125
Porocercospora Amaradasa, Amundsen, Madrid & Crous (1)*
Prathoda Subram. (2)*
Pseudoyuconia Lar.N. Vassiljeva (1)
Pyrenophora Fr. (= Marielliottia Shoemaker) (ca. 95)
Stemphylium Wallr. (ca. 96)*
Tamaricicola Thambug., Camporesi & K.D. Hyde (1)
Typhicola Crous (1)

Pseudoastrosphaeriellaceae Phook. & K.D. Hyde


Carinispora K.D. Hyde (2)
Pseudoastrosphaeriella Phook., Z.L. Luo & K.D. Hyde (6)
Pseudoastrosphaeriellopsis Devadatha, Wanas., Jeewon & V.V. Sarma (1)*

Pseudoberkleasmiaceae Phukhams. & K.D. Hyde


Pseudoberkleasmium Tibpromma & K.D. Hyde (3)

Pseudocoleodictyosporaceae Doilom & K.D. Hyde


Pseudocoleodictyospora Doilom & K.D. Hyde (3)
Subglobosporium Doilom & K.D. Hyde (1)

Pseudolophiotremataceae K.D. Hyde & Hongsanan


Clematidis Tibpromma, Camporesi & K.D. Hyde (1)
Pseudolophiotrema A. Hashim. & Kaz. Tanaka (1)

Pseudomassarinaceae Phukhams & K.D. Hyde


Pseudomassarina Phukhams. & K.D. Hyde (1)

Pseudopyrenochaetaceae Valenz.-López, Crous, Stchigel, Guarro & J.F. Cano


Pseudopyrenochaeta Valenzuela-López, Crous, Stchigel, Guarro & Cano (2)

Pyrenochaetopsidaceae Valenz.-López, Crous, Cano, Guarro & Stchigel


Pyrenochaetopsis Gruyter, Aveskamp & Verkley (7)
Neopyrenochaetopsis Valenz-López, Cano, Guarro & Stchigel (1)
Xenopyrenochaetopsis Valenz.-Lopez, Crous, Stchigel, Guarro & Cano (1)

Roussoellaceae Jian K. Liu, Phook., D.Q. Dai & K.D. Hyde


Appendispora K.D. Hyde (2)
Cytoplea Bizz. & Sacc. (5)
Elongatopedicellata Jin F. Zhang, Jian K. Liu, K.D. Hyde & Zi Y. Liu (1)
Immotthia M.E. Barr (2)*
Neoroussoella Jian K. Liu, Phook. & K.D. Hyde (7)
Pararoussoella Wanas., E.B.G. Jones & K.D. Hyde (3)*
Pseudoneoconiothyrium Wanas., Phukhams., Camporesi & K.D. Hyde (1)
Pseudoroussoella Mapook & K.D. Hyde (2)
Roussoella Sacc. (38)
Roussoellopsis I. Hino & Katum. (3)
Setoarthopyrenia Mapook & K.D. Hyde (1)
Xenoroussoella Mapook & K.D. Hyde (1)

Salsugineaceae K.D. Hyde & Tibpromma


Acrocordiopsis Borse & K.D. Hyde (2)
Salsuginea K.D. Hyde (1)

1126
Shiraiaceae Y.X. Liu, Zi Y. Liu & K.D. Hyde
Grandigallia M.E. Barr, Hanlin, Cedeño, Parra & R. Hern. (1)
Rubroshiraia D.Q. Dai & K.D. Hyde (1)
Shiraia Henn. (1)

Sporormiaceae Munk
Chaetopreussia Locq.-Lin. (1)
Forliomyces Phukhams., Camporesi & K.D. Hyde (1)
Pleophragmia Fuckel (1)
Preussia Fuckel (51)
Sparticola Phukhams., Ariyaw., Camporesi & K.D. Hyde (4)
Sporormia De Not. (29)
Sporormiella Ellis & Everh. (60)*
Sporormurispora Wanas., Bulgakov, Gafforov & K.D. Hyde (2)
Westerdykella Stolk (50)

Striatiguttulaceae S.N. Zhang, K.D. Hyde & J.K. Liu


Longicorpus S.N. Zhang, K.D. Hyde & J.K. Liu (1)
Striatiguttula S.N. Zhang, K.D. Hyde & J.K. Liu (2)

Sulcatisporaceae Kaz. Tanaka & K. Hiray.


Anthosulcatispora Phukhams. & K.D. Hyde (2)
Magnicamarosporium Kaz. Tanaka & K. Hiray. (2)
Neobambusicola Crous & M.J. Wingf. (2)
Parasulcatispora Phukhams. & K.D. Hyde (1)
Pseudobambusicola Hern.-Restr. & Crous (1)*
Sulcatispora Kaz. Tanaka & K. Hiray. (2)

Teichosporaceae M.E. Barr


Asymmetrispora Thambug. & K.D. Hyde (2)
Aurantiascoma Thambug. & K.D. Hyde (1)
Chaetomastia (Sacc.) Berl. (10)
Floricola Kohlm. & Volkm.-Kohlm. (2)
Loculohypoxylon M.E. Barr (1)
Magnibotryascoma Thambug. & K.D. Hyde (2)
Misturatosphaeria Mugambi & Huhndorf (2)
Paulkirkia Wijayaw., Wanas., Tangthir., Camporesi & K.D. Hyde (1)
Pseudoaurantiascoma Thambug. & K.D. Hyde (1)
Pseudomisturatosphaeria Thambug. & K.D. Hyde (1)
Ramusculicola Thambug. & K.D. Hyde (1)
Sinodidymella J.Z. Yue & O.E. Erikss. (5)
Teichospora Fuckel (35)

Testudinaceae Arx
Angustospora Abdel-Aziz (1)
Halotestudina Dayar. & K.D. Hyde (1)
Lepidosphaeria Parg.-Leduc (1)
Lojkania Rehm (10)
Muritestudina Wanas., E.B.G. Jones & K.D. Hyde (1)
Neotestudina Segretain & Destombes (3)
Testudina Bizz. (1)
Ulospora D. Hawksw., Malloch & Sivan. (1)

1127
Verruculina Kohlm. & Volkm.-Kohlm. (1)

Tetraplosphaeriaceae Kaz. Tanaka & K. Hiray


Byssolophis Clem. (3)
Ernakulamia Subram. (2)
Polyplosphaeria Kaz. Tanaka & K. Hiray. (5)
Pseudotetraploa Kaz. Tanaka & K. Hiray. (4)
Quadricrura Kaz. Tanaka, K. Hiray. & Sat. Hatak. (3)
Shrungabeeja V.G. Rao & K.A. Reddy (5)
Tetraploa Berk. & Broome (19)
Triplosphaeria Kaz. Tanaka & K. Hiray (4)

Thyridariaceae Q. Tian & K.D. Hyde


Chromolaenomyces Mapook & K.D. Hyde (1)
Cycasicola Wanas., E.B.G. Jones & K.D. Hyde (2)
Liua Phook. & K.D. Hyde (1)*
Parathyridaria Jaklitsch & Voglmayr (5)
Pseudothyridariella Mapook & K.D. Hyde (2)
Thyridaria Sacc. (52)
Thyridariella Devadatha, V.V. Sarma, K.D. Hyde, Wanas. & E.B.G Jones (2)

Torulaceae Corda
Dendryphion Wallr. (68)
Neotorula Ariyaw., Z.L. Luo & K.D. Hyde (2)
Rostriconidium Z.L. Luo, K.D. Hyde & H.Y. Su (2)
Rutola J.L. Crane & Schokn. (1)
Sporidesmioides Jun F. Li, Phook. & K.D. Hyde (1)
Torula Pers. (12)

Trematosphaeriaceae K.D. Hyde, Y. Zhang ter, Suetrong & E.B.G. Jones


Bryosphaeria Döbbeler (9)
Falciformispora K.D. Hyde (4)
Hadrospora Boise (2)
Halomassarina Suetrong, Sakay., E.B.G. Jones, Kohlm., Volkm.-Kohlm. & C.L. Schoc (1)
Raghukumaria Devadatha, V.V Sarma & E.B.G Jones (1)
Trematosphaeria Fuckel (20)

Tzeananiaceae H.A. Ariyaw., A.J.L. Phillips & Chuang


Tzeanania H.A. Ariyaw., A.J.L. Phillips & Chuang (1)

Wicklowiaceae Ariyaw. & K.D. Hyde


Wicklowia Raja, A. Ferrer & Shearer (2)

Zopfiaceae G. Arnaud ex D. Hawksw.


Celtidia J.M. Janse (1)
Coronopapilla Kohlm. & Volkm.-Kohlm. (2)
Rechingeriella Petr. (2)
Richonia Boud. (1)
Zopfia Rabenh. (5)
Zopfiofoveola D. Hawksw. (1)

1128
Pleosporales genera incertae sedis
Acuminatispora S.N. Zhang, K.D. Hyde & J.K. Liu (1)
Aegeanispora E.B.G. Jones & Abdel-Wahab (1)
Antealophiotrema A. Hashim. & Kaz. Tanaka (1)
Ascorhombispora L. Cai & K.D. Hyde (1)
Atradidymella Davey & Currah (1)
Briansuttonia R.F. Castañeda, Minter & Saikawa (1)
Camarographium Bubák (7)
Chaetodiplodia P. Karst. (9)
Chaetophoma Cooke (ca. 30)
Cheiromoniliophora Tzean & J.L. Chen (4)
Crassiperidium M. Matsum. & Kaz. Tanaka (2)
Cyclothyrium Petr. (2)
Dangeardiella Sacc. & P. Syd. (2)
Daruvedia Dennis (1)
Dokmaia I. Promputtha (1)
Farasanispora Abdel-Wahab, Bahkali & E.B.G. Jones (1)
Glaxoa P.F. Cannon (1)
Homostegia Fuckel (2)
Hobus Jaklitsch & Voglmayr (1)
Inflatispora Y. Zhang ter, J. Fourn. & K.D. Hyde (2)
Isthmosporella Shearer & J.L. Crane (1)
Megacapitula J.L. Chen & Tzean (1)
Megatomentella D.A.C. Almeida, Gusmão & A.N. Mill. (1)
Neocurreya Thambug. & K.D. Hyde (5)
Ostropella (Sacc.) Höhn. (5)
Paraepicoccum Matsush. (1)
Paraliomyces Kohlm. (1)
Parameliola Hongsanan, Peršoh & K.D. Hyde (2)
Perthomyces Crous (1)
Phialophorophoma Linder (1)
Pleosphaerellula Naumov & Czerepan. (2)
Pseudohendersonia Crous & M.E. Palm (2)
Pseudopassalora Crous (1)
Pyrenochaeta De Not. (5)
Rebentischia P. Karst. (16)
Repetophragma Subram. (38)
Scleroramularia Batzer & Crous (6)
Scolecobasidium E.V. Abbott (64)
Setophaeosphaeria Crous & Y. Zhang ter (6)
Sirodesmium De Not. (ca. 25)
Spiroplana Voglmayr, M.J. Park & H.D. Shin (1)
Stuartella Fabre (6)
Xenolophium Syd. (ca. 5)

Pleosporomycetidae genus incertae sedis


Hysterographium Corda (3)

Dothideomycetes orders incertae sedis


Abrothallales Pérez-Ort. & Suija [= Lichenoconiales Diederich, Lawrey & K.D. Hyde]
Lichenoconiaceae Diederich & Lawrey [= Abrothallaceae Pérez-Ort. & Suija]*
Abrothallus De Not (=Epinephroma Zhurb.; Vouauxiomyces Dyko & D. Hawks.) (42)*

1129
Lichenoconium Petr. & Syd. (15)

Acrospermales Minter, Peredo & A.T. Watson


Acrospermaceae Fuckel
Acrospermum Tode (12)
Gonatophragmium Deighton (17)
Oomyces Berk. & Broome (7)

Acrospermales genus incertae sedis


Pseudovirgaria H.D. Shin, U. Braun, Arzanlou & Crous (2)

Asterinales M.E. Barr ex D. Hawksw. & O.E. Erikss. (= Asterotexales Firmino et al.)
Asterinaceae Hansf.
Asterina Lév. (ca. 1085)
Asterinella Theiss. (ca. 39)
Asterolibertia G. Arnaud (ca. 30)
Asterostomella Speg.
Batistinula Arx (1)
Cirsosia G. Arnaud (18)
Dothidasteromella Höhn. (11)
Echidnodella Theiss. & Syd. (35)
Halbania Racib. (3)
Meliolaster Höhn. (3)
Parasterinopsis Bat. (3)
Platypeltella Petr. (3)
Prillieuxina G. Arnaud (66)
Pycnocarpon Theiss.
Schenckiella Henn. (1)
Trichasterina G. Arnaud (11)
Trichopeltospora Bat. & Cif. (2)
Uleothyrium Petr. (3)
Vizellopsis Bat., J.L. Bezerra & T.T. Barros (1)

Asterotexaceae Firmino, O.L. Pereira & Crous


Asterotexis Arx (2)

Hemigraphaceae D.Q. Dai & K.D. Hyde*


Hemigrapha (Müll. Arg.) D. Hawksw. (8)

Lembosiaceae Hosag.
Lembosia Lév. (ca. 200)

Melaspileellaceae D.Q. Dai & K.D. Hyde*


Melaspileella (P. Karst.) Vain. (1)

Morenoinaceae Hongsanan & K.D. Hyde


Morenoina Theiss. (ca. 25)

Neobueliellaceae Hongsanan & K.D. Hyde


Neobueliella Hongsanan & K.D. Hyde

1130
Stictographaceae D.Q. Dai & K.D. Hyde*
Buelliella Fink (12)
Karschia Körb. (4)
Labrocarpon Etayo & Pérez-Ort. (1)
Melaspileopsis (Müll. Arg.) Ertz & Diederich (1)
Stictographa Mudd (2)

Asterinales genera incertae sedis


Andamanomyces Hosag. (1)
Caribaeomyces Cif. (1)
Discopycnothyrium Hongsanan & K.D. Hyde (1)
Hazslinszkya Körb. (4)
Inocyclus Theiss. & Syd. (6)
Melanographa Müll. Arg. (1)
Pirozynskiella S. Hughes (3)
Vishnumyces Hosag. (1)

Botryosphaeriales C.L. Schoch, Crous & Shoemaker


Aplosporellaceae Slippers, Boissin & Crous
Alanomyces Roh. Sharma (1)*
Aplosporella Speg. (= Bagnisiella Speg.) (10)*

Botryosphaeriaceae Theiss. & Syd. (= Endomelanconiopsidaceae Tao Yang & Crous)*


Alanphillipsia Crous & M.J. Wingf. (5)
Barriopsis A.J.L. Phillips, A. Alves & Crous (5)
Botryobambusa Phook., J.K. Liu & K.D. Hyde (2)
Botryosphaeria Ces. & De Not. (13)
Cophinforma Doilom, J.K. Liu & K.D. Hyde (2)
Diplodia Fr. (more than 1000 names in MycoBank, 30 known from culture)
Dothiorella Sacc. (389 names in MycoBank, 38 known from culture) (= Spencermartinsia
A.J.L. Phillips, A. Alves & Crous)*
Endomelanconiopsis Rojas & Samuels (2)*
Eutiarosporella Crous (7)*
Lasiodiplodia Ellis & Everh. (35)
Macrophomina Petr. (2)
Marasasiomyces Crous (1)*
Mucoharknessia Crous, R.M. Sánchez & Bianchin. (2)*
Neodeightonia Booth (6)
Neofusicoccum Crous, Slippers & A.J.L. Phillips (4)
Neoscytalidium Crous & Slippers (3)*
Oblongocollomyces Tao Yang & Crous (1)*
Phaeobotryon Theiss. & Syd. (4)
Sakireeta Subram. & K. Ramakr. (1)*
Sardiniella Linaldeddu, A. Alves & A.J.L. Phillips (1)*
Sphaeropsis Sacc. (more than 600 names in MycoBank, 4 known from culture)
Tiarosporella Höhn. (2)

Melanopsaceae Phillips A.J.L., Slippers, Boissin & Crous


Melanops Nitschke ex Fuckel (105 names in MycoBank, 4 known from culture)

Phyllostictaceae Fr. (= Pseudofusicoccumaceae Tao Yang & Crous)


Phyllosticta Pers. (ca. 53)

1131
Pseudofusicoccum Mohali, Slippers & M.J. Wingf. (7)*

Planistromellaceae M.E. Barr


Kellermania Ellis & Everh. (ca. 16)*
Mycosphaerellopsis Höhn. (1)
Planistroma A.W. Ramaley (6)
Umthunziomyces Crous & M.J. Wingf. (1)*

Saccharataceae Slippers, Boissin & Crous (= Septorioideaceae Wyka & Broders)


Neoseptorioides Crous, Jacq. Edwards & Pascoe (1)*
Pileospora Tanney & Seifert (1)
Saccharata Denman & Crous (20)
Septorioides Quaedvl., Verkley & Crous (2)*

Botryosphaeriales genera incertae sedis


Auerswaldiella Theiss. & Syd. (7)
Coccostromella Petr. (1)
Leptoguignardia E. Müll. (1)
Metameris Theiss. & Syd. (5)
Phyllachorella Syd. (8)
Pilgeriella Henn. (2)
Sivanesania W.H. Hsieh & Chi Y. Chen (1)
Vestergrenia Rehm (3)

Catinellales Ekanayaka, K.D. Hyde & Ariyaw.


Catinellaceae Ekanayaka, K.D. Hyde & Ariyaw.
Catinella Boud. (1 or 2)

Cladoriellales Crous
Cladoriellaceae Crous
Cladoriella Crous (5)

Collemopsidiales Pérez-Ort., Garrido-Ben. & Grube


Xanthopyreniaceae Zahlbr.
Collemopsidium Nyl. (27)
Didymellopsis (Sacc.) Clem. & Shear (6)
Frigidopyrenia Grube (1)
Xanthopyrenia Bachm. (2)
Zwackhiomacromyces Etayo & van den Boom (2)
Zwackhiomyces Grube & Hafellner (35)

Dyfrolomycetales K.L. Pang, K.D. Hyde & E.B.G. Jones


Pleurotremataceae Walt. Watson
Dyfrolomyces K.D. Hyde, K.L. Pang, Alias, Suetrong & E.B.G. Jones (8)
Melomastia Nitschke ex Sacc. (4)
Pleurotrema Müll. Arg. (1)

Eremithallales Lücking & Lumbsch


Melaspileaceae W. Watson (= Eremithallaceae Lücking & Lumbsch)
Encephalographa A. Massal. (1)
Melaspilea Nyl. (1 + c. 75 orphaned) (= Eremithallus Lücking et al.)

1132
Eremomycetales Pem & Hyde
Eremomycetaceae Malloch & Cain
Eremomyces Malloch & Cain (2)
Rhexothecium Samson & Mouch. (1)

Eremomycetales genus incertae sedis


Arthrographis G. Cochet ex Sigler (12)

Jahnulales K.L. Pang, Abdel-Wahab, El-Shar., E.B.G. Jones & Sivichai


Aliquandostipitaceae Inderbitzin
Aliquandostipite Inderbitzin (7)
Brachiosphaera Nawawi (2)
Jahnula Kirschst. (19)
Megalohypha A. Ferrer & Shearer (1)
Neojahnula W. Dong, H. Zhang & K.D. Hyde (1)
Pseudojahnula W. Dong, H. Zhang & K.D. Hyde (1)
Xylomyces Goos, R.D. Brooks & Lamore (8)*

Manglicolaceae Suetrong & E.B.G. Jones


Manglicola Kohlm. & E. Kohlm. (1)*

Kirschsteiniotheliales Hern.-Restr., R.F. Castañeda, Gené & Crous


Kirschsteiniotheliaceae Boonmee & K.D. Hyde
Kirschsteiniothelia D. Hawksw. (29)

Kirschsteiniotheliales genera incertae sedis


Brachysporiella Bat. (Brachysporiella s. lato.) (15)
Taeniolella S. Hughes sensu lato*

Lembosinales Crous
Lembosinaceae Crous
Lembosina Theiss. (21)

Lichenotheliales K. Knudsen, Muggia & K.D. Hyde


Lichenotheliaceae Henssen
Lichenothelia D. Hawksw. (27)
Endococcus Nyl. (44)

Microthyriales G. Arnaud
Microthyriaceae Sacc.
Arnaudiella Petr. (12)
Calothyriopsis Höhn. (4)
Chaetothyriothecium Hongsanan & K.D. Hyde (1)
Hamatispora L.T.H. Yen, K. Yamag. & K. Ando (1)
Microthyrium Desm. (ca. 180)
Neoanungitea Crous (1)
Paramicrothyrium H.X. Wu & K.D. Hyde (1)
Pseudomicrothyrium X.Y. Zeng, S. Hongsanan & K.D. Hyde (1)
Pseudopenidiella Crous & Koukol (1)
Seynesiella G. Arnaud (5)
Tumidispora Hongsanan & K.D. Hyde (1)

1133
Microthyriales genera incertae sedis
Heliocephala V. Rao, K.A. Reddy & de Hoog (7)
Mitopeltis Speg. (2)
Neoscolecobasidium Crous (1)
Parazalerion Madrid, Gené & Cano (1)*
Thyriodictyella Cif (1)
Tothia Bat. (2)

Minutisphaerales Raja, Oberlies, Shearer & A.N. Mill.


Acrogenosporaceae Jayasiri & K.D. Hyde*
Acrogenospora M.B. Ellis (12)

Minutisphaeraceae Raja, Oberlies, Shearer & A.N. Mill.


Minutisphaera Shearer, A.N. Mill. & A. Ferrer (4)

Monoblastiales Lücking, M.P. Nelsen & K.D. Hyde


Monoblastiaceae Walt. Watson
Acrocordia A. Massal. (6)
Anisomeridium (Müll. Arg.) M. Choisy (ca. 80)
Caprettia Bat. & H. Maia (8)
Megalotremis Aptroot (12)
Monoblastia Riddle (11)
Trypetheliopsis Asahina (6)

Murramarangomycetales Crous
Murramarangomycetaceae Crous
Murramarangomyces Crous (1)

Muyocopronales Mapook, Boonmee & K.D. Hyde


Muyocopronaceae K.D. Hyde
Arxiella Papendorf (3)
Leptodiscella Papendorf (5)
Muyocopron Speg. (51)
Mycoleptodiscus Ostaz. (18)
Neocochlearomyces Pinruan, Sommai, Suetrong, J.Z. Groenew. & Crous (1)
Neomycoleptodiscus Hern.-Restr., J.D.P. Bezerra & Crous (1)
Paramycoleptodiscus Crous & M.J. Wingf. (1)
Pseudopalawania Mapook & K.D. Hyde (1)
Setoapiospora Mapook & K.D. Hyde (1)

Natipusillales Raja, Shearer, A.N. Mill. & K.D. Hyde


Natipusillaceae Raja, Shearer & A.N. Mill.
Natipusilla A. Ferrer, A.N. Mill. & Shearer (4)

Parmulariales D.Q. Dai & K.D. Hyde*


Parmulariaceae E. Müll. & Arx ex M.E. Barr
Aldona Racib. (3)
Aldonata Sivan. & A.R.P. Sinha (1)
Antoniomyces Inácio (1)
Aulacostroma Syd. & P. Syd. (5)
Campoa Speg. (4)
Cirsosiopsis Butin & Speer (1)

1134
Cocconia Sacc. (13)
Cycloschizon P. Henn. (13)
Cyclostomella Pat. (4)
Dothidasteroma Höhn. (4)
Ferrarisia Sacc. (ca. 8)
Hysterostomella Speg. (23)
Kiehlia Viégas (2)
Mintera Inácio & P.F. Cannon (1)
Pachypatella Theiss. & Syd. (1)
Palawaniella Doidge (7)
Parmularia Lév. (6)
Parmulariopsella Sivan. (1)
Parmulariopsis Petr. (1)
Parmulina Theiss. & Syd. (6)
Placoasterella Sacc. ex Theiss. & Syd. (4)
Placosoma Syd. (2)
Placostromella Petr. (3)
Pleiostomellina Bat., J.L. Bezerra & H. Maia (1)
Polycyclina Theiss. & Syd. (1)
Polycyclus Höhn. (2)
Protothyrium G. Arnaud (4)
Pseudolembosia Theiss. (4)
Rhagadolobiopsis Guatim. & R.W. Barreto (1)
Rhagadolobium P. Henn. & Lindau (10)
Rhipidocarpon (Theiss.) Theiss. & Syd. (1)
Symphaeophyma Speg. (1)
Syrropeltis Bat., J.L. Bezerra & Matta (1)
Thallomyces H.J. Swart (1)
Viegasella Inácio & P.F. Cannon (1)

Patellariales D. Hawksw. & O.E. Erikss.


Patellariaceae Corda
Baggea Auersw. (1)
Banhegyia L. Zeller & Tóth (2)
Colensoniella Hafellner (1)
Endotryblidium Petr. (1)
Glyphium Nitschke ex F. Lehm. (ca. 4)
Haematomyxa Sacc (2)
Holmiella Petrini, Samuels & E. Müll. (4)
Hysteropatella Rehm (3)
Hysteropeltella Petr. (1)
Lahmiomyces Cif. & Tomas. (1)
Lecanidiella Sherwood (1)
Lirellodisca Aptroot (1)
Murangium Seaver (1)
Patellaria Fr. (12)
Poetschia Körb. (4)
Pseudoparodia Theiss. & Syd. (1)
Rhizodiscina Hafellner (1)
Rimula Velen. (1)
Schrakia Hafellner (1)
Stratisporella Hafellner (1)

1135
Tryblidaria (Sacc.) Rehm (9)

Phaeotrichales Ariyaw., Jian K. Liu & K.D. Hyde


Phaeotrichaceae Cain
Echinoascotheca Matsush. (1)
Phaeotrichum Cain & M.E. Barr (2)
Trichodelitschia Munk (4)

Stigmatodiscales Voglmayr & Jaklitsch


Stigmatodiscaceae Voglmayr & Jaklitsch
Stigmatodiscus Voglmayr & Jaklitsch (= Asterodiscus Voglmayr et al.) (6)*

Strigulales Lücking, M.P. Nelsen & K.D. Hyde


Strigulaceae Zahlbr. (= Phyllobatheliaceae Bitter & F. Schill.)
Dichoporis Clem. (18)
Flagellostrigula Lücking & S.H. Jiang (1)
Flavobathelium Lücking, Aptroot & G. Thor (1)
Phyllobathelium (Müll. Arg.) Müll. Arg (8)
Phyllocharis Fée (1)
Phyllocraterina Sérus. & Aptroot (= Phyllocratera Sérus. & Aptroot) (2)
Phylloporis Clem. (ca. 10)
Puiggariella Speg. (3)
Raciborskiella Höhnel (2)
Racoplaca Fée (5)
Serusiauxiella S.H. Jiang, Lücking & J.C.Wei (3)
Strigula Fr. (ca. 30)
Swinscowia S.H. Jiang & Lücking (33)

Tenuitholiascaceae S.H. Jiang, Lücking & J.C. Wei*


Tenuitholiascus S.H. Jiang, Lücking & J.C. Wei. (1)

Superstratomycetales van Nieuwenh., Miądl., Houbraken, Adan, Lutzoni & Samson


Superstratomycetaceae van Nieuwenh., Miądl., Houbraken, Adan, Lutzoni & Samson
Superstratomyces van Nieuwenh., Miądl. & Samson (4)

Trypetheliales Lücking, Aptroot & Sipman


Polycoccaceae Ertz, Hafellner & Diederich
Clypeococcum D. Hawksw. (ca. 10)
Polycoccum Saut. ex Körb. (ca. 60)

Trypetheliaceae Zenker
Alloarthopyrenia Phukhams., Lücking & K.D. Hyde (1)
Aptrootia Lücking & Sipman (3)
Architrypethelium Aptroot (8)
Astrothelium Eschw. (= Campylothelium Müll.) (ca. 275)
Bathelium Ach. (16)
Bogoriella Zahlbr. (=Distothelia Aptroot) (29)
Constrictolumina Lücking, M.P. Nelsen & Aptroot (9)
Dictyomeridium Aptroot, M.P. Nelsen & Lücking (7)
Macroconstrictolumina Lücking, R. Miranda & Aptroot (5)
Marcelaria Aptroot (= Buscalionia Sambo) (3)
Nigrovothelium Lücking, M.P. Nelsen & Aptroot (3)

1136
Novomicrothelia Aptroot, M.P. Nelsen & Lücking (1)
Polymeridium (Müll. Arg.) R.C. Harris (51)
Polypyrenula D. Hawksw. (1)
Pseudobogoriella Lücking, R. Miranda & Aptroot (15)
Pseudopyrenula Müll. Arg. (21)
Schummia Lücking, R. Miranda & Aptroot (1)
Trypethelium Sprengel (16)
Viridothelium Lücking, M.P. Nelsen & Aptroot (= ?Exiliseptum R.C. Harris fide Hongsanan
et al. 2020) (11)

Tubeufiales Boonmee & K.D. Hyde (= Bezerromycetales J.D.P. Bezerra et al.; =


Wiesneriomycetales J.D.P. Bezerra et al.)
Bezerromycetaceae J.D.P. Bezerra, Souza-Motta & Crous
Bezerromyces J.D.P. Bezerra, Souza-Motta & Crous (2)
Neorhamphoria Boonmee, E. Hüseyin & F. Selçuk (1)
Xiliomyces J.D.P. Bezerra, Souza-Motta & Crous (1)

Tubeufiaceae M.E. Barr


Acanthohelicospora Boonmee & K.D. Hyde (4)
Acanthophiobolus Berl. (6)
Acanthostigma De Not. (64)
Acanthostigmina Höhn. (7)
Acanthotubeufia Y.Z. Lu & K.D. Hyde (1)
Aquaphila Goh, K.D. Hyde & W.H. Ho (2)
Artocarpomyces Subram. (1)
Berkleasmium Zobel (ca. 40)
Bifrontia Norman (2)
Boerlagiomyces Butzin (9)
Camporesiomyces D.P. Wei & K.D. Hyde (1)
Chaetosphaerulina I. Hino (6)
Chlamydotubeufia Boonmee & K.D. Hyde (8)
Dematiohelicoma Y.Z. Lu, J.C. Kang & K.D. Hyde (2)
Dematiohelicomyces Y.Z. Lu, Boonmee & K.D. Hyde (1)
Dematiohelicosporum Y.Z. Lu, J.K. Liu & K.D. Hyde (1)
Dematiotubeufia Y.Z. Lu, Boonmee & K.D. Hyde (1)
Dictyospora Brahaman., Y.Z. Lu, Boonmee & K.D. Hyde (1)
Discotubeufia Jayasiri, E.B.G. Jones & K.D. Hyde (1)
Helicangiospora Boonmee, Bhat & K.D. Hyde (1)
Helicoarctatus Y.Z. Lu, J.C. Kang & K.D. Hyde (1)
Helicodochium J.S. Monteiro, R.F. Castañeda, A.C. Cruz & Gusmão (2)
Helicohyalinum Y.Z. Lu, J.K. Liu & K.D. Hyde (2)
Helicoma Corda (ca. 40)
Helicomyces Link (14)
Helicosporium Nees (ca. 15)
Helicotruncatum Y.Z. Lu, J.C. Kang & K.D. Hyde (1)
Helicotubeufia Y.Z. Lu & J.K. Liu (3)
Kamalomyces R.K. Verma, N. Sharma & Soni (5)
Kevinhydea N.G. Liu, Y.Z. Lu & J.K. Liu (1)
Manoharachariella Bagyan., N.K. Rao & Kunwar (4)
Muripulchra Z.L. Luo, Hong Y. Su & K.D. Hyde (1)
Neoacanthostigma Boonmee, Bhat & K.D. Hyde (8)
Neochlamydotubeufia Y.Z. Lu, Boonmee & K.D. Hyde (2)

1137
Neohelicoma Y.Z. Lu, Boonmee & K.D. Hyde (1)
Neohelicomyces Z.L. Luo, Bhat & K.D. Hyde (3)
Neohelicosporium Y.Z. Lu, J.C. Kang & K.D. Hyde (7)
Neotubeufia Chaiwan, Boonmee, Y.Z. Lu & K.D. Hyde (1)
Pleurohelicosporium Y.Z. Lu, J.C. Kang & K.D. Hyde (1)
Podonectria Petch (11)
Pseudohelicomyces Y.Z. Lu, J.K. Liu & K.D. Hyde (5)
Pseudohelicoon Y.Z. Lu & K.D. Hyde (2)
Tamhinispora Rajeshkumar & Rahul Sharma (2)
Thaxteriella Petr. (15)
Thaxteriellopsis Sivan., Panwar & S.J. Kaur (3)
Tubeufia Penz. & Sacc. (ca. 60)

Wiesneriomycetaceae Suetrong, Rungjind., Somrith. & E.B.G. Jones


Parawiesneriomyces Crous & M.J. Wingf. (1)
Phalangispora Nawawi & J. Webster (3)
Pseudogliophragma Phadke & V.G. Rao (1)
Setosynnema D.E. Shaw & B. Sutton (3)
Speiropsis Tubaki (8)
Wiesneriomyces Koord. (4)

Valsariales Jaklitsch, K.D. Hyde & Voglmayr


Valsariaceae Jaklitsch, K.D. Hyde & Voglmayr
Bambusaria Jaklitsch, D.Q. Dai, K.D. Hyde & Voglmayr (1)
Myrmaecium Nitschke ex Fuckel (ca. 3)
Valsaria Ces. & De Not. (140 epithets)

Venturiales Y. Zhang ter, C.L. Schoch & K.D. Hyde


Sympoventuriaceae Y. Zhang ter, C.L. Schoch & K.D. Hyde
Acroconidiellina M.B. Ellis (4)
Clavatispora Boonmee & K.D. Hyde (1)
Fusicladium Bonord. (75)
Matsushimaea Subram. (4)
Mycosisymbrium Carris (1)
Ochroconis de Hoog & Arx (28)
Sympoventuria Crous & Seifert (3)
Veronaeopsis Arzanlou & Crous (1)
Verruconis Samerp., H.J. Choi, van den Ende, Horré & de Hoog (5)
Yunnanomyces Tibpromma & K.D. Hyde (2)

Venturiaceae E. Müll. & Arx ex M.E. Barr


Apiosporina Höhn. (6)
Atopospora Petr. (4)
Caproventuria U. Braun (2)
Coleroa (Fr.) Rabenh. (56)
Dimeriella Speg. (51)
Dimerosporiopsis Henn. (1)
Magnohelicospora R.F. Castañeda, Hern.-Restr., Gené & Guarro (2)
Metacoleroa Petr. (1)
Neocoleroa Petr. (6)
Protoventuria Berl. & Sacc. (45)
Pseudoanungitea Crous (3)*

1138
Pseudoparodiella F. Stevens (1)
Tyrannosorus Unter. & Malloch (1)
Venturia Sacc. (ca. 60)

Venturiales genera incertae sedis


Cylindrosympodioides Crous & M.J. Wingf. (1)
Cylindrosympodium W.B. Kendr. & R.F. Castañeda (12)
Lasiobotrys Kunze (9)

Zeloasperisporiales Hongsanan & K.D. Hyde


Zeloasperisporiaceae Crous
Zeloasperisporium R.F. Castañeda (8)

Dothideomycetes families incertae sedis


Alinaceae Boonmee & K.D. Hyde
Alina Racib. (1)

Argynnaceae Shearer & J.L. Crane


Argynna Morgan (1)
Lepidopterella Shearer & J.L. Crane (2)

Ascoporiaceae Kutorga & D. Hawksw.


Ascoporia Samuels & A.I. Romero (1)

Aulographaceae Luttr. ex P.M. Kirk, P.F. Cannon & J.C. David


Aulographum Lib. (ca. 30)
Echidnodes Theiss. & Syd. (31)
Lembosiella Sacc. (1)
Thyriopsis Theiss. & Syd. (3)

Balladynaceae Boonmee & K.D. Hyde


Balladyna Racib. (41)
Balladynocallia Bat. (3)
Balladynopsis Theiss. & Syd. (10)

Cleistosphaeraceae Boonmee & K.D. Hyde


Cleistosphaera Syd. & P. Syd. (1)

Coccoideaceae P. Henn. ex Sacc. & D. Sacc.


Coccoidea P. Henn. (4)
Coccoidella Höhn. (9)
Englerodothis Theiss. & Syd. (3)

Cookellaceae Höhn. ex Saccardo & Trotter


Cookella Sacc. (4)
Pycnoderma Syd. & P. Syd. (2)

Dimeriaceae E. Müll. & Arx ex Arx & E. Müll.


Dimerium (Sacc. & P. Syd.) McAlpine (79)

Dubujianaceae D. Pem, Doilom & K.D. Hyde


Dubujiana D.R. Reynolds & G.S. Gilbert (1)

1139
Dysrhynchisceae Boonmee & K.D. Hyde
Dysrhynchis Clem. (4)

Endosporiaceae D. Pem
Endosporium Tsuneda (2)

Englerulaceae P. Henn.
Allosoma Syd. (5)
Digitosarcinella S. Hughes (1)
Englerula P. Henn. (13)
Goosia B. Song (1)
Parenglerula Höhn. (7)
Rhytidenglerula Höhn. (11)
Sarcinella Sacc. (ca. 70)
Thrauste Theiss. (3)

Eriomycetaceae Huanraluek & K.D. Hyde


Eriomyces Huanraluek, Thambugala & K.D. Hyde (1)
Funbolia Crous & Seifert (1)
Heleiosa Kohlm., Volkm.-Kohlm. & O.E. Erikss. (1)
Phellinocrescentia Crous & Decock (1)
Pseudopassalora Pseudopassalora Crous (1)

Homortomycetaceae Thambug., A.J.L. Phillips & K.D. Hyde


Homortomyces Crous & M.J. Wingf. (2)

Hyalomeliolinaceae Boonmee & K.D. Hyde


Hyalomeliolina F. Stevens (2)

Leptopeltidaceae Höhn. ex Trotter


Dothiopeltis E. Müll. (2)
Leptopeltis Höhn. (11)
Ronnigeria Petr. (1)
Staibia Bat. & Peres (1)

Macrovalsariaceae D. Pem, Doilom & K.D Hyde


Macrovalsaria Petr. (1)

Meliolinaceae S. Hughes
Briania D.R. Reynolds (1)
Meliolina Syd. & P. Syd. (ca. 40)

Mesnieraceae Arx & E. Müll.


Bondiella Piroz. (1)
Mesniera Sacc. & P. Syd. (1)
Stegasphaeria Syd. & P. Syd. (3)

Naetrocymbaceae Höhn. ex R.C. Harris


Bonaria Bat. (4)
Jarxia D. Hawksw. (2)
Leptorhaphis Körb. (14)
Naetrocymbe Körb. (1)

1140
Tomasellia A. Massal. (ca. 5)

Nematotheciaceae Boonmee & K.D. Hyde


Nematothecium Syd. & P. Syd. (5)
Nematostigma Syd. & P. Syd. (5)
Ophioparodia Petr. & Cif. (1)

Neoparodiaceae Boonmee & K.D. Hyde


Neoparodia Petr. & Cif. (1)

Palawaniaceae Mapook & K.D. Hyde


Palawania Syd. & P. Syd. (2)

Paranectriellaceae S. Boonmee & K.D. Hyde


Paranectriella (Henn. ex Sacc. & D. Sacc.) Magnus. (= Araneomyces Höhn.) (9)
Puttemansia Henn. (18)

Parodiellaceae Theiss. & H. Syd. ex M.E. Barr


Parodiella Speg. (4)

Perisporiopsidaceae E. Müll. & Arx ex R. Kirschner & T.A. Hofm. (= Parodiopsidaceae Toro)
Asteronia (Sacc.) Henn. (2)
Byssocallis Syd. (3)
Chevalieropsis G. Arnaud (1)
Parodiellina Henn. ex G. Arnaud (1)
Perisporiopsis Henn. (22)

Phaeodimeriellaceae Boonmee, Mapook & K.D. Hyde


Phaeodimeriella Speg. (30)

Pododimeriaceae Boonmee & K.D. Hyde


Chaetoscutula E. Müll. (1)
Pododimeria E. Müll. (4)

Polyclypeolinaceae Boonmee & K.D. Hyde


Polyclypeolina Bat. & I.H. Lima (1)

Polystomellaceae Theiss. & H. Syd.


Dermatodothella Viégas (1)
Dothidella Speg. (2)
Munkiella Speg. (3)
Parastigmatea Doidge (3)

Protoscyphaceae Kutorga & D. Hawksw.


Protoscypha Syd. (2)

Pseudoperisporiaceae Toro
Bryomyces Döbbeler (12)
Eudimeriolum Speg. (8)
Lasiostemma Theiss. (5)
Nematostoma Syd. & P. Syd. (13)

1141
Pseudorobillardaceae Crous
Pseudorobillarda M. Morelet (12)

Pyrenidiaceae Zahlbr.
Pyrenidium Nyl. (11)

Seynesiopeltidaceae K.D. Hyde


Seynesiopeltis F. Stevens & R.W. Ryan (1)

Stomatogeneceae Boonmee & K.D. Hyde


Stomatogene Theiss. (3)

Thyrinulaceae X.Y. Zeng, S. Hongsanan & K.D. Hyde


Blastacervulus H.J. Swart (2)
Paraopeba V.P. Abreu, A.A.M. Gomes, Firmino & O.L. Pereira (1)
Thyrinula Petr. & Syd. (= Alysidiella Crous) (1)

Toroaceae Boonmee & K.D. Hyde


Toroa Syd. (2)

Trichopeltinaceae Bat., C.A.A. Costa & Cif.


Acrogenotheca Cif. & Bat. (3)
Brefeldiella Speg. (4)
Saccardinula Speg. (11)
Trichopeltella Höhn. (1)
Trichopeltheca Bat. (2)
Trichopeltina Theiss. (2)
Trichothyrinula Petr. (2)

Trichothyriaceae Theiss.
Lichenopeltella Höhn. (48)
Macrographa Etayo (1)
Pachythyrium G. Arnaud ex Spooner & P.M. Kirk (1)
Trichothyrium Speg. (12)

Vizellaceae H.J. Swart


Acarella Syd. (1)
Blasdalea Sacc. & P. Syd. (1)
Vizella Sacc. (11)

Dothideomycetes genera incertae sedis


Acanthorus Bat. & Cavalc. (1)
Acanthostigmella Höhn. (6)
Achorella Theiss. & Syd. (10)
Actinomyxa Syd. & P. Syd. (1)
Alascospora Raja, Violi & Shearer (1)
Ampullifera Deighton (6)
Anguillosporella U. Braun (2)
Anopeltis Bat. & Peres (1)
Arkoola J. Walker & Stovold (1)
Armata W. Yamam. (1)
Ascominuta Ranghoo & K.D. Hyde (2)

1142
Asterinema Bat. & Gayão (3)
Asterodothis Theiss. (1)
Asteromassaria Höhn. (12)
Asteromella Pass. & Thüm. (ca. 265)
Asteroporum Müll. Arg. (7)
Auerswaldia Sacc. (ca. 20)
Bactrodesmium Cooke (ca. 50)
Bahusakala Subram. (4)
Brachyconidiella R.F. Castañeda & W.B. Kendr. (1)
Brooksia Hansf. (1)
Bryorella Döbbeler (10)
Bryostroma Döbbeler (8)
Bryothele Döbbeler (2)
Byssogene Syd. (2)
Callebaea Bat. (1)
Calyptra Theiss. & Syd. (5)
Capillataspora K.D. Hyde (1)
Caryosporella Kohlm. (1)
Catulus Malloch & Rogerson (1)
Ceramoclasteropsis Bat. & Cavalc. (2)
Ceratophoma Höhn. (2)
Cercidospora Körb. (101)
Cerodothis Muthappa (1)
Chaetocrea Syd. (1)
Chaetosticta Petr. & Syd. (3)
Chionomyces Deighton & Piroz. (7)
Chuppia Deighton (2)
Cilioplea Munk (ca. 10)
Cirsosina Bat. & J.L. Bezerra (2)
Clavariopsis De Wild. (ca. 5)
Clypeostroma Theiss. & Syd. (ca. 3)
Cocciscia Norman (2)
Coccochora Höhn. (4)
Coccochorina Hara (2)
Coccodothis Theiss. & Syd. (2)
Comesella Speg. (1)
Comminutispora A.W. Ramaley (1)
Coniosporium Link (ca. 20)
Crauatamyces Viégas (1)
Crotone Theiss. & Syd. (1)
Cryomyces Selbmann, de Hoog, Mazzaglia, Friedmann & Onofri (4)
Cyclotheca Theiss. (9)
Dactuliophora C.L. Leakey (5)
Dawsomyces Döbbeler (2)
Dawsophila Döbbeler (3)
Dermatodothis Racib. ex Theiss. & Syd. (6)
Dianesea Inácio & P.F. Cannon (1)
Dictyoasterina Hansf. (1)
Dictyodochium Sivan. (1)
Dictyopeltis Theiss. (6)
Dictyostomiopelta Viégas (1)
Dictyothyriella Speg. (1)

1143
Dictyothyrina Theiss. (1)
Dictyothyrium Theiss. (2)
Didymocyrtidium Vain. (2)
Didymolepta Munk (2)
Didymopleella Munk (3)
Diplochorina Gutner (1)
Dothichiza Lib. ex Roum. (15)
Dothideopsella Höhn. (1, but more epithets exist)
Dothivalsaria Petr. (1)
Dubitatio Speg. (1)
Echinothecium Zopf (2)
Elmerinula Syd. (1)
Epibelonium E. Müll. (1)
Eriomycopsis Speg. (13)
Eriothyrium Speg. (1, but more epithets exist)
Eupelte Syd. (5)
Excipulariopsis P.M. Kirk & Spooner (1)
Extrusothecium Matsush. (2)
Fusicladiella Höhn. (5)
Gibbera Fr. (ca. 28)
Gilletiella Sacc. & P. Syd. (3)
Globoa Bat. & H. Maia (2)
Globulina Speg. (1 fide Kirk et al. 2008)
Gloeodiscus Dennis (1)
Govindua Bat. & H. Maia (1)
Griggsia F. Stevens & Dalbey (1)
Halokirschsteiniothelia Boonmee & K.D. Hyde (1)
Hansfordiella S. Hughes (8)
Hansfordiellopsis Deighton (5)
Hansfordiopsis Bat. (1)
Harknessiella Sacc. (1)
Helminthopeltis Sousa da Câmara (1)
Heptameria Rehm & Thuem. (2)
Heptaster Cif., Bat. & Nascim. (3)
Heteroconium Petr. (21)
Heterosphaeriopsis Hafellner (1)
Hidakaea I. Hino & Katum. (2)
Hyalocrea Syd. & P. Syd. (4)
Hyaloscolecostroma Bat. & J. Oliveira (1)
Hyalosphaera F. Stevens (4)
Hyalotheles Speg. (1)
Hypobryon Döbbeler (7)
Hysteropsis Rehm (4)
Isomunkia Theiss. & Syd. (1)
Isthmospora F. Stevens (3)
Jaffuela Speg. (1)
Kabatia Bubák (ca. 10)
Keratosphaera H.P. Upadhyay (6)
Kriegeriella Höhn. (4)
Krishnamyces Hosag. (1)
Kullhemia P. Karst. (2)
Kusanobotrys P. Henn. (2)

1144
Lanatosphaera Matzer (2)
Lautitia S. Schatz (1)
Lazarenkoa Zerova (1)
Lembosiopeltis Bat. & J.L. Bezerra (2)
Leptomeliola Höhn. (13)
Letendraeopsis K.F. Rodriguez & Samuels (1)
Leveillella Theiss. & Syd. (1)
Leveillina Theiss. & Syd. (2)
Lichenotubeufia Etayo (5)
Licopolia Sacc., Syd. & P. Syd. (2)
Lignosphaeria Boonmee, Thambug. & K.D. Hyde (2)
Limaciniopsis Mend. (1)
Lineolata Kohlm. & Volkm.-Kohlm. (1)
Linopeltis I. Hino & Katum. (2)
Lophionema Sacc. (9)
Lucidascocarpa A. Ferrer, Raja & Shearer (1)
Macowaniella Doidgeb (2)
Maheshwaramyces Hosag. (2)
Maireella Syd. & Maire (ca. 5)
Malacaria Syd. (2)
Manginula G. Arnaud (ca. 5)
Marquesius L.B. Conç., R.F. Castañeda & Gusmão (1)
Massariola Füisting (2)
Maublancia G. Arnaud (1)
Melioliphila Speg. (7)
Mendoziopeltis Bat. (4)
Microcyclella Theiss. (1)
Microdothella Syd. & P. Syd. (2)
Monoblastiopsis R.C. Harris & C.A. Morse (2)
Monodictys S. Hughes (ca. 50)
Monorhizina Theiss. & Syd. (1)
Montagnella Speg. (9)
Moriolomyces Cif. & Tomas. (1)
Muricopeltis Viégas (1)
Muroia I. Hino & Katum. (1)
Mycocryptospora J. Reid & C. Booth (1)
Mycodidymella C.Z. Wei, Y. Harada & Katum. (1)
Mycoglaena Höhn. (16)
Mycoporellum Müll. Arg. (7)
Mycoporopsis Müll. Arg. (ca. 10)
Mycothyridium Petr. (30)
Myriangiopsis P. Henn. (2)
Myriostigmella G. Arnaud (1)
Mytilostoma P. Karst. (2)
Myxophora Döbbeler & Poelt (7)
Nannfeldtia Petr. (2)
Neodactylaria Guevara-Suarez, Deanna A. Sutton, Wiederh. & Gené (1)
Neopeckia Sacc. (1 fide Kirk et al. 2008)
Neosporidesmium Mercado & J. Mena (15)
Neottiosporina Subram. (11)
Neoventuria Syd. & P. Syd. (1)
Ocala Raja & Shearer (1)

1145
Oletheriostrigula Huhndorf & R.C. Harris (1)
Omphalospora Theiss. & Syd. (2)
Oncopodiella G. Arnaud ex Rifai (13)
Ophioirenina Sawada & W. Yamam. (1)
Ophiotrichum Kunze (3)
Otthia Nitschke ex Fuckel (11)
Parmulariella P. Henn. (1)
Paropodia Cif. & Bat. (1)
Passeriniella Berl. (7)
Passerinula Sacc. (1)
Pauahia F. Stevens (1)
Peltaster Syd. & P. Syd. (8)
Peltasterella Bat. & H. Maia (1)
Pendulispora M.B. Ellis (1)
Perischizon P. Syd. (3)
Peroschaeta Bat. & A.F. Vital (1)
Petrakina Cif. (3)
Petrakiopeltis Bat., A.F. Vital & Cif. (1)
Phacidina Höhn. (1)
Phaeocyrtidula Vain. (2)
Phaeopeltosphaeria Berl. & Peglion (2)
Phaeosclera Sigler, Tsuneda & J.W. Carmich. (1)
Phaeosperma Nitschke ex Fuckel (1)
Phaeostigme Syd. & P. Syd. (6)
Phaeotomasellia Katum. (1)
Phanerococculus Cif. (1)
Philobryon Döbbeler (1)
Philonectria Hara (3)
Phragmaspidium Bat. (3)
Phragmogibbera Samuels & Rogerson (3)
Phragmoscutella Woron. & Abramov ex Woron. (1)
Phragmosperma Theiss. & Syd. (1)
Phycorella Döbbeler (1)
Physalosporopsis Bat. & H. Maia (1)
Pirozynskia Subram. (1)
Placoasterina Toro (1)
Placodothis Syd. (1)
Placomelan Cif. (1)
Placosphaeria (De Not.) Sacc. (1, but several other epithets exist)
Plagiostromella Höhn. (1)
Plejobolus (E. Bommer et al.) O.E. Erikss. (1 or 2 species)
Plenotrichaius Bat. & Valle (1)
Pleomerium Speg. (1)
Pleotrichiella Sivan. (1)
Polycyclinopsis Bat., A.F. Vital & I.H. Lima (1)
Polyrhizon Theiss., Syd. & P. Syd. (2)
Polysporidiella Petr. (1)
Polystomellopsis F. Stevens (1)
Proliferosphaera T.P. Devi (1)
Pseudoarthrographis Crous & Thangavel (1)
Pseudomorfea Punith. (1)
Pseudopleospora Petr. (1)

1146
Punctillum Petr. & Syd. (1)
Pyrenobotrys Theiss. & Syd. (1)
Pyrenochium Link (1)
Pyrenocyclus Petr. (1)
Pyrenostigme Syd. (1)
Radulidium Arzanlou, W. Gams & Crous (3)
Rhizotexis Theiss. & Syd. (1)
Rhopographus Nitschke ex Fuckel (6)
Rosellinula R. Sant. (4)
Rosenscheldia Speg. (1)
Roumegueria (Sacc.) P. Henn. (1)
Rupestriomyces Lei Su, Li Y. Guo & Xing Z. Liu (3)
Sapucchaka K. Ramakr. (2)
Saxomyces L. Selbmann & D. Isola (2)
Scleroconidioma Tsuneda, Currah & Thormann (1)
Scolecobonaria Bat. (2)
Scolecoxyphium Cif. & Bat. (5)
Scolionema Theiss. & Syd. (1)
Semisphaeria K. Holm & L. Holm (1)
Septoidium G. Arnaud (ca. 7)
Shearia Petr. (2)
Shivamyces Hosag. (2)
Sivanesaniella Gawande & D.K. Agarwal (1)
Solicorynespora R.F. Castañeda & W.B. Kendr. (29)*
Soloacrosporiella Crous & M.J. Wingf. (1)
Spilodochium Syd. (4)
Spissiomyces Lei Su, Li Y. Guo & Xing Z. Liu (2)
Stegothyrium Höhn. (2)
Stephanotheca Syd. & P. Syd. (4)
Stigmatodothis Syd. & P. Syd. (1)
Stigmatophragmia Tehon & G.L. Stout (1)
Symphaster Theiss. & Syd. (1)
Taphrophila Scheuer (4)
Teichosporella (Sacc.) Sacc. (26)
Teratoschaeta Bat. & Fons.) (1)
Tetracrium Henn. (7)
Thalassoascus Ollivier (3)
Thelenidia Nyl. (1)
Thryptospora Petr. (1)
Tilakiella Srinivas. (1)
Tomeoa I. Hino (1)
Torulopsiella Bender (2)
Trematosphaeriopsis Elenkin (1)
Tretospora M.B. Ellis (8)
Trichodothella Petr. (1)
Trichodothis Theiss. & Syd. (3)
Trichometasphaeria Munk (8)
Trichothyriella Theiss. (1)
Troposporella P. Karst. (4)
Uredinophila Rossman (2)
Wentiomyces Koord. (ca. 50)
Westea H.J. Swart (1)

1147
Wettsteinina Höhn. (30)
Xenomeris Syd. (11)
Xenosporium Penz. & Sacc. (18)
Xenostomella Syd. (2)
Xylopezia Höhn. (ca. 3)
Yoshinagaia Henn. (1)
Yoshinagella Höhn. (4)

Eurotiomycetes Tehler ex O.E. Eriksson & K. Winka


Chaetothyriomycetidae Doweld
Chaetothyriales M.E. Barr
Chaetothyriaceae Hansf. ex M.E. Barr
Actinocymbe Höhn. (3)
Aithaloderma Syd. & P. Syd. (13)
Aphanophora Réblová & Unter. (1)
Arthrophiala (D.J. Soares, R.W. Barreto & U. Braun) W.S. Lisboa, Meir. Silva & R.W.
Barreto (1)*
Beelia F. Stevens & R.W. Ryan (3)
Camptophora Réblová & Unter. (2)*
Ceramothyrium Bat. & H. Maia (35)
Ceratocarpia Rolland (2)
Chaetothyriomyces Pereira-Carv., Inácio & Dianese (1)
Chaetothyrium Speg. (51)
Cyphellophoriella Crous & A.J. Sm. (1)
Euceramia Bat. & Cif. (3)
Longihyalospora D.S.Tennakoon, C.H Kuo & K.D. Hyde (2)
Microcallis Syd. (10)
Nullicamyces Crous (1)
Phaeosaccardinula P. Henn. (27)
Stanhughesia Constant. (1)
Treubiomyces Höhn. (7)
Vonarxia Bat. (2)
Yatesula Syd. & P. Syd. (2)

Coccodiniaceae Höhn. ex O.E. Erikss.


Coccodinium A. Massal. (4)
Dennisiella Bat. & Cif. (= Microxiphium (Harv. ex Berk. & Desm.) Thüm.) (9)
Limacinula Höhn. (17)

Cyphellophoraceae Réblová & Unter.


Anthopsis Fil. March., A. Fontana & Luppi Mosca (2)*
Cyphellophora G.A. de Vries (25)*

Epibryaceae S. Stenroos & Gueidan


Epibryon Döbbeler (ca. 40)

Herpotrichiellaceae Munk
Aculeata W. Dong, H. Zhang & K.D. Hyde (1)*
Brycekendrickomyces Crous & M.J. Wingf. (1)
Capronia Sacc. (ca. 81)
Cladophialophora Borelli (35)*
Exophiala J.W. Carmich. (51)*

1148
Fonsecaea Negroni (8)*
Marinophialophora J.F. Li, Phook. & K.D. Hyde (1)
Melanoctona Qing Tian, Doilom & K.D. Hyde (1)
Metulocladosporiella Crous, Schroers, J.Z. Groenew., U. Braun & K. Schub. (6)
Minimelanolocus R.F. Castañeda & Heredia (33)*
Phialophora Medlar (7)*
Pleomelogramma Speg. (2)
Rhinocladiella Nannf. (17)
Sorocybe Fr. (3)
Thysanorea Arzanlou, W. Gams & Crous (2)*
Veronaea Cif. & Montemart. (20)

Lyrommataceae Lücking
Lyromma Bat. (7)

Microtheliopsidaceae O.E. Erikss.


Microtheliopsis Müll. Arg. (4)

Paracladophialophoraceae Crous
Paracladophialophora Crous (2)*

Pyrenotrichaceae Zahlbr
Pyrenothrix Riddle (2)
Neophaeococcomyces Crous & M.J. Wingf. (2)

Trichomeriaceae Chomnunti & K.D. Hyde (= Strelitzianaceae Crous & M.J. Wingf.)
Arthrocladium Papendorf (4)*
Bradymyces Hubka, Réblová, Selbmann & M. Kolařík (3)*
Knufia L.J. Hutchison & Unter. (13)*
Lithohypha Selbmann & Isola (1)
Lithophila Selbmann & Isola (1)*
Neostrelitziana Crous & M.J. Wingf. (1)
Strelitziana Arzanlou & Crous (8)
Trichomerium Speg. (28)

Chaetothyriales genera incertae sedis


Atrokylindriopsis Y.R. Ma & X.G. Zhang (1)
Bacillicladium Hubka, Réblová & Thureborn (1)*
Lichenodiplis Dyko & D. Hawksw. (= Laeviomyces D. Hawksw.) (13)
Lichenodiplisiella S.Y. Kondr. & Kudratov (1)
Melnikomyces Crous & U. Braun (1)
Minutoexcipula V. Atienza & D. Hawksw. (7)
Muellerella Hepp (14)*
Pleostigma Kirschst. (9)
Sarcinomyces Lindner (5)
Uncispora R.C. Sinclair & Morgan-Jones (3)

Phaeomoniellales K.H. Chen, A.E. Arnold, Gueidan & Lutzoni


Celotheliaceae Lücking, Aptroot & Sipman (= Phaeomoniellaceae P.M. Kirk)
Aequabiliella Crous (1)
Celerioriella Crous (3)
Celothelium A. Massal. (8)

1149
Minutiella Crous (1)
Moristroma A.I. Romero & Samuels (4)
Neophaeomoniella Rooney-Latham & Crous (3)
Paraphaeomoniella Crous (1)
Phaeomoniella Crous & W. Gams (2)
Pseudophaeomoniella Nigro, Antelmi & Crous (2)
Xenocylindrosporium Crous & Verkley (1)

Pyrenulales Fink ex D. Hawksw. & O.E. Erikss.


Pyrenulaceae Rabenh.
Anthracothecium Hampe ex A. Massal. (5)
Blastodesmia A. Massal. (1)
Clypeopyrenis Aptroot (2)
Distopyrenis Aptroot (8)
Granulopyrenis Aptroot (6)
Lithothelium Müll. Arg. (28)
Mazaediothecium Aptroot (4)
Pyrenographa Aptroot (1)
Pyrenowilmsia R.C. Harris & Aptroot (1)
Pyrenula Ach. (= Heufleridium Müll. Arg.; = Stromatothelium Trevis.) (ca. 225)
Pyrgillus Nyl. (8)
Sulcopyrenula H. Harada (5)

Pyrenulales genera incertae sedis


Rhaphidicyrtis Vain. (1)
Xenus Kohlm. & Volkm.-Kohlm. (1)

Verrucariales Mattick ex D. Hawksw. & O.E. Erikss.


Adelococcaceae Triebel
Adelococcus Theiss. & Syd. (4)
Pseudopyrenidium Nav.-Ros., Zhurb. & Cl. Roux (1)
Sagediopsis Sacc. ex Vain. (10)

Sarcopyreniaceae Nav.-Ros. & Cl. Roux


Sarcopyrenia Nyl. (11)

Verrucariaceae Zenker
Agonimia Zahlbr. (ca. 20)
Anthracocarpon Breuss (1)
Atla S. Savić & Tibell (10)
Awasthiella Kr.P. Singh (1)
Bagliettoa A. Massal. (17)
Bellemerella Nav.-Ros. & Cl. Roux (4)
Catapyrenium Flot. (6)
Clauzadella Nav.-Ros. & Cl. Roux (1)
Clavascidium Breuss (9)
Dermatocarpon Eschw. (20)
Endocarpon Hedw. (ca. 75)
Flakea O.E. Erikss. (1)
Glomerilla Norman (1)
Haleomyces D. Hawksw. & Essl. (1)
Halospora (Zschacke) Tomas. & Cif. (4)

1150
Henrica de Lesd. (4)
Heterocarpon Müll. Arg. (1)
Heteroplacidium Breuss (12)
Hydropunctaria C. Keller, Gueidan & Thüs (8)
Involucropyrenium Breuss (9)
Mastodia Hook.f. & Harv. (= Turgidosculum Kohlm. & E. Kohlm.) (5)
Moriola Norman (ca. 15)
Neocatapyrenium H. Harada (5)
Normandina Nyl. (= Lauderlindsaya J.C. David & D. Hawksw.) (3)
Norrlinia Theiss. & Syd. (2)
Parabagliettoa Gueidan & Cl. Roux (3)
Phaeospora Hepp ex Stein (14)
Phylloblastia Vain. (12)
Placidiopsis Beltr. (20)
Placidium A. Massal. (28)
Placocarpus Trevis. (5)
Placopyrenium Breuss (22)
Placothelium Müll. Arg. (1)
Plurisperma Sivan. (1)
Polyblastia A. Massal. (ca. 40 + ca. 50 orphaned)
Psoroglaena Müll. Arg. (17)
Rhabdopsora Müll. Arg. (2)
Scleropyrenium H. Harada (2)
Servitia M.S. Christ. & Alstrup (1)
Spheconisca (Norman) Norman (ca. 20)
Sporodictyon A. Massal. (5)
Staurothele Norman (ca. 40)
Telogalla Nik. Hoffm. & Hafellner (2)
Thelidiopsis Vain. (4)
Thelidium A. Massal. (ca. 50 + ca. 50 orphaned)
Trimmatothele Norman ex Zahlbr. (3)
Verrucaria Schrad. (ca. 300)
Verrucula J. Steiner (22)
Verruculopsis Gueidan, Nav.-Ros. & Cl. Roux (ca. 10)
Wahlenbergiella Gueidan & Thüs (3)
Willeya Müll. Arg. (12)

Verrucariales genera incertae sedis


Botryolepraria Canals, Hern.-Mar., Gómez-Bolea & Llimona (2)
Gemmaspora D. Hawksw. & Halici (1)
Kalbiana Henssen (1)
Merismatium Zopf (10)

Chaetothyriomycetidae family incertae sedis


Rhynchostomataceae Winka & O.E. Erikss.
Rhynchomeliola Speg. (3)
Rhynchostoma P. Karst. (23)

Coryneliomycetidae A.R. Wood, Damm, J.Z. Groenew., Cheew. & Crous


Coryneliales Seaver & Chardon
Coryneliaceae Sacc. ex Berl. & Voglino
Caliciopsis Peck (36)

1151
Corynelia Ach. (16)
Coryneliopsis Butin (2)
Coryneliospora Fitzp. (2)
Fitzpatrickella Benny, Samuelson & Kimbr. (1)
Lagenulopsis Fitzp. (1)
Tripospora Sacc. ex Berl. & Vogl. (5)

Eremascaceae Engl. & E. Gilg


Eremascus Eidam (2)

Eurotiomycetidae Geiser & Lutzoni


Arachnomycetales Gibas, Sigler & Currah
Arachnomycetaceae Gibas, Sigler & Currah
Arachnomyces Massee & E.S. Salmon (10)
Onychocola Sigler (4)

Eurotiales G.W. Martin ex Benny & Kimbr.


Aspergillaceae Link (= Monascaceae J. Schröt.)
Aspergillago Samson, Houbraken & Frisvad (1)
Aspergillus P. Micheli ex Haller (428)
Dichlaena Durieu & Mont. (4)
Hamigera Stolk & Samson (9)
Leiothecium Samson & Mouch. (2)
Monascus Tiegh. (38)
Penicillago Guevara-Suarez, Gené & Dania García (1)
Penicilliopsis Solms (15)
Penicillium Link (467)
Phialomyces P.C. Misra & P.H.B. Talbot (5)
Pseudopenicillium Guevara-Suarez, Cano & Guarro (2)
Sclerocleista Subram. (2)
Sclerocleista Subram. (2)
Xerochrysium Pitt (2)
Xeromyces Fraser (1)

Elaphomycetaceae Tul. ex Paol.


Elaphomyces Nees (101)
Pseudotulostoma O.K. Miller & T. Henkel (2)

Thermoascaceae Apinis
Paecilomyces Bainier (10)
Thermoascus Miehe (5)

Trichocomaceae E. Fisch.
Chaetotheca Zukal (2)
Dendrosphaera Pat. (1)
Rasamsonia Houbraken & Frisvad (11)
Sagenomella W. Gams (8)
Talaromyces C.R. Benj. (149)
Thermomyces Tsikl. (6)
Trichocoma Jungh. (2)

Onygenales Cif. ex Benny & Kimbr.

1152
Ajellomycetaceae Unter., J.A. Scott & Sigler
Blastomyces Gilchrist & W.R. Stokes (=Ajellomyces McDonough & A.L Lewis; Emmonsia
Cif. & Montemart.) (9)
Emmonsiellopsis Y. Marín, Stchigel, Guarro & Cano (2)
Emergomyces Dukik, Sigler & de Hoog (5)*
Histoplasma Darling (4 epithets in Index Fungorum 2020)
Lacazia Taborda, V.A. Taborda & McGinnis (1)
Paracoccidioides F.P. Almeida (6)

Arthrodermataceae Currah
Arthroderma Curr. & Berk. (32)
Ctenomyces Eidam (7)
Epidermophyton Sabour. (1)
Guarromyces Y Gräser & de Hoog (1)
Lophophyton Matr. & Dassonv. (1)
Microsporum Gruby (3)
Nannizzia Stockdale (9)
Paraphyton Y Gräser, Dukik & de Hoog (3)
Shanorella R.K. Benj. (1)
Trichophyton Malmsten (16)

Ascosphaeraceae L.S. Olive & Spiltoir


Arrhenosphaera Stejskal (1)
Ascosphaera L.S. Olive & Spiltoir (27)
Bettsia Skou (2)

Gymnoascaceae Baran.
Aciascus Doweld (1)
Amaurascopsis Guarro, Gené & De Vroey (1)
Arachniotus J. Schröt. (21)
Gymnascella Peck (9)
Gymnoascoideus G.F. Orr, K. Roy & G.R. Ghosh (1)
Gymnoascus Baran. (=Narasimhella Thirum. & P.N. Mathur) (26)
Kraurogymnocarpa Udagawa & Uchiyama (1)
Mallochia Arx & Samson (4)
Oncocladium Wallr. (1)
Orromyces Sur & G.R. Ghosh (1)

Nannizziopsidaceae Guarro, Stchigel, Deanna A. Sutton & Cano


Nannizziopsis Currah (16)

Onygenaceae Berk.
Amauroascus J. Schröt. (ca. 15)
Aphanoascus Zukal (18)
Apinisia La Touche (3)
Arachnotheca Arx (1)
Ascocalvatia Malloch & Cain (1)
Auxarthron G.F. Orr & Kuehn (13)
Auxarthronopsis Rah. Sharma, Y. Gräser & S.K. Singh (2)
Bifidocarpus Cano, Guarro & R.F. Castañeda (2)
Byssoonygena Guarro, Punsola & Cano (1)
Castanedomyces Cano, L.B. Pitarch & Guarro (1)

1153
Chlamydosauromyces Sigler, Hambl. & Paré (1)
Chrysosporium Corda (66)
Coccidioides G.W. Stiles (6)
Kuehniella G.F. Orr (2)
Leucothecium Arx & Samson (3)
Malbranchea Sacc. (23)
Myotisia Kubátová, M. Kolařík & Hubka (1)
Myriodontium Samson & Polon. (1)
Neoarachnotheca Ulfig, Cano & Guarro (1)
Neogymnomyces G.F. Orr (2)
Onygena Pers. (10)
Ophidiomyces Sigler, Hambl. & Paré (1)
Paranannizziopsis Sigler (4)
Pectinotrichum Varsavsky & G.F. Orr (2)
Polytolypa J.A. Scott & Malloch (1)
Pseudoamauroascus Cano, M. Solé & Guarro (1)
Renispora Sigler & J.W. Carmich. (2)
Sporendonema Desm. (2)
Testudomyces Cano, M. Solé & Guarro (1)
Uncinocarpus Sigler & G.F. Orr (2)
Xanthothecium Arx & Samson (1)

Spiromastigaceae Guarro, Cano & Stchigel


Pseudospiromastix Guarro, Stchigel & Cano (1)
Sigleria Hirooka, Tanney & Seifert (2)
Spiromastigoides Doweld (8)
Spiromastix Kuehn & G.F. Orr (5)

Onygenales genera incertae sedis


Arthropsis Sigler, M.T. Dunn & J.W. Carmich. (4)
Ovadendron Sigler & J.W. Carmich. (1)*
Sphaerosporium Schwein. sensu lato (2)*

Eurotiomycetidae genera incertae sedis


Azureothecium Matsush. (1)
Calyptrozyma Boekhout & Spaay (1)
Pisomyxa Corda (1)
Samarospora Rostr. (1)
Veronaia Benedek (2)

Mycocaliciomycetidae Tibell
Mycocaliciales Tibell & Wedin
Mycocaliciaceae A.F.W. Schmidt (= Sphinctrinaceae M. Choisy)
Brunneocarpos Giraldo & Crous (1)
Chaenothecopsis Vain. (ca. 40)
Mycocalicium Vain. ex Reinke (12)
Phaeocalicium A.F.W. Schmidt (11)
Pyrgidium Nyl. (3)
Sphinctrina Fr. (ca. 9)
Stenocybe (Nyl.) Körb. (14)

Sclerococcomycetidae Réblová, Unter. & W. Gams

1154
Sclerococcales Réblová, Unter. & W. Gams
Dactylosporaceae Bellem. & Hafellner (= Sclerococcaceae Réblová, Unter. & W. Gams)
Cylindroconidiis H. Zhang & X.D. Yu (1)
Fusichalara S. Hughes & Nag Raj (5)*
Longimultiseptata H. Zhang & W. Dong (2)
Rhopalophora Réblová, Unter. & W. Gams (1)
Sclerococcum Fr. (= Dactylospora Körb.) (ca. 80)*

Eurotiomycetes genus incertae sedis


Neocladophialophora Crous & R.K. Schumach. (1)

Geoglossomycetes Zheng Wang, C.L. Schoch & Spatafora


Geoglossales Zheng Wang, C.L. Schoch & Spatafora
Geoglossaceae Corda
Geoglossum Pers. (40)
Glutinoglossum Hustad, A.N. Mill., Dentinger & P.F. Cannon (13)
Hemileucoglossum Arauzo (5)
Leucoglossum S. Imai (2)
Maasoglossum K.S. Thind & R. Sharma (2)
Sabuloglossum Hustad, A.N. Mill., Dentinger & P.F. Cannon (1)
Trichoglossum Boud. (19)

Geoglossomycetes genera incertae sedis


Nothomitra Maas Geest. (3)*
Sarcoleotia S. Ito & S. Imai (3)*

Laboulbeniomycetes Engler
Herpomycetales Haelew. & Pfister*
Herpomycetaceae I.I. Tav.
Herpomyces Thaxt. (26)

Laboulbeniales Lindau
Ceratomycetaceae S. Colla
Autoicomyces Thaxt. (28)
Ceratomyces Thaxt. (32)
Drepanomyces Thaxt. (1)
Eusynaptomyces Thaxt. (5)
Helodiomyces F. Picard (1)
Phurmomyces Thaxt. (1)
Plectomyces Thaxt. (1)
Rhynchophoromyces Thaxt. (8)
Synaptomyces Thaxt. (1)
Tettigomyces Thaxt. (16)
Thaumasiomyces Thaxt. (3)
Thripomyces Speg. (2)

Euceratomycetaceae I.I. Tav.


Cochliomyces Speg. (2)
Colonomyces R.K. Benj. (2)
Euceratomyces Thaxt. (1)
Euzodiomyces Thaxt. (2)
Pseudoecteinomyces W. Rossi (1)

1155
Laboulbeniaceae G. Winter
Acallomyces Thaxt. (3)
Acompsomyces Thaxt. (6)
Acrogynomyces Thaxt. (6)
Amorphomyces Thaxt. (15)
Amphimyces Thaxt. (1)
Apatelomyces Thaxt. (1)
Apatomyces Thaxt. (1)
Aphanandromyces W. Rossi (1)
Aporomyces Thaxt. (11)
Arthrorhynchus Kolen. (3)
Asaphomyces Thaxt. (2)
Autophagomyces Thaxt. (17)
Benjaminiomyces I.I. Tav. (4)
Blasticomyces I.I. Tav. (3)
Bordea Maire (15)
Botryandromyces I.I. Tav. & T. Majewski (2)
Camptomyces Thaxt. (8)
Cantharomyces Thaxt. (29)
Capillistichus Santam. (1)
Carpophoromyces Thaxt. (1)
Cesariella W. Rossi & Santam. (1)
Chaetarthriomyces Thaxt. (3)
Chaetomyces Thaxt. (2)
Chitonomyces Peyr. (ca. 98)
Clematomyces Thaxt. (5)
Clonophoromyces Thaxt. (2)
Columnomyces R.K. Benj. (1)
Compsomyces Thaxt. (7)
Coreomyces Thaxt. (22)
Corethromyces Thaxt. (ca. 85)
Corylophomyces R.K. Benj. (5)
Cryptandromyces Thaxt. (= Peyerimhoffiella Maire) (19)
Cucujomyces Speg. (20)
Cupulomyces R.K. Benj. (= Balazucia R.K. Benj.) (1)
Dermapteromyces Thaxt. (1)
Diandromyces Thaxt. (2)
Diaphoromyces Thaxt. (5)
Diclonomyces Thaxt. (3)
Dimeromyces Thaxt. (118)
Dimorphomyces Thaxt. (32)
Dioicomyces Thaxt. (32)
Diphymyces I.I. Tav. (25)*
Diplomyces Thaxt. (3)
Diplopodomyces W. Rossi & Balazuc (6)
Dipodomyces Thaxt. (2)
Distolomyces Thaxt. (3)
Dixomyces I.I. Tav. (14)
Ecteinomyces Thaxt. (1)
Enarthromyces Thaxt. (1)
Eucantharomyces Thaxt. (26)
Euhaplomyces Thaxt. (1)

1156
Eumonoicomyces Thaxt. (2)
Euphoriomyces Thaxt. (15)
Filariomyces Shanor (1)
Gloeandromyces Thaxt. (4)
Haplomyces Thaxt. (3)
Hesperomyces Thaxt. (8)*
Histeridomyces Thaxt. (6)
Homaromyces R.K. Benj. (1)
Hydraeomyces Thaxt. (1)
Hydrophilomyces Thaxt. (12)
Idiomyces Thaxt. (1)
Ilyomyces F. Picard (2)
Ilytheomyces Thaxt. (15)
Kainomyces Thaxt. (3)
Kleidiomyces Thaxt. (4)
Kruphaiomyces Thaxt. (1)
Kyphomyces I.I. Tav. (14)
Laboulbenia Mont. & C.P. Robin (= Scalenomyces I.I. Tav.) (ca. 633)*
Limnaiomyces Thaxt. (3)
Majewskia Y.B. Lee & Sugiyama (1)
Meionomyces Thaxt. (6)
Microsomyces Thaxt. (2)
Mimeomyces Thaxt. (16)
Misgomyces Thaxt. (4)
Monandromyces R.K. Benj. (11)
Monoicomyces Thaxt. (47)
Nanomyces Thaxt. (48)
Neohaplomyces R.K. Benj. (3)
Nycteromyces Thaxt. (2)
Opilionomyces Santam., Enghoff, Gruber & Reboleira (1)*
Ormomyces I.I. Tav. (1)
Osoriomyces Terada (1)
Parvomyces Santam. (1)
Peyritschiella Thaxt. (47)
Phalacrichomyces R.K. Benj. (2)
Phaulomyces Thaxt. (14)
Picardella I.I. Tav. (2)
Polyandromyces Thaxt. (1)
Polyascomyces Thaxt. (1)
Porophoromyces Thaxt. (1)
Prolixandromyces R.K. Benj. (20)
Pselaphidomyces Speg. (1)
Rhachomyces Thaxt. (ca. 75)
Rhipidiomyces Thaxt. (1)
Rhizomyces Thaxt. (10)
Rhizopodomyces Thaxt. (7)
Rickia Cavara (144)
Rodaucea W. Rossi & Santam. (2)
Rossiomyces R.K. Benj. (1)
Sandersoniomyces R.K. Benj. (1)
Scaphidiomyces Thaxt. (5)
Scelophoromyces Thaxt. (1)

1157
Scepastocarpus Santam. (1)
Siemaszkoa I.I. Tav. & Maj. (7)
Smeringomyces Thaxt. (4)
Sphaleromyces Thaxt. (3)
Stemmatomyces Thaxt. (2)
Stichomyces Thaxt. (7)
Stigmatomyces H. Karst. (= Fanniomyces T. Majewski) (150)
Sugiyamaemyces I.I. Tav. & Balazuc (1)
Symplectromyces Thaxt. (3)
Sympodomyces R.K. Benj. (1)
Synandromyces Thaxt. (9)
Tavaresiella T. Majewski (4)
Teratomyces Thaxt. (11)
Tetrandromyces Thaxt. (6)
Thaxterimyces Santam., Reboleira & Enghoff (1)
Trenomyces Chatton & F. Picard (11)
Triainomyces W. Rossi & A. Weir (1)
Triceromyces T. Majewski (5)
Trochoideomyces Thaxt. (1)
Troglomyces S. Colla (8)
Zeugandromyces Thaxt. (4)
Zodiomyces Thaxt. (4)

Laboulbeniales genera incertae sedis


Cainomyces Thaxt. (1)
Coreomycetopsis Thaxt. (1)
Gliocephalis Matr. (2)

Pyxidiophorales P.F. Cannon


Pyxidiophoraceae Arnold
Mycorhynchidium Malloch & Cain (1)
Pleurocatena G. Arnaud ex Aramb., Gamundí, W. Gams & G.R.W. Arnold (3)
Pyxidiophora Bref. & Tavel (17)

Laboulbeniomycetes genus incertae sedis


Laboulbeniopsis Thaxt. (1)

Lecanoromycetes O.E. Erikss. & Winka


Acarosporomycetidae V. Reeb, Lutzoni & Cl. Roux
Acarosporales V. Reeb, Lutzoni & Cl. Roux
Acarosporaceae Zahlbr.
Acarospora A. Massal. (200)
Caeruleum Arcadia (2)
Glypholecia Nyl. (1)
Lithoglypha Brusse (1)
Myriospora Nägeli ex Uloth (9)
Pleopsidium Körb. (4)
Polysporina Vězda (10)
Sarcogyne Flot. (28)
Thelocarpella Nav.-Ros. & Cl. Roux (1)
Timdalia Hafellner (1)
Trimmatothelopsis Zschacke (1)

1158
Eigleraceae Hafellner
Eiglera Hafellner (2)

Lecanoromycetidae P.M. Kirk, P.F. Cannon, J.C. David & Stalpers ex Miądl., Lutzoni &
Lumbsch
Caliciales Bessey
Caliciaceae Chevall.
Acolium (Ach.) Gray (5)
Acroscyphus Lév. (1)
Allocalicium M. Prieto & Wedin (1)
Amandinea M. Choisy ex Scheid. & M. Mayrhofer (35)
Australiaena Matzer, H. Mayrhofer & Elix (1)
Baculifera Marbach (14)
Buellia De Not. (= Dirinastrum Müll. Arg.) (300)
Caliciella Vain. (1)
Calicium Pers. (= Cyphelium Ach.) (ca. 30)
Chrismofulvea Marbach (4)
Ciposia Marbach (1)
Cratiria Marbach (ca. 20)
Culbersonia Essl. (1)*
Dermatiscum Nyl. (3)
Dermiscellum Hafellner, H. Mayrhofer & Poelt (1)
Dimelaena Norman (10)
Diploicia A. Massal. (ca. 12)
Diplotomma Flot. (ca. 30)
Dirinaria (Tuck.) Clem. (ca. 35)
Endohyalina Marbach (10)
Fluctua Marbach (1)
Gassicurtia Fée (30)
Hypoflavia Marbach (3)
Monerolechia Trevis. (4)
Orcularia (Malme) Kalb & Giralt (4)
Pseudothelomma M. Prieto & Wedin (2)
Pyxine Fr. (ca. 75)
Redonia C.W. Dodge (2)
Santessonia Hale & Vobis (10)
Sculptolumina Marbach (4)
Sphinctrinopsis Woron. (1)
Stigmatochroma Marbach (9)
Tetramelas Norman (16)
Texosporium Nádv. ex Tibell & Hofsten (1)
Thelomma A. Massal. (5)
Tholurna Norman (1)

Physciaceae Zahlbr.
Anaptychia Körb. (ca. 15)
Coscinocladium Kunze (2)
Heterodermia Trevis. (ca. 90)
Hyperphyscia Müll. Arg. (9)
Kashiwadia S.Y. Kondr. (1)
Leucodermia Kalb (10)
Mischoblastia A. Massal. (3)

1159
Mobergia H. Mayrhofer & Sheard (1)
Oxnerella S.Y. Kondr., Lőkös & Hur (1)
Phaeophyscia Mob. (66)
Phaeorrhiza H. Mayrhofer & Poelt (2)
Physcia (Schreb.) Michaux (ca. 80)
Physciella Essl. (4)
Physconia Poelt (ca. 25)
Polyblastidium Kalb (18)
Rinodina (Ach.) Gray (ca. 300)
Rinodinella H. Mayrhofer & Poelt (6)
Tornabea Oesth. (1)

Lecanorales Nannf.
Bruceomycetaceae Rikkinen & A.R. Schmidt
Bruceomyces Rikkinen (4)
Resinogalea Rikkinen & A.R. Schmidt (1)*

Catillariaceae Hafellner
Austrolecia Hertel (1)
Catillaria A. Massal. (ca. 30 + several orphaned names)
Placolecis Trevis. (1)
Solenopsora A. Massal. (11)
Xanthopsorella Kalb & Hafellner (1)

Cladoniaceae Zenker (= Squamarinaceae Hafellner, = Stereocaulaceae Chevall.)*


Calathaspis I.M. Lamb & W.A. Weber (1)
Carassea S. Stenroos (1)
Cetradonia J.C. Wei & Ahti (1)
Cladia Nyl. (ca. 27)
Cladonia Hill ex P. Browne (ca. 500)
Gymnoderma Nyl. (3)
Herteliana P. James (3)
Hertelidea Printzen & Kantvilas (6)
Heteromyces Müll. Arg. (1)
Lepraria Ach. (76)
Metus D.J. Galloway & P. James (3)
Notocladonia S. Hammer (2)
Paralecia Brackel, Greiner, Peršoh & Rambold (1)
Pilophorus Th. Fr. (17)
Pulchrocladia S. Stenroos, Pino-Bodas, Lumbsch & Ahti (3)
Pycnothelia Duf. (2)
Sphaerophoropsis Vain. (2)
Squamarina Poelt (25)
Squamella S. Hammer (1)
Stereocaulon Hoffm. (ca. 140)
Thysanothecium Mont. & Berk. (3)
Xyleborus R.C. Harris & Ladd (1)

Gypsoplacaceae Timdal
Gypsoplaca Timdal (5)

Haematommataceae Hafellner

1160
Haematomma A. Massal. (ca. 50)

Lecanoraceae Körb. (= Carbonicolaceae Bendiksby & Timdal)


Adelolecia Hertel & Hafellner (4)*
Ameliella Fryday & Coppins (2)
Bryodina Hafellner (2)
Bryonora Poelt (11)
Carbonicola Bendiksby & Timdal (3)
Cladidium Hafellner (2)
Claurouxia D. Hawksw. (1)
Clauzadeana Cl. Roux (1)
Edrudia W.P. Jordan (1)
Frutidella Kalb (3)*
Huea C.W. Dodge & G.E. Baker (=Carbonea (Hertel) Hertel) (20)
Japewia Tønsberg (3)*
Japewiella Printzen (7)
Lecanora Ach. (ca. 550)
Lecidella Körb. (80)
Maronina Hafellner & R.W. Rogers (6)
Maronora Kalb & Aptroot (1)
Miriquidica Hertel & Rambold (30)
Myriolecis Clements (43)
Palicella Rodr. Flakus & Printzen (4)
Protoparmeliopsis Choisy (= Sedelnikovaea S.Y. Kondr., M.H. Jeong & Hur) (ca. 20)
Psorinia Gotth. Schneid. (2)
Punctonora Aptroot (2)
Pyrrhospora Körb. (7)
Rhizoplaca Zopf (11)
Sagema Poelt & Grube (1)
Traponora Aptroot (8)
Vainionora Kalb (9)

Malmideaceae Kalb, Rivas Plata & Lumbsch


Cheiromycina B. Sutton (4)
Crustospathula Aptroot (4)*
Kalbionora Sodamuk, S.D. Leav. & Lumbsch (1)
Malmidea Kalb, Rivas Plata & Lumbsch (52)
Savoronala Ertz, Eb. Fisch., Killmann, Razafindr. & Sérus (1)
Sprucidea M.Cáceres, Aptroot & Lücking (4)
Zhurbenkoa Flakus, Etayo, Pérez-Ortega & Rodr. Flakus (3)

Megalariaceae Hafellner
Catillochroma Kalb (2)
Megalaria Hafellner (ca. 30)*

Parmeliaceae Zenker
Alectoria Ach. (= Gowardia Halonen, Myllys, Velmala & Hyvärinen) (9)
Allantoparmelia (Vain.) Essl. (3)
Anzia Stizenb. (34)
Arctoparmelia Hale (5)
Asahinea W.L. Culb. & C.F. Culb. (2)
Austromelanelixia Divakar, A. Crespo & Lumbsch (5)

1161
Austroparmelina A. Crespo, Divakar & Elix (13)
Brodoa Goward (3)
Bryocaulon Kärnefelt (4)
Bryoria Brodo & D. Hawksw. (ca. 52)
Bulbothrix Hale (62)
Canoparmelia Elix & Hale (35)
Cetraria Ach. (= Allocetraria Kurok. & M.J. Lai, = Cetrariella Kärnefelt & Thell, =
Usnocetraria M.J. Lai & J.C. Wei, = Vulpicida Mattson & M.J. Lai) (35)
Cetrelia W.L. Culb. & C.F. Culb. (18)
Coelopogon Brusse & Karnefelt (2)
Cornicularia (Schreb.) Ach. (1)
Dactylina Nyl. (2)
Davidgallowaya Aptroot (1)
Dolichousnea (Y. Ohmura) Articus (3)
Emodomelanelia Divakar & A. Crespo (1)
Esslingeriana Hale & M.J. Lai (1)
Eumitria Stirt. (13)
Evernia Ach. (10)
Everniopsis Nyl. (1)
Flavoparmelia Hale (32)
Flavopunctelia Hale (5)
Himantormia I.M. Lamb (2)
Hypogymnia (Nyl.) Nyl. (90)
Hypotrachyna (Vain.) Hale (262)
Imshaugia F.C. Mey. (1)
Letharia (Th. Fr.) Zahlbr. (9)
Lethariella (Motyka) Krog (11)
Masonhalea Kärnefelt (2)*
Melanelia Essl. (2)
Melanelixia O. Blanco, A. Crespo, Divakar, Essl., D. Hawksw. & Lumbsch (11)
Melanohalea O. Blanco, A. Crespo, Divakar, Essl., D. Hawksw. & Lumbsch (22)
Menegazzia A. Massal. (70)
Montanelia Divakar, A. Crespo, Wedin & Essl. (5)
Myelochroa (Asahina) Elix & Hale (30)
Neoprotoparmelia Garima Singh, Lumbsch & I. Schmitt (14)
Nephromopsis Müll. Arg. (= Ahtiana Goward; = Arctocetraria Kärnefelt & Thell; =
Cetrariopsis Kurok.; = Flavocetraria Kärnefelt & Thell; = Flavocetrariella D.D. Awasthi; =
Kaernefeltia Thell & Goward; = Tuckermanella Essl.; = Tuckermannopsis Gyeln.) (62)
Nesolechia A. Massal. (ca. 2)
Nipponoparmelia (Kurok.) K.H. Moon, Y. Ohmura & Kashiw. (4)
Nodobryoria Common & Brodo (3)
Notoparmelia A. Crespo, Ferencová & Divakar (16)
Omphalodium Meyen & Flot. (4)
Omphalora T.H. Nash & Hafellner (1)
Oropogon Th. Fr. (42)
Pannoparmelia (Müll. Arg.) Darb. (5)
Parmelia Ach (43)
Parmelina Hale (10)
Parmelinella Elix & Hale (8)
Parmeliopsis (Nyl.) Nyl. (3)
Parmotrema A. Massal. (= Crespoa (D. Hawksw.) Lendemer & B.P. Hodk.) (255)
Parmotremopsis Elix & Hale (2)

1162
Phacopsis Tul. (10)
Platismatia W.L. Culb. & C.F. Culb. (11)
Pleurosticta Petr. (2)
Protoparmelia M. Choisy (11)
Protousnea (Motyka) Krog (8)
Pseudephebe M. Choisy (2)
Pseudevernia Zopf (4)
Pseudoparmelia Lynge (15)
Psiloparmelia Hale (13)
Punctelia Krog (48)
Relicina (Hale & Kurok.) Hale (59)
Remototrachyna Divakar & A. Crespo (19)
Raesaenenia D. Hawksw. (1)
Sulcaria Bystr. (5)
Usnea Dill. ex Adans. (355)
Xanthoparmelia (Vain.) Hale (822)

Pilocarpaceae Zahlbr.
Aquacidia Aptroot (3)*
Badimiella Malcolm & Vězda (1)
Baflavia Lücking (1)
Bapalmuia Sérus. (22)
Barubria Vězda (2)
Brasilicia Lücking, Kalb & Serus. (6)
Bryogomphus Lücking, W.R. Buck, Sérus. & L.I. Ferraro (1)
Byssolecania Vain. (7)
Byssoloma Trevis. (60)
Calopadia Vězda (27)
Calopadiopsis Lücking & R. Sant. (2)
Eugeniella Lücking, Sérus. & Kalb (11)
Fellhanera Vězda (ca. 100)
Fellhaneropsis Sérus. & Coppins (9)
Kantvilasia P.M. McCarthy, Elix & Sérus. (1)
Lasioloma R. Sant. (9)
Leimonis R.C. Harris (2)
Loflammia Vězda (5)
Loflammiopsis Lücking & Kalb (1)
Logilvia Vězda (1)
Micarea Fr. (102)
Podotara Malcolm & Vězda (1)
Pseudocalopadia Lücking (1)
Roccellinastrum Follmann (7)
Schadonia Körb. (4)*
Septotrapelia Aptroot & Chaves (4)
Sporopodiopsis Sérus. (2)
Sporopodium Mont. (24)
Szczawinskia A. Funk (5)
Tapellaria Müll. Arg. (23)
Tapellariopsis Lücking (1)

Psilolechiaceae S. Stenroos, Miądl. & Lutzoni


Psilolechia A. Massal. (4)

1163
Psoraceae Zahlbr.
Brianaria S. Ekman & M. Svensson (4)
Glyphopeltis Brusse (1)
Protoblastenia (Zahlbr.) J. Steiner (30)
Protomicarea Hafellner (2)
Psora Hoffm. (35)
Psorula Gotth. Schneid. (1)

Ramalinaceae C. Agardh*
Auriculora Kalb (1)
Bacidia De Not. (= Bacidiopsora Kalb) (230)*
Bacidina Vězda (12)
Badimia Vězda (20)
Bellicidia Kistenich, Timdal, Bendiksby & Ekman (1)*
Biatora Fr. (= Myrionora R.C. Harris; = Ivanpisutia S.Y. Kondr., Lőkös & Hur) (42)*
Bibbya J.H. Willis (10)*
Bilimbia De Not. (= Myxobilimbia Hafellner) (6)
Cenozosia A. Massal. (1)
Cliostomum Fr. (25)
Echidnocymbium Brusse (1)
Eschatogonia Trevis. (7)
Heppsora D.D. Awasthi & K. Singh (1)
Jarmania Kantvilas (2)
Kiliasia Hafellner (9)*
Krogia Timdal (7)
Lecania A. Massal. (50)
Lueckingia Aptroot & Umana (1)
Mycobilimbia Rehm (5)*
Myelorrhiza Verdon & Elix (2)*
Niebla Rundel & Bowler (23)
Parallopsora Kistenich, Timdal & Bendiksby (3)*
Phyllopsora Müll. Arg. (= Crocynia (Ach.) A. Massal.) (75)*
Physcidia Tuck. (10)
Ramalina Ach. (230)
Rolfidium Moberg (3)
Scutula Tul. (=Karsteniomyces D. Hawksw.; = Libertiella Speg. & Roum.) (43)*
Sporacestra A. Massal. (1)*
Stirtoniella D.J. Galloway, Hafellner & Elix (1)
Thalloidima A. Massal. (17)*
Thamnolecania (Vain.) Gyeln. (1)
Tibellia Vězda & Hafellner (1)
Toninia A. Massal. (= Arthrosporum A. Massal.) (85)*
Toniniopsis Frey (7)
Tylothallia P. James & H. Kilias (3)*
Waynea Moberg (7)

Ramboldiaceae S. Stenroos, Miądl. & Lutzoni


Ramboldia Kantvilas & Elix (34)

Scoliciosporaceae Hafellner
Scoliciosporum A. Massal. (15)

1164
Sphaerophoraceae Fr.
Austropeltum Henssen, H. Döring & Kantvilas (1)
Bunodophoron A. Massal. (25)
Calycidium Stirt. (2)
Leifidium Wedin (1)
Neophyllis F. Wilson (2)
Sphaerophorus Pers. (8)

Tephromelataceae Hafellner
Calvitimela Hafellner (11)
Mycoblastus Norman (10)
Tephromela M. Choisy (ca. 30)
Violella T. Sprib. (2)

Lecanorales genera incertae sedis


Catinaria Vain. (2)*
Compsocladium I.M. Lamb (2)*
Coronoplectrum Brusse (1)
Corticiruptor Wedin & Hafellner (2)
Lichenosticta Zopf (5)
Myochroidea Printzen, T. Sprib. & Tønsberg (4)*
Neopsoromopsis Gyeln. (1)
Nimisiostella Calat., Barreno & O.E. Erikss. (1)
Psoromella Gyeln. (1)
Puttea S. Stenroos & Huhtinen (4)
Ramalea Nyl. (4)
Tasmidella Kantvilas, Hafellner & Elix (1)*
Umbilithecium Etayo (1)
Umushamyces Etayo (1)

Lecideales Vain.
Lecideaceae Chevall.
Amygdalaria Norman (11)
Bahianora Kalb (1)
Bellemerea Hafellner & Cl. Roux (10)
Bryobilimbia Fryday (6)*
Catarrhospora Brusse (2)
Cecidonia Triebel & Rambold (2)
Clauzadea Hafellner & Bellem. (7)
Cryptodictyon A. Massal. (2)
Eremastrella Vogel (2)
Farnoldia Hertel (6)
Immersaria Rambold & Pietschm. (8)
Koerberiella Stein (2)
Labyrintha Malcolm, Elix & Owe-Larss. (1)
Lecidea Ach. (ca. 100)
Lecidoma Gotth. Schneid. & Hertel (1)
Melanolecia Hertel (7)
Pachyphysis R.C. Harris & Ladd (1)
Paraporpidia Rambold & Pietschm. (3)
Poeltiaria Hertel (8)
Poeltidea Hertel & Hafellner (3)

1165
Porpidia Körb. (51)
Porpidinia Timdal (1)
Pseudopannaria (B. de Lesd.) Zahlbr. (1)
Rhizolecia Hertel (1)
Romjularia Timdal (1)
Schizodiscus Brusse (1)
Stenhammarella Hertel (1)
Stephanocyclos Hertel (1)
Xenolecia Hertel (2)

Lopadiaceae Hafellner
Lopadium Körb. (10)

Leprocaulales Lendemer & B.P. Hodk.


Leprocaulaceae Lendemer & B.P. Hodk.
Halecania M. Mayrhofer (22)
Leprocaulon Nyl. (ca. 10)
Speerschneidera Trevis. (1)

Peltigerales W. Watson
Coccocarpiaceae Henssen ex Henssen
Coccocarpia Pers. (ca. 50)
Peltularia R. Sant. (4)
Spilonema Bornet (4)

Collemataceae Zenker
Blennothallia Trevis. (4)
Callome Otálora & Wedin (1)
Collema F.H. Wigg. (ca. 35)
Enchylium (Ach.) Gray (11)
Lathagrium (Ach.) Gray (10)
Leptogium (Ach.) Gray (ca. 110)
Pseudoleptogium Müll. Arg. (1)
Rostania Trevis. (3 + 4 orphaned species)*
Scytinium (Ach.) Gray (49)

Koerberiaceae T. Sprib. & Muggia


Henssenia Ertz, R.S. Poulsen & Søchting (4)*
Koerberia A. Massal. (2)
Vestergrenopsis Gyeln. (2)

Massalongiaceae Wedin, P.M. Jørg. & E. Wiklund.


Leptochidium M. Choisy (2)
Massalongia Körb. (2 + 6 orphaned species)
Polychidium (Ach.) Gray (1)

Pannariaceae Tuck.
Austrella P.M. Jørg. (3)
Degelia Arv. & D.J. Galloway (16)
Erioderma Feé (32)
Fuscoderma (D.J. Galloway & P.M. Jørg.) P.M. Jørg. & D.J. Galloway (5)
Fuscopannaria P.M. Jørg. (= Kroswia P.M. Jørg.) (58)

1166
Gibbosporina Elvebakk, S.G. Hong & P.M. Jørg. (13)
Homothecium A. Massal. (4)
Joergensenia Passo, S. Stenroos & Calvelo (1)
Leciophysma Th. Fr. (2)
Leightoniella Henssen (1)
Leioderma Nyl. (7)
Lepidocollema Vain. (22)
Leptogidium Nyl. (3)
Nebularia P.M. Jørg. (2)
Nevesia P.M.Jørg, L. Lindblom, Wedin & S. Ekman (1)
Pannaria Del. ex Bory (ca. 40)
Parmeliella Müll. Arg. (ca. 40)
Pectenia P.M. Jørg. (4)
Physma A. Massal. (12)
Protopannaria (Gyeln.) P.M. Jørg. & S. Ekman (7)
Psoroma Michaux (ca. 70)
Psoromaria Nyl. ex Nyl. (= Degeliella P.M. Jørg.) (2)
Psoromidium Stirt. (2)
Ramalodium Nyl. (6)
Siphulastrum Müll. Arg. (4)
Staurolemma Körb. (3)
Steineropsis T. Sprib. & Muggia (1)

Peltigeraceae Dumort. (= Lobariaceae Chevall.; = Nephromataceae Wetm. ex J.C. David & D.


Hawksw.)*
Crocodia Link (5)
Dendriscosticta Moncada & Lücking (5)
Lobaria (Schreb.) Hoffm. (ca. 60)
Lobariella Yoshim. (35)
Lobarina Nyl. ex Cromb. (15)
Nephroma Ach. (ca. 36)
Parmostictina Nyl. (15)
Peltigera Willd. (ca. 100)
Podostictina Clem. (5)
Pseudocyphellaria Vain. (ca. 100)
Ricasolia De Not. (15)
Solorina Ach. (ca. 10)
Sticta (Schreb.) Ach. (ca. 200)
Yarrumia D.J. Galloway (2)
Yoshimuriella Moncada & Lücking (8)

Placynthiaceae Å.E. Dahl


Hertella Henssen (3)
Placynthiopsis Zahlbr. (1)
Placynthium (Ach.) Gray (ca. 20)

Vahliellaceae Wedin
Vahliella P.M. Jørg. (10)

Peltigerineae genus incertae sedis


Erinacellus T. Sprib., Muggia & Tønsberg (2)

1167
Rhizocarpales Miądl. & Lutzoni ex Miądl. & Lutzoni ex Miadl. & Lutzoni
Rhizocarpaceae M. Choisy & Hafellner
Catolechia Flot. (1)
Epilichen Clem. (2)
Poeltinula Hafellner (2)
Rhizocarpon Ramond ex DC. (225)

Sporastatiales Lumbsch & Leavitt*


Sporastatiaceae Bendiksby & Timdal
Sporastatia A. Massal. (4)
Toensbergia Bendiksby & Timdal (1)

Teloschistales D. Hawksw. & O.E. Erikss.


Brigantiaeaceae Hafellner & Bellem. (= Letrouitiaceae Bellem. & Hafellner)*
Brigantiaea Trevis. (26)
Letrouitia Hafellner & Bellem. (18)

Megalosporaceae Vězda ex Hafellner & Bellem.


Megaloblastenia Sipman (2)
Megalospora Meyen (36)
Sipmaniella Kalb (1)

Teloschistaceae Zahlbr.
Amundsenia Søchting, Garrido-Ben., Arup & Frödén (2)
Apatoplaca Poelt & Hafellner (1)
Athallia Arup, Frödén & Søchting (= ?Coppinsiella S. Y. Kondr. et al.; = ?Fominiella S. Y.
Kondr., Upreti & Hur) (17)
Austroplaca Søchting, Frödén & Arup (10)
Blastenia A. Massal. (11)
Brownliella S.Y. Kondr., Kärnefelt, Elix, A. Thell & Hur (4)
Bryoplaca Søchting, Frödén & Arup (3)
Calogaya Arup, Frödén & Søchting (= Lazarenkoella S.Y. Kondr. et al.; = Seawardiella
S.Y. Kondr. et al.) (19)
Caloplaca Th. Fr. (351)
Catenarina Søchting, Søgaard, Arup, Elvebakk & Elix (3)
Cephalophysis (Hertel) H. Kilias (1)
Cerothallia Arup, Frödén & Søchting (4)
Charcotiana Søchting, Garrido-Ben. & Arup (1)
Dijigiella S.Y. Kondr. & L. Lőkös (2)
Dufourea Ach. (= Xanthodactylon P.A. Duvign.) (25)
Eilifdahlia S.Y. Kondr., Kärnefelt, Elix, A. Thell & Hur (2)
Fauriea S.Y. Kondr., Lőkös & Hur (2)
Filsoniana S.Y. Kondr., Kärnefelt, Elix, A. Thell & Hur (= Harusavskia S.Y. Kondr.; =
Nevilleiella S.Y.Kondr. & Hur; = Thelliana S.Y. Kondr. et al.) (9)
Flavoplaca Arup, Søchting & Frödén (28)
Follmannia C.W. Dodge (2)
Franwilsia S.Y. Kondr., Kärnefelt, Elix, A. Thell & Hur (3)
Gondwania Søchting, Frödén & Arup (4)
Gyalolechia A. Massal. (= Hanstrassia S.Y. Kondr.; = Laundonia S. Y. Kondr., L. Lőkös &
Hur; = Lazarenkoiopsis S.Y. Kondr., L. Lőkös & Hur; = Opeltia S.Y. Kondr. & L. Lőkös; =
Oxneriopsis S.Y. Kondr., D. Upreti & Hur) (40)
Haloplaca Arup, Søchting & Frödén (31)

1168
Hosseusiella S.Y. Kondr., L. Lőkös, Kärnefelt & A. Thell (3)
Huneckia S.Y. Kondr., Elix, Kärnefelt, A. Thell & Hur (2)
Ioplaca Poelt (2)
Jasonhuria S.Y. Kondr., Lőkös & S.O. Oh (1)
Josefpoeltia S.Y. Kondr. & Kärnefelt (3)
Kaernefia S.Y. Kondr., Elix, A. Thell & Hur (3)
Leproplaca (Nyl.) Nyl. (7)
Loekoesia S.Y. Kondr., S.O. Oh & Hur (1)
Marchantiana S.Y. Kondr., Kärnefelt, Elix, A. Thell & Hur (= Streimanniella S.Y. Kondr.
et al.) (5)
Olegblumia S.Y. Kondr., Lőkös & Hur (1)
Orientophila Arup, Søchting & Frödén (4)
Pachypeltis Søchting, Arup & Frödén (4)
Parvoplaca Arup, Søchting & Frödén (6)
Polycauliona Hue (= ? Tomnashia S.Y. Kondr. & Hur) (18)
Pyrenodesmia A. Massal. (6)
Rehmaniella S.Y. Kondr. et al. (1)
Rufoplaca Arup, Søchting & Frödén (6)
Rusavskia S.Y. Kondr. & Kärnefelt (= ? Zeroviella S.Y. Kondr. & J.-S. Hur) (19)
Scutaria Søchting, Arup & Frödén (1)
Seirophora Poelt (ca. 8)
Shackletonia Søchting, Frödén & Arup (5)
Sirenophila Søchting, Arup & Frödén (= Elixjohnia S.Y. Kondr. & Hur; = Tarasginia S.Y.
Kondr. et al.) (14)
Solitaria Arup, Søchting & Frödén (1)
Squamulea Arup, Søchting & Frödén (= Huriella S.Y. Kondr. & D. Upreti) (8)
Stellarangia Frödén, Arup & Søchting (3)
Tassiloa S.Y. Kondr., Kärnefelt, A. Thell, Elix & Hur (2)
Teloschistes Norman (ca. 24)
Teloschistopsis Frödén, Søchting & Arup (3)
Teuvoahtiana S.Y. Kondr. & Hur (3)
Upretia S.Y. Kondr., A. Thell & Hur
Usnochroma Søchting, Arup & Frödén (2)
Variospora Arup, Søchting & Frödén (16)
Villophora Søchting, Arup & Frödén (= Tayloriella S.Y. Kondr. et al.; = Tayloriellina S.Y.
Kondr. et al.) (4)
Wetmoreana Arup, Søchting & Frödén (3)
Xanthocarpia A. Massal. & De Not. (12)
Xanthomendoza S.Y. Kondr. & Kärnefelt (20)
Xanthopeltis R. Sant. (1)
Xanthoria (Fr.) Th. Fr. (10)
Yoshimuria S.Y. Kondr., Kärnefelt, Elix, A. Thell & Hur (= Ikaeria S.Y. Kondr., D. Upreti
& Hur) (4)

Teloschistales genus incertae sedis


Malcolmiella Vězda (1)

Lecanoromycetidae familis incertae sedis


Biatorellaceae M. Choisy ex Hafellner & Casares-Porcel
Biatorella De Not. (ca. 30)

Helocarpaceae Hafellner

1169
Helocarpon Fr. (3)

Pachyascaceae Poelt ex P.M. Kirk, P.F. Cannon & J.C. David


Pachyascus Poelt & Hertel (1)

Ostropomycetidae V. Reeb, Lutzoni & Cl. Roux


Baeomycetales Lumbsch, Huhndorf & Lutzoni. (= Arctomiales S. Stenroos, Miądl. & Lutzoni; =
Hymeneliales S. Stenroos, Miądl. & Lutzoni; = Trapeliales B.P. Hodk. & Lendemer)*
Arctomiaceae Th. Fr.
Arctomia Th. Fr. (14)
Gregorella Lumbsch (1)
Steinera Zahlbr. (14)*
Wawea Henssen & Kantvilas (1)

Arthrorhaphidaceae Poelt & Hafellner


Arthrorhaphis Th. Fr. (13)

Baeomycetaceae Dumort.
Ainoa Lumbsch & I. Schmitt (2)
Anamylopsora Timdal (1)
Baeomyces Pers. (10)
Parainoa Resl & T. Sprib. (1)
Phyllobaeis Gierl & Kalb (6)

Cameroniaceae Kantvilas & Lumbsch


Cameronia Kantvilas (2)

Hymeneliaceae Körb.
Hymenelia Kremp. (26)
Ionaspis Th. Fr. (7)
Tremolecia M. Choisy (6)

Protothelenellaceae Vězda, H. Mayrhofer & Poelt (= Thrombiaceae Poelt & Vězda ex J.C. David
& D. Hawksw.)*
Mycowinteria Sherwood (3)
Protothelenella Räsenen (11)
Thrombium Wallr. (5)

Trapeliaceae M. Choisy ex Hertel


Amylora Rambold (1)
Aspiciliopsis (Müll. Arg.) M. Choisy (1)
Coppinsia Lumbsch & Heibel (1)
Ducatina Ertz & Søchting (1)*
Lignoscripta B.D. Ryan (1)
Orceolina Hertel (2)
Placopsis (Nyl.) Linds. (ca. 60)
Placynthiella Elenkin (7)
Rimularia Nyl. (4)*
Sarea Fr. (2)
Trapelia M. Choisy (24)
Trapeliopsis Hertel & Gotth. Schneid. (20)

1170
Xylographaceae Tuck.
Lambiella Hertel (12)
Lithographa Nyl. (10)
Ptychographa Nyl. (1)
Xylographa (Fr.) Fr. (2)

Graphidales Bessey
Diploschistaceae Zahlbr.
Acanthothecis Clem. (ca. 60)
Acanthotrema Frisch (5)
Aggregatorygma M. Cáceres, Aptroot & Lücking (1)
Ampliotrema Kalb ex Kalb (12)
Asteristion Leight. (9)
Austrotrema I. Medeiros, Lücking & Lumbsch (3)
Borinquenotrema Merc.-Díaz, Lücking & Parnmen (1)
Byssotrema M. Cáceres (1)
Carbacanthographis Staiger & Kalb (28)
Compositrema Rivas Plata, Lücking & Lumbsch (4)
Corticorygma M. Cáceres, S.C. Feuerst., Aptroot & Lücking (1)
Diploschistes Norman (33)
Fibrillithecis A. Frisch (15)
Gintarasia Kraichak, Lücking & Lumbsch (5)
Glaucotrema Rivas Plata & Lumbsch (5)
Gyrotrema A. Frisch (6)
Heiomasia Nelsen, Lücking & Rivas Plata (3)
Melanotopelia Lumbsch & Mangold (4)
Melanotrema A. Frisch (12)
Myriochapsa M. Cáceres, Lücking & Lumbsch (3)
Myriotrema Fée (55)
Nadvornikia Tibell (5)
Nitidochapsa Parnmen, Lücking & Lumbsch (5)
Ocellularia G. Mey. (ca. 400)
Phaeographopsis Sipman (3)
Pseudoramonia Kantvilas & Vězda (4)
Redingeria A. Frisch (8)
Reimnitzia Kalb (1)
Rhabdodiscus Vain. (35)
Sanguinotrema Lücking (1)
Schizotrema Mangold & Lumbsch (6)
Stegobolus Mont. (16)
Topeliopsis Kantvilas & Vězda (20)
Wirthiotrema Rivas Plata, Kalb, Frisch & Lumbsch (5)
Xalocoa Kraichak, Lücking & Lumbsch (1)

Fissurinaceae (Rivas Plata, Lücking & Lumbsch) B.P. Hodk.


Clandestinotrema Rivas Plata, Lücking & Lumbsch (17)
Cruentotrema Rivas Plata, Papong, Lumbsch & Lücking (6)
Dyplolabia A. Massal. (5)
Enigmotrema Lücking (1)
Fissurina Fée (ca. 155)
Pycnotrema Rivas Plata & Lücking (2)

1171
Gomphillaceae Walt. Watson
Actinoplaca Müll. Arg. (2)
Aderkomyces Bat. (30)
Aplanocalenia Lücking, Sérus. & Vězda (1)
Arthotheliopsis Vain. (5)
Asterothyrium Müll. Arg. (32)
Aulaxina Fée (14)
Calenia Müll. Arg. (30)
Caleniopsis Vězda & Poelt (2)
Corticifraga D. Hawksw. & R. Sant. (7)*
Diploschistella Vain. (4)
Echinoplaca Fée (40)
Ferraroa Lücking, Sérus. & Vězda (1)
Gomphillus Nyl. (6)
Gyalectidium Müll. Arg. (52)
Gyalidea Lettau (50)
Gyalideopsis Vězda (91)
Hippocrepidea Sérus. (1)
Jamesiella Lücking, Sérus. & Vězda (4)
Lithogyalideopsis Lücking, Sérus. & Vězda (4)
Paratricharia Lücking (1)
Paragyalideopsis Etayo (4)
Phyllogyalidea Lücking & Aptroot (2)
Psorotheciopsis Rehm (7)
Rolueckia Papong, Thammath. & Boonpr. (2)
Taitaia Suija, Kaasalainen, Kirika & Rikkinen (1)*
Tricharia Fée (ca. 30)

Graphidaceae Dumort.
Allographa Chevall. (183)*
Amazonotrema Kalb & Lücking (1)
Anomalographis Kalb (2)
Anomomorpha Nyl. ex Hue (8)
Creographa A. Massal. (1)
Cryptoschizotrema Aptroot, Lücking & M. Cáceres (1)
Diaphorographis A.W. Archer & Kalb (2)
Diorygma Eschw. (74)
Flegographa A. Massal. (1)
Glyphis Ach. (7)
Graphis Adans. (ca. 275)
Halegrapha Rivas Plata & Lücking (9)
Hemithecium Trevis. (ca. 50)
Kalbographa Lücking (5)
Leiorreuma Eschw. (18)
Malmographina M. Cáceres, Rivas Plata & Lücking (1)
Mangoldia Lücking, Parnmen & Lumbsch (2)
Pallidogramme Staiger, Kalb & Lücking (13)
Phaeographis Müll. Arg. (ca. 180)
Platygramme Fée (30)
Platythecium Staiger (27)
Pliariona A. Massal. (= Phaeographina Müll. Arg.) (1)
Polistroma Clemente (1)

1172
Pseudochapsa Parnmen, Lücking & Lumbsch (18)
Pseudotopeliopsis Parnmen, Lücking & Lumbsch (4)
Sarcographa Fée (37)
Sarcographina Müll. Arg. (6)
Schistophoron Stirt. (5)
Thalloloma Trevis. (20)
Thecaria Fée (4)
Thecographa A. Massal. (3)

Redonographaceae (Lücking, Tehler & Lumbsch) Lumbsch (Bas.: Redonographoideae Lücking,


Tehler & Lumbsch, Am. J. Bot. 100: 846 2013)
Gymnographopsis C.W. Dodge (2)
Redonographa Lücking, Tehler & Lumbsch (4)

Thelotremataceae Stizenb.
Astrochapsa Parnmen, Lücking & Lumbsch (28)
Chapsa A. Massal. (ca. 60)
Chroodiscus (Müll. Arg.) Müll. Arg. (17)
Crutarndina Parnmen, Lücking & Lumbsch (1)
Leucodecton A. Massal. (31)
Paratopeliopsis Merc.-Díaz, Lücking & Parnmen (1)
Thelotrema Ach. (= Tremotylium Nyl.) (106)

Gyalectales Henssen ex D. Hawksw. & O.E. Erikss.


Coenogoniaceae (Fr.) Stizenb.
Coenogonium Ehrenb. ex Nees (ca. 91)

Gyalectaceae (A. Massal.) Stizenb.


Gyalecta Ach. (= Cryptolechia A. Massal.) (50)
Ramonia Stizenb. (24)
Semigyalecta Vain. (1)

Phlyctidaceae Poelt & Vězda ex J.C. David & D. Hawksw.


Phlyctis (Wallr.) Flot. (20)
Psathyrophlyctis Brusse (1)

Sagiolechiaceae Baloch, Lücking, Lumbsch & Wedin


Rhexophiale Th. Fr. (1)
Sagiolechia A. Massal. (3)

Trichotheliaceae Bitter & F. Schill. (= Porinaceae Walt. Watson; = Porinaceae Rchb.)


Clathroporina Müll. Arg. (ca. 25)
Flabelloporina Sobreira, M. Cáceres & Lücking (1)
Myeloconis P.M. McCarthy & Elix (4)
Porina Müll. Arg. (ca. 145)
Pseudosagedia (Müll. Arg.) Choisy (80)
Segestria Fr. (70)
Trichothelium Müll. Arg. (40)

Ostropales Nannf.
Odontotremataceae D. Hawksw. & Sherwood
Claviradulomyces P.R. Johnst., D.C. Park, H.C. Evans, R.W. Barreto & D.J. Soares (1)

1173
Coccomycetella Höhn. (2)
Odontotrema Nyl. (7)
Odontura Clem. (1)
Paschelkiella Sherwood (1)
Potriphila Döbbeler (3)
Rogellia Döbbeler (2)
Stromatothecia D.E. Shaw & D. Hawksw. (1)
Tryblis Clem. (2)
Xerotrema Sherwood & Coppins (2)

Phaneromycetaceae Gamundí & Spinedi


Phaneromyces Speg. & Har. ex Speg. (2)

Spirographaceae Flakus, Etayo & Miadlikowska


Spirographa Zahlbr. (5)

Stictidaceae Fr.
Absconditella Vězda (12)
Acarosporina Sherwood (5)
Biostictis Petr. (5)
Carestiella Bres. (1)
Conotremopsis Vězda (1)
Cryptodiscus Corda (= Lettauia D. Hawksw. & R. Sant.) (9)*
Cyanodermella O.E. Erikss. (2)
Delpontia Penz. & Sacc. (1)
Dendroseptoria Alcalde (3)
Fitzroyomyces Crous (1)
Geisleria Nitschke (1)
Glomerobolus Kohlm. & Volkm.-Kohlm. (1)
Ingvariella Guderley & Lumbsch (1)
Karstenia Fr. (10)
Lillicoa Sherwood (4)
Nanostictis M.S Christ. (ca. 8)
Neofitzroyomyces Crous (1)
Ostropa Fr. (1)
Propoliopsis Rehm (1)
Robergea Desm. (8)
Schizoxylon Pers. (ca. 35)
Sphaeropezia Sacc. (= Lethariicola Grummann) (19)
Stictis Pers. (4)
Stictophacidium Rehm (3)
Thelopsis Nyl. (9)
Topelia P.M. Jørg. & Vězda (6)
Trinathotrema Lücking, Rivas Plata & Mangold (3)
Xyloschistes Vain. ex Zahlbr. (1)

Ostropales genera incertae sedis


Aabaarnia Diederich (1)
Biazrovia Zhurb. & Etayo (1)
Elongaticonidia W.J. Li, E. Camporesi & K.D. Hyde (1)
Epicladonia D. Hawksw. sensu lato (2)*
Normanogalla Diederich (1)

1174
Paralethariicola Calat., Etayo & Diederich (1)

Pertusariales M. Choisy ex D. Hawksw. & O.E. Erikss.


Agyriaceae Corda (= Miltideaceae Hafellner)*
Agyrium Fr. (3)
Miltidea Stirt. (1)

Coccotremataceae Henssen ex J.C. David & D. Hawksw.


Coccotrema Müll. Arg. (16)
Gyalectaria I. Schmitt, Kalb & Lumbsch (3)
Parasiphula Kantvilas & Grube (7)

Icmadophilaceae Triebel
Dibaeis Clem. (ca. 14)
Endocena Cromb. (=Chirleja Lendemer & B.P. Hodk.) (2)
Icmadophila Trevis. (4)
Pseudobaeomyces M. Satì (1)
Siphula Fr. (26)
Siphulella Kantvilas, Elix & P. James (1)
Thamnolia Ach. ex Schaerer (4)

Megasporaceae Lumbsch
Aspicilia A. Massal. (ca. 200)
Circinaria Link (ca. 40)
Lobothallia (Clauzade & Cl. Roux) Hafellner (12)
Megaspora (Clauzade & Cl. Roux) Hafellner & V. Wirth (4)
Sagedia Ach. (ca. 30)
Teuvoa Sohrabi & S. Leavitt (5)

Microcaliciaceae Tibell*
Microcalicium Vain. (4)

Ochrolechiaceae R.C. Harris ex Lumbsch & I. Schmitt


Ochrolechia A. Massal. (60)

Pertusariaceae Körb. ex Körb.


Loxosporopsis Henssen (1)
Pertusaria DC. (ca. 400)*
Thamnochrolechia Aptroot & Sipman (1)

Varicellariaceae B.P. Hodk., R.C. Harris & Lendemer ex Lumbsch & Leavitt
Varicellaria Nyl. (8)*

Variolariaceae Fée ex Zenker


Lepra Scop. (= Marfloraea S.Y. Kondr., Lőkös & Hur) (94)

Sarrameanales B.P. Hodk. & Lendemer


Sarrameanaceae Hafellner
Loxospora A. Massal. (13)
Sarrameana Vězda & P. James (1)

Schaereriales Lumbsch & Leavitt

1175
Schaereriaceae M. Choisy ex Hafellner
Schaereria Körb. (= Hafellnera Houmeau & Cl. Roux) (16)

Thelenellales Lumbsch & Leavitt


Thelenellaceae O.E. Erikss. ex H. Mayrhofer
Aspidothelium Vain. (17)
Chromatochlamys Trevis. (3)
Thelenella Nyl. (30)

Ostropomycetidae family incertae sedis


Epigloeaceae Zahlbr.
Epigloea Zukal (12)

Ostropomycetidae genera incertae sedis


Amphorothecium P.M. McCarthy, Kantvilas & Elix (1)
Anzina Scheid. (1)
Aspilidea Hafellner (1)
Bachmanniomyces D. Hawksw. (8)
Dictyocatenulata Finley & E.F. Morris (1)
Malvinia Döbbeler (1)
Pleiopatella Rehm (1)

Umbilicariomycetidae Bendiksby, Hestmark & Timdal


Umbilicariales J.C. Wei & Q.M. Zhou
Elixiaceae Lumbsch
Elixia Lumbsch (2)
Meridianelia Kantvilas & Lumbsch (1)

Fuscideaceae Hafellner
Fuscidea V. Wirth & Vězda (ca. 40)
Hueidea Kantvilas & P.M. McCarthy (1)
Maronea A. Massal. (13)
Orphniospora Körb. (4)

Ophioparmaceae R.W. Rogers & Hafellner


Boreoplaca Timdal (1)
Hypocenomyce M. Choisy (3)
Ophioparma Norman (9)

Ropalosporaceae Hafellner
Ropalospora A. Massal. (9)

Umbilicariaceae Chevall.
Fulgidea Bendiksby & Timdal (2)
Umbilicaria Hoffm. (= Lasallia Mérat) (ca. 90) *
Xylopsora Bendiksby & Timdal (2)

Lecanoromycetes order incertae sedis


Micropeltidales X.Y. Zeng, H.X. Wu & K.D. Hyde
Micropeltidaceae Clem. & Shear*
Anariste Syd. (1)
Caudella Syd. & P. Syd. (2)

1176
Cyclopeltella Petr. (1)
Dictyopeltella Bat. & I.H. Lima (2)
Haplopeltheca Bat., J.L. Bezerra & Cavalc. (1)
Micropeltis Mont. (ca. 110)
Neopeltella Petr. (1)
Scolecopeltidium F. Stevens & Manter (ca. 80)
Stomiopeltis Theiss. (25)
Stomiopeltopsis Bat. & Cavalc. (2)
Stomiotheca Bat. (2)

Turquoiseomycetales Crous
Turquoiseomycetaceae Crous
Turquoiseomyces Crous (1)

Lecanoromycetes genera incertae sedis


Argopsis Th. Fr. (1)
Ascographa Velen. (1)
Bartlettiella D.J. Galloway & P.M. Jørg. (1)
Bouvetiella Øvstedal (1)
Buelliastrum Zahlbr. (1)
Haploloma Trevis. (1)
Hosseusia Gyeln. (3)
Korfiomyces Iturr. & D. Hawksw. (1)
Maronella M. Steiger (1)
Notolecidea Hertel (1)
Petractis Fr. (3)
Piccolia A. Massal. (ca. 7)
Ravenelula Speg. (1)
Robincola Velen. (1)
Roburnia Velen. (1)

Leotiomycetes O.E. Erikss. & Winka


Chaetomellales Crous & Denman
Chaetomellaceae Baral, P.R. Johnst. & Rossman
Chaetomella Fuckel (26)
Pilidium Kunze (23)
Sphaerographium Sacc. (23)
Synchaetomella Decock & Seifert (3)

Cyttariales Luttr. ex Gamundí


Cyttariaceae Speg.
Cyttaria Berk. (13)

Helotiales Nannf. ex Korf & Lizoň


Amorphothecaceae Parbery*
Amorphotheca Parbery (21 fide Baral 2016)

Arachnopezizaceae Hosoya, J.G. Han & Baral


Arachnopeziza Fuckel (35)
Arachnoscypha Boud. (1)
Austropezia Spooner (1)
Eriopezia (Sacc.) Rehm (21)

1177
Parachnopeziza Korf (8)

Ascocorticiaceae J. Schrot
Ascocorticiellum Julich & B. de Vries (1)
Ascocorticium Bref. (2)
Ascosorus P. Henn. & Ruhland (1)

Ascodichaenaceae D. Hawksw. & Sherwood


Ascodichaena Butin (2)
Delpinoina Kuntze (1)

Bloxamiaceae Locq.
Bloxamia Berk. & Broome (10)

Bryoglossaceae Ekanayaka & Hyde


Bryoclaviculus L. Ludw., P.R. Johnst. & Steel (1)
Bryoglossum Redhead (2)
“Crocicreas” multicuspidatum (1)
Neocudoniella S. Imai (3)
“Roseodiscus” formosus (1)

Calloriaceae Marchand
Aivenia Svrcek (4)
Calloria Fr. (28)
Chaetonaevia Arx (3)
Diplonaevia Sacc. (33)
Duebenia Fr. (6)
Eupropolella Hohn. (8)
Hyalacrotes (Korf & L.M. Kohn) Raitv. (5)
Iridinea Velen. (2)
Laetinaevia Nannf. (19)
Loricella Velen. (6)
Micropodia Boud. (15)
Naeviella (Rehm) Clem. (3)
Naeviopsis B. Hein (14)
Ploettnera Henn. (6)

Cenangiaceae Rehm (= Hemiphacidiaceae Korf)*


Cenangiopsis Rehm (9)
Cenangium Fr. (47)
Chlorencoelia J.R. Dixon (4)
Crumenulopsis J.W. Groves (3)
Encoelia (Fr.) P. Karst. (38)
Fabrella Kirschst. (1)
Heyderia Link (4)
Rhabdocline Syd. (7)
Sarcotrochila Hohn. (7)
Trochila Fr. (33)
Velutarina Korf (3)

Chlorociboriaceae Baral & P.R. Johnst.*


Chlorociboria Seaver ex C.S. Ramamurthi, Korf & L.R. Batra (23)

1178
Chlorospleniaceae Ekanayaka & Hyde
Chlorosplenium Fr. (17)

Chrysodiscaceae Baral & Haelew.*


Chrysodisca Baral, Polhorský & G. Marson (1)

Cordieritidaceae Sacc.
Ameghiniella Speg. (2)
Annabella Fryar, Haelew. & D.E.A. Catches. (1)
Austrocenangium Gamundí (2)
Cordierites Mont. (5)
Diplocarpa Massee (1)
Diplolaeviopsis Giralt & D. Hawksw. (3)
“Encoelia” fimbriata Spooner & Trigaux (1)
“Encoelia” heteromera (Mont.) Nannf. (1)
Ionomidotis E.J. Durand ex Thaxt. (4)
Llimoniella Hafellner & Nav.-Ros. (21)
Macroskyttea Etayo, Flakus, Suija & Kukwa (1)
Midotiopsis Henn. (2)
Rhymbocarpus Zopf (10)
Sabahriopsis Crous & M.J. Wingf. (1)
Skyttea Sherwood, D. Hawksw. & Coppins (30)
Skyttella D. Hawksw. & R. Sant. (2)
Thamnogalla D. Hawksw. (1)
Unguiculariopsis Rehm (29)

Dermateaceae Fr.
Coleophoma Hohn. (= Parafabraea Chen Chen et al.) (30)*
Corniculariella P. Karst. (3)
Dermea Fr. (24)
Gelatinoamylaria Prasher & R. Sharma (1)*
Neodermea W.J. Li, D.J. Bhat & K.D. Hyde (1)
Neofabraea H.S. Jacks. (9)
Neogloeosporidina W.J. Li, Camporesi & K.D. Hyde (1)
Pezicula Tul. & C. Tul. (92)
Phlyctema Desm. (60)
Pseudofabraea Chen Chen, Verkley & Crous (1)
Rhizodermea Verkley & Zijlstra (1)
Schizothyrioma Hohn (4)
Verkleyomyces Y. Marın & Crous (1)
Xenochalara M.J. Wingf. & Crous (1)

Discinellaceae Ekanayaka & K.D. Hyde*


Articulospora Ingold (6)
Cladochasiella Marvanova (1)
Discinella Boud. (13)
Fontanospora Dyko (4)
Gyoerffyella Kol (10)
Lemonniera De Wild. (8)
Margaritispora Ingold (2)
Naevala B. Hein (5)
Pezoloma Clem. (14)

1179
Pseudopezicula Korf (2)
Tetrachaetum Ingold (1)
Varicosporium W. Kegel (9)

Drepanopezizaceae Baral*
Blumeriella Arx (7)
Diplocarpon F.A. Wolf (7)
Drepanopeziza (Kleb.) Hohn. (5)
Felisbertia Viegas (7)
Leptotrochila P. Karst. (15)
Pseudopeziza Fuckel (2)
Spilopodia Boud. (4)
Spilopodiella E. Mull. (1)

Erysiphaceae Tul. & C. Tul.


Arthrocladiella Vassilkov (1)
Blumeria Golovin ex Speer (1)
Brasiliomyces Viegas (6)
Bulbomicroidium Marm., S. Takam. &U. Braun (1)*
Caespitotheca S. Takam. & U. Braun (1)
Cystotheca Berk. & Curtis (9)
Erysiphe DC. (478)
Golovinomyces (U. Braun) Heluta (66)
Leveillula G. Arnaud (49)
Microidium (To-anun & S. Takam.) To-anun & S. Takam. (3)
Neoerysiphe U. Braun (15)
Parauncinula S. Takam. & U. Braun (4)
Phyllactinia Lev. (117)
Pleochaeta Sacc. & Speg. (5)
Podosphaera Kunze (124)
Pseudoidium Y.S. Paul & J.N. Kapoor (80)
Queirozia Viegas & Cardoso (1)
Sawadaea Miyabe (10)
Takamatsuella U. Braun & A. Shi (1)
Typhulochaeta Ito & Hara (4)

Gelatinodiscaceae S.E. Carp


Ascocoryne J.W. Groves & D.E. Wilson (8)
Ascotremella Seaver (2)
Chloroscypha Seaver (14)
Didymocoryne Sacc. & Trotter (1)
Neobulgaria Petr. (11)
Ombrophila Fr. (11)
Phaeangellina Dennis (1)
Skyathea Spooner & Dennis (1)
Xerombrophila Baral (1)

Godroniaceae Baral
Ascocalyx Naumov (4)
Atropellis Zeller & Goodd. (4)
Godronia Moug. & Lev. (30)
Gremmeniella M. Morelet (3)

1180
Grovesiella M. Morelet (2)*

Helotiaceae Rehm
Ascoconidium Seaver (3)
Bisporella Sacc. (19)
Bryoscyphus Spooner (19)
Calycella (Sacc.) Sacc. (1)
Cudoniella Sacc. (31)
Cyathicula De Not. (30)
Dicephalospora Spooner (4)
Dimorphospora Tubaki (1)
Discorehmia Kirschst. (5)
Eubelonis Hohn. (2)
Filosporella Nawawi (6)
Geniculospora Sv. Nilsson ex Marvanová & Sv. Nilsson (2)
Glarea Bills & Palaez (2)
Gloeotinia M. Wilson, Noble & E.G. Gray (2)
Graddonia Dennis (7)
Gremmenia Korf (4)
Helicodendron Peyronel (3)
Hymenoscyphus Gray (170)
Hymenotorrendiella P.R. Johnst., Baral & R. Galán (9)
Muscicola Velen. (1)
Mycofalcella Marvanová, Om-Kalth. & J. Webster (2)
Mytilodiscus Kropp & S.E. Carp. (1)
Neocrinula Crous (2)
Phaeohelotium Kanouse (41)
Pithyella Boud. (8)
Pseudohelotium Fuckel (50)
Pseudoniptera Velen. (25)
Roesleria Thüm. & Pass. (4)*
Scytalidium Pesante (30)
Symphyosirinia E.A. Ellis (6)
Tatraea Svrcek (2)
Tricladium Ingold (25)
Xylogramma Wallr. (18)

Heterosphaeriaceae Rehm
Heterosphaeria Grev. (7)

Hyaloscyphaceae Nannf.*
Aeruginoscyphus Dougoud (7)*
Ambrodiscus S.E. Carp. (1)
Amicodisca Svrcek (6)
Arbusculina Marvanova & Descals (3)
Asperopilum Spooner (1)
"Chalara" longipes (Preuss) Cooke (1)
Clathrosphaerina Beverw. (2)
Crucellisporiopsis Nag Raj (3)
Dematioscypha Svrcek (4)
Dimorphotricha Spooner (1)
Echinula Graddon (1)

1181
Endoscypha Syd. (1)
Fuscolachnum J.H. Haines (7)
Gamarada D.J. Midgley & Tran-Dinh (1)
Graddonidiscus Raitv. & R. Galan (3)
Grahamiella Spooner (3)
Haplographium Berk. & Broome (15)
Hegermila Raitv. (4)
Hyalopeziza Fuckel (15)
Hyaloscypha Boud. (45)
Hyphodiscus Kirschst. (16)
Hyphopeziza J.G. Han, Hosoya & H.D. Shin (1)
Incrupila Raitv. (10)
Meliniomyces Hambl. & Sigler (3)
Mycoarthris Marvanova & P.J. Fisher (1)
Olla Velen. (2)
Polaroscyphus Huhtinen (1)
Proprioscypha Spooner (1)
Protounguicularia Raitv. & Galan (10)
Pseudaegerita J.L. Crane & Schokn. (7)*
Psilocistella Svrcek (10)
Rhizoscyphus W.Y. Zhuang & Korf (1)
Scolecolachnum Guatim., R.W. Barreto & Crous (2)*
Thindiomyces Arendh. & R. Sharma (1)
Unguiculariella K.S. Thind & R. Sharma (1)
Unguiculella Hohn (17)
Venturiocistella Raitv (7)

Lachnaceae (Nannf.) Raitv.


Albotricha Raitv. (19)
Belonidium Mont. & Dur. (1)
Brunnipila Baral (10)
Capitotricha (Raitv.) Baral (10)
Dasyscyphella Tranzschel (1)
Erioscyphella Kirschst. (10)
Incrucipulum Baral (6)
Lachnellula P. Karst. (40)
Lachnopsis Guatim., R.W. Barreto & Crous (2)*
Lachnum Retz. (50)
Lasiobelonium Ellis & Everh. (20)
Neodasyscypha Sukova & Spooner (2)
Perrotia Boud. (19)
Proliferodiscus J.H. Haines & Dumont (8)
Solenopezia Sacc. (7)
Trichopeziza Fuckel (30)
Tubolachnum Velen (2)
Velebitea I. Kušan, Matočec & Jadan (1)

Leptodontidiaceae Hern.-Restr., Crous & Gené


Leptodontidium de Hoog. (11)

Loramycetaceae Dennis ex Digby & Goos


Loramyces W. Weston (2)

1182
Obtectodiscus E. Müll., Petrini & Samuels (2)

Mitrulaceae Rchb.
Mitrula Fr. (16)

Mollisiaceae Rehm
Barrenia E. Walsh & N. Zhang (2)*
Bulbomollisia Graddon (1)
Cheirospora Moug. & Fr. (2)*
Cystodendron Bubak (2)
Discocurtisia Nannf. (12)
Fuscosclera Hern.-Restr., J. Mena & Gené (1)*
Mollisia (Fr.) P. Karst. (130)
Neotapesia E. Mull. & Hutter (3)
Niptera Fr. (10)
Nipterella Starback ex Dennis (2)
Phialocephala W.B. Kendr. (37)*
Pseudonaevia Dennis & Spooner (2)
Sarconiptera Raitv. (1)
Scutobelonium Graddon (1)
Scutomollisia Nannf. (14)
Tapesia (Pers.) Fuckel (110)*
Trimmatostroma Corda (30)
Variocladium Descals & Marvanova (1)

Myxotrichaceae Currah*
Byssoascus Arx (1)
“Malbranchea” flavorosea Sigler & J.W. Carmich. (1)
Myxotrichum Kunze (17)
Oidiodendron Robak (26)

Neolauriomycetaceae Crous*
Exochalara W. Gams & Hol.-Jech. (3)
Lareunionomyces Crous & M.J. Wingf. (4)
Neolauriomyces Crous (1)

Pezizellaceae Velen.
Allophylaria (P. Karst.) P. Karst. (6)
Antinoa Velen. (8)
Calycellina Hohn (45)
Calycina Nees ex Gray (30)
Chalara (Corda) Rabenh. (99)
Ciliolarina Svrcek (1)
Curviclavula G. Delgado, F.A. Fernández & A.N. Mill. (1)
Hamatocanthoscypha Svrcek (3)
Hyalodendriella Crous (1)
Micropeziza Fuckel (12)
Microscypha Syd. & P. Syd. (6)
Mollisina Hohn. ex Weese (11)
Mollisinopsis Arendh. & R. Sharma (3)
Moserella Poder & Scheuer (1)
Phaeoscypha Spooner (1)

1183
Phialina Höhn. (6)*
Poculinia Spooner (1)
Psilachnum Hohn. (28)
Rodwayella Spooner (3)
Scleropezicula Verkley (6)
Velutaria Fuckel (1)
Xenopolyscytalum Crous (1)
Zymochalara Guatim., R.W. Barreto & Crous (2)*

Ploettnerulaceae Kirschst.
Cadophora Lagerb. & Melin (15)
Collembolispora Marvanova & Pascoal (2)
Cylindrosporium Grev. (168)*
Dennisiodiscus Svrcek (10)
Lasiomollisia Raitv. & Vesterh. (1)
Mastigosporium Riess (4)
Mycochaetophora Hara & Ogawa (2)
Nothophacidium J. Reid & Cain (1)
Oculimacula Crous & W. Gams (6)
Pirottaea Sacc. (28)
Pyrenopeziza Fuckel (3)
Rhynchosporium Heinsen ex A.B. Frank (5)

Rutstroemiaceae Holst-Jensen, L.M. Kohn & T. Schumach.*


Bicornispora Checa, Barrasa, M.N. Blanco & A.T. Martínez (2)
Dencoeliopsis Korf (2)
Lambertella Hohn. (6)
Lanzia Sacc. (1)
Pseudolanzia Baral & G. Marson (1)*
Rutstroemia P. Karst. (100)
Torrendiella Boud. & Torrend (3)

Sclerotiniaceae Whetzel ex Whetzel


Amphobotrys Hennebert (1)
Botrytis P. Micheli ex Pers. (3)
Ciboria Fuckel (21)
Ciborinia Whetzel (16)
Cristulariella Hohn. (5)
Cudoniopsis Speg. (1)
Dumontinia L.M. Kohn (5)
Elliottinia L.M. Kohn (1)
Grovesinia M.N. Cline, J.L. Crane & S.D. Cline (2)
Haradamyces Masuya, Kusunoki, Kosaka & Aikawa (1)
Kohninia Holst-Jensen, Vrålstad & T. Schumach. (1)
Martininia Dumont & Korf (1)
Monilinia Honey (30)
Mycopappus Redhead & G.P. White (3)*
Myrioconium Syd. & P. Syd. (10)
Myriosclerotinia N.F. Buchw. (10)
Ovulinia Weiss (9)
Phaeosclerotinia Hori (1)
Piceomphale Svrcek (1)

1184
Pseudociboria Kanouse (1)
Pycnopeziza W.L. White & Whetzel (5)
Redheadia Y. Suto & Suyama (1)
Sclerencoelia Pärtel & Baral (3)*
Scleromitrula S. Imai (6)
Sclerotinia Fuckel (15)
Sclerotium Tode (100)
Seaverinia Whetzel (2)
Septotinia Whetzel ex J.W. Groves & M.E. Elliott (2)
Streptotinia Whetzel (3)
Stromatinia (Boud.) Boud. (15)
Valdensia Peyronel (3)

Vibrisseaceae Korf
Acephala Grunig & T.N. Sieber (2)
Chlorovibrissea L.M. Kohn (4)
Leucovibrissea (A. Sanchez) Korf (1)
Pocillum De Not. (1)
Vibrissea Fr (34)

Helotiales genera incertae sedis


Acidea Hujslova & M. Kolarık (1)
Acidomelania E. Walsh & N. Zhang (1)
Algincola Velen. (1)
Amylocarpus Curr. (1)
Angelina Fr. (1)
Apiculospora Wijayaw., Camporesi, A.J.L. Phillips & K.D. Hyde (1)
Aquadiscula Shearer & J.L. Crane (2)
Aquapoterium Raja & Shearer (1)
Ascluella DiCosmo, Nag Raj & W.B. Kendr. (1)
Ascoclavulina Otani (8)
Banksiamyces G. Beaton (4)
Belonioscyphella Hohn. (4)
Benguetia Syd. & P. Syd. (1)
Bioscypha Syd. (2)
Brachyalara Reblova & W. Gams (1)
Brefeldochium Verkley (1)
Bulgariella P. Karst. (4)
Bulgariopsis Henn. (2)
Calycellinopsis W.Y. Zhuang (1)
Capillipes R. Sant. (1)
Capricola Velen. (1)
Cashiella Petr. (3)
Cejpia Velen. (3)
Cenangiumella J. Frohl. & K.D. Hyde (1)
Chloroepilichen Etayo (1)
Chlorospleniella P. Karst. (1)
Chondroderris Maire (1)
Ciliella Sacc. & P. Syd. (1)
Cistella Quel. (50)
Clathrosporium Nawawi & Kuthub. (1)
Coleosperma Ingold (1)

1185
Colipila Baral & Guy Garcıa (2)
Comesia Sacc. (3)
Cornuntum Velen. (1)
Coronellaria P. Karst. (4)
Criserosphaeria Speg. (1)
Crocicreas Fr. (4)
Crucellisporium M.L. Farr (3)
Crumenella P. Karst. (1)
Cryptohymenium Samuels & L.M. Kohn (1)
Cryptopezia Hohn. (1)
Dactylaria Sacc. (100)
Dawsicola Dobbeler (1)
Dermateopsis Nannf. (2)
Didonia Velen. (5)
Didymascella Maire & Sacc. (5)
Discomycella Hohn. (1)
Durella Tul. & C. Tul. (22)*
Echinodiscus Etayo & Diederich (2)
Encoeliopsis Nannf. (4)
Episclerotium L.M. Kohn (2)
Erikssonopsis M. Morelet (1)
Fulvoflamma Crous (1)
Gloeopeziza Zukal (8)
Godroniopsis Diehl & E.K. Cash (3)
Gorgoniceps (P. Karst.) P. Karst. (3)
Grimmicola Dobbeler & Hertel (1)
Grovesia Dennis (1)
Hemiglossum Pat. (2)
Humicolopsis Cabral & S. Marchand (2)
Hydrocina Scheuer (1)
Hymenobolus Durieu & Mont. (3)
Hyphoscypha Velen. (1)
Hysteronaevia Nannf. (12)
Hysteropezizella Hohn. (26)
Hysterostegiella Hohn. (10)
Infundichalara Reblova & W. Gams (2)
Involucroscypha Raitv. (10)
Jacobsonia Boedijn (1)
Korfia J. Reid & Cain (1)
Lareunionomyces Crous & M.J. Wingf. (2)
Larissia Raitv. (1)
Lasseria Dennis (1)
Lemalis Fr. (3)
Libartania Nag Raj (2)
Livia Velen. (1)
Masseea Sacc. (4)
Melanopeziza Velen. (1)
Merodontis Clem. (1)
Microdiscus Sacc. (1)
Mitrulinia Spooner (1)
Monochaetiellopsis B. Sutton & DiCosmo (2)
Mycosphaerangium Verkley (3)

1186
Obconicum Velen. (2)
Obscurodiscus Raitv. (1)
Orbiliopsis (Sacc. & D. Sacc.) Syd. & P. Syd. (2)
Otwaya G. Beaton (12)
Pachydisca Boud. (32)*
Parencoelia Petr. (4)
Patellariopsis Dennis (5)
Patinellaria H. Karst. (1)
Peltigeromyces Möller (3)
Pestalopezia Seaver (3)
Pezolepis Syd. (2)
Pezomela Syd. (1)
Phacidiella P. Karst. (1)
Phaeofabraea Rehm (1)
Phaeopyxis Rambold & Triebel (1)
Phragmonaevia Rehm (16)*
Piceomphale Svrček (1)
Pleoscutula Vou. (3)
Podophacidium Niessl (2)
Polydesmia Boud. (7)
Polyphilus D.G. Knapp, Ashrafi, W. Maier & Kovács (2)
Potridiscus Dobbeler & Triebel (1)
Pseudohelotium Fuckel (50)
Pseudolachnum Velen. (1)
Pseudomitrula Gamundı (1)
Pseudopeltis L. Holm & K. Holm (1)
Pseudotryblidium Rehm (1)
Psilophana Syd. (1)
Pteromyces E. Bommer, M. Rousseau & Sacc. (1)
Pubigera Baral, Gminder & Svrček (1)
Radotinea E. Bommer, M. Rousseau & Sacc. (1)
Rhexocercosporidium U. Braun (2)
Rhizocladosporium Crous & U. Braun (1)
Rhizothyrium Naumov (1)
Rommelaarsia Baral & Haelew. (1)
Roseodiscus Baral (4)
Sageria A. Funk (1)
Sambucina Velen. (1)
Sarcomyces Massee (1)
Sclerocrana Samuels & L.M. Kohn (4)
Scutulopsis Velen. (1)
Soosiella Hujslova & M. Kolarık (1)
Sorokina Sacc. (1)
Sorokinella J. Frohl. & K.D. Hyde (2)
Spirosphaera Beverw. (8)
Stamnaria Fuckel (7)
Stilbopeziza Speg. (1)
Strossmayeria Schulzer (20)
Tetracladium De Wild. (10)
Thedgonia B. Sutton (6)
Themisia Velen. (8)
Tovariella Syd. (1)

1187
Trichohelotium Killerm. (2)
Triposporium Corda (14)
Unguicularia Hohn. (7)*
Urceolella Boud. (44)
Vandijckella Sand.-Den. (1)
Waltonia Saho (1)
Woodiella Sacc. & P. Syd. (3)
Xeromedulla Korf & W.Y. Zhuang (3)
Zugazaea Korf, Iturr. & Lizoň (1)

Lahmiales O.E. Erikss.


Lahmiaceae O.E. Erikss.
Lahmia Korb. (2)

Lauriomycetales Hern.-Restr., R.F. Castañeda & Guarro


Lauriomycetaceae Hern.-Restr., R.F. Castañeda & Guarro
Lauriomyces R.F. Castaneda (11)

Leotiales Korf & Lizoň


Cochlearomycetaceae Crous
Cochlearomyces Crous (1)*
Satchmopsis B. Sutton & Hodges (4)

Leotiaceae Corda
Halenospora E.B.G. Jones (1)
Leotia Pers. (23)
Microglossum Gillet (26)
Miniancora Marvanova & Barl. (1)

Mniaeciaceae Baral*
Epithamnolia Zhurb. (7)*
Mniaecia Boud. (3)

Tympanidaceae Baral & Quijada


Claussenomyces Kirschst. (15)*
Collophorina Damm & Crous (7)
Durandiella Seaver (15)
Gelatinosporium Peck (12)*
Myriodiscus Boedijn (2)
Pragmopora A. Massal. (8)
Tympanis Fr. (64)

Leotiales genera incertae sedis


Aotearoamyces P.R. Johnst., J.A. Cooper & Quijada (1)*
Alatospora Ingold (4)
Flagellospora Ingold (6)*

Lichinodiales M. Prieto, M. Schultz, Olariaga & Wedin


Lichinodiaceae M. Prieto, M. Schultz, Olariaga & Wedin
Lichinodium Nyl. (4)*

1188
Marthamycetales. R. Johnst. & Baral*
Marthamycetaceae Baral, Lantz, Hustad & Minter
Cyclaneusma DiCosmo, Peredo & Minter (2)
Marthamyces Minter (18)
Mellitiosporiella Hohn. (3)
Mellitiosporium Corda (10)
Naemacyclus Fuckel (13)
Phragmiticola Sherwood (1)
Propolina Sacc. (1)
Propolis (Fr.) Corda (8)
Ramomarthamyces P.R. Johnst. (4)*

Medeolariales Korf
Medeolariaceae Korf
Medeolaria Thaxt (1)

Micraspidales Quijada & Tanney*


Micraspidaceae Quijada & Tanney
Micraspis Darker (3)

Phacidiales C.E. Bessey*


Helicogoniaceae Baral
Calloriopsis Syd. & P. Syd. (1)
Eleutheromycella Hohn. (1)
Eleutheromyces Fuckel (2)
Gelatinipulvinella Hosoya & Y. Otani (1)
Gelatinopsis Rambold & Triebe (8)
Geltingia Alstrup & D. Hawksw. (1)
Helicogonium W.L. White (19)

Phacidiaceae Fr.
Allantophomopsiella Crous (1)
Allantophomopsis Petr. (4)
Bulgaria Fr. (12)
Darkera H.S. Whitney, J. Reid & Piroz. (5)
Lophophacidium Lagerb. (2)
Phacidiopycnis Potebnia (6)
Phacidium Fr. (40)
Pseudophacidium P. Karst. (11)
Starbaeckia Rehm ex Starback (1)

Phacidiales genus incertae sedis


Coma Nag Raj & W.B. Kendr. (1)

Rhytismatales M.E. Barr ex Minter


Cudoniaceae P.F. Cannon
Cudonia Fr. (20)
Spathularia Pers. (10)

Rhytismataceae Chevall.
Bifusella Hohn. (9)
Bifusepta Darker (1)

1189
Bivallum P.R. Johnst. (7)
Canavirgella W. Merr, Wenner & Dreisbach (1)
Cavaraella Speg. (1)
Ceratophacidium J. Reid & Piroz. (1)
Cerion Massee (2)
Coccomyces De Not. (119)
Colpoma Wallr. (14)
Criella (Sacc.) Sacc. & P. Syd. (2)
Cryptomyces Grev. (3)
Davisomycella Darker (11)
Discocainia J. Reid & A. Funk (4)
Duplicaria Fuckel (1)
Duplicariella B. Erikss. (1)
Elytroderma Darker (3)
Gelineostroma H.J. Swart (2)
Heufleria Auersw. (2)*
Hypoderma De Not. (56)
Hypodermella Tubeuf (3)
Hypodermellina Höhn. (1)
Hypohelion P.R. Johnst. (4)
Lasiostictella Sherwood (1)
Lirula Darker (12)
Lophodermella Hohn. (9)
Lophodermium Chevall. (185)
Macroderma Hohn. (2)
Meloderma Darker (5)
Moutoniella Penz. & Sacc. (1)
Mycomelanea Velen. (1)
Myriophacidium Sherwood (6)
Nematococcomyces C.L. Hou, M. Piepenbr. & Oberw. (2)
Neococcomyces Y.R. Lin, C.T. Xiang & Z.Z. Li (3)
Neophacidium Petr. (2)
Nothorhytisma Minter, P.F. Cannon, A.I. Romero & Peredo (1)
Parvacoccum R.S. Hunt & A. Funk (1)
Phaeophacidium P. Henn. & Lindau (3)
Ploioderma Darker. (8)
Propolidium Sacc. (15)
Pseudographis Nyl. (10)*
Pseudorhytisma Juel (1)
Pureke P.R. Johnst. (1)
Rhytisma Fr. (30)
Soleella Darker (7)
Sporomega Corda (1)
Terriera B. Erikss. (34)
Therrya Sacc. (7)
Tryblidiopsis P. Karst. (5)
Virgella Darker (1)
Vladracula P.F. Cannon, Minter & Kamal (2)
Xyloschizon Syd. (2)
Zeus Minter & Diamandis (1)

Triblidiaceae Rehm*

1190
Huangshania O.E. Erikss. (2)
Triblidium Rebent. (13)

Rhytismatales genera incertae sedis


Apiodiscus Petr. (1)
Bonanseja Sacc. (1)
Didymascus Sacc. (2)
Haplophyse Theiss. (1)
Irydyonia Racib. (1)
Laquearia Fr. (2)
Mycosymbioces J.L. Frank (1)
Nymanomyces P. Henn. (2)
Pseudotrochila Hohn. (1)

Thelebolales P.F. Cannon


Pseudeurotiaceae Malloch & Cain
Connersia Malloch (1)
Geomyces Traaen (9)
Gymnostellatospora Udagawa, Uchiy. & Kamiya (6)*
Leuconeurospora Malloch & Cain (2)
Neelakesa Udaiyan & Hosag. (3)
Pleuroascus Massee & E.S. Salmon (3)
Pseudeurotium J.F.H. Beyma (8)
Pseudogymnoascus Raillo (12)*

Thelebolaceae (Brumm.) Eckblad


Antarctomyces Stchigel & Guarro (2)
Ascophanus Boud. (56)
Ascozonus (Renny) E.C. Hansen (9)
Caccobius Kimbr. (1)
Cleistothelebolus Malloch & Cain (1)
Coprobolus Cain & Kimbr. (1)
Leptokalpion Brumm. (1)
Pseudascozonus Brumm. (1)
Ramgea Brumm. (2)
Thelebolus Tode (16)

Leotiomycetes genera incertae sedis


Adelodiscus Syd. (1)
Bagnisimitrula S. Imai (1)
Callerascus Whitton, K.D. Hyde & McKenzie (1)
Deltopyxis Baral & G. Marson (1)
Epicladonia D. Hawksw. (5)
Gorgomyces M. Gonczol & Revay (2)
Helicocentralis Sri-indr., Chuaseehar., Boonyuen, K. Yamag., Suetrong & C.K.M. Tsui (1)
Helotiella Sacc. (17)
Holwaya Sacc. (2)
Leohumicola N.L. Nick. (7)
Melanormia Korb. (1)
Metapezizella Petr. (1)
Ocotomyces H.C. Evans & Minter (1)
Patinella Sacc. (25)*

1191
Phyllopezis Petr. (1)
Physmatomyces Rehm (1)
Polydiscina Syd. (1)
Psilothecium Clem. (1)
Schnablia Sacc. & P. Syd. (1)
Trullula Ces. (5)

Lichinomycetes V. Reeb, Lutzoni & Cl. Roux


Lichinales Henssen & Büdel
Gloeoheppiaceae Henssen
Gloeoheppia Gyeln. (5)
Gudelia Henssen (1)
Pseudopeltula Henssen (1)

Lichinaceae Nyl.
Anema Nyl. ex Forssell (21)
Calotrichopsis Vain. (4)
Corynecystis Brusse (1)
Cryptothele Th. Fr. (7)
Digitothyrea P. Moreno & Egea (3)
Edwardiella Henssen (1)
Ephebe Fr. (13)
Finkia Vain. (1)
Gyrocollema Vain. (2)
Heppia Nägeli (4)
Jenmania W. Wächt. (2)
Lecidopyrenopsis Vain. (1)
Lemmopsis (Vain.) Zahlbr. (3)
Lempholemma Körb. (35)
Leprocollema Vain. (3)
Lichina C. Agardh (9)
Lichinella Nyl. (30)
Mawsonia C.W. Dodge (1)
Metamelanea Henssen (3)
Paulia Feé (10)
Peccania A. Massal. ex Arnold (3)
Phloeopeccania J. Steiner (4)
Phylliscidiopsis Sambo (1)
Phylliscidium Forssell (1)
Phyllisciella Henssen & Büdel (3)
Phylliscum Nyl. (8)
Porocyphus Körb. (8)
Pseudarctomia Gyeln. (1)
Pseudoheppia Zahlbr. (1)
Pseudopaulia M. Schultz (1)
Psorotichia A. Massal. (50)
Pterygiopsis Vain. (17)
Pyrenocarpon Trevis. (1)
Pyrenopsis Nyl. (40)
Solorinaria (Vain.) Gyeln. (1)
Stromatella Henssen (1)
Synalissa Fr. (30)

1192
Thallinocarpon A.E. Dahl (2)
Thelignya A. Massal. (2)
Thermutis Fr. (2)
Thermutopsis Henssen (1)
Thyrea A. Massal. (13)
Zahlbrucknerella Herre (10)

Peltulaceae Büdel
Peltula Nyl. (32)

Orbiliomycetes O.E. Erikss. & Baral


Orbiliales Baral, O.E. Erikss., G. Marson & E. Weber
Orbiliaceae Nannf.
Arthrobotrys Corda (ca. 100+)
Dactylella Grove (31)
Dactylellina M. Morelet (= Gamsylella M. Scholler et al.) (26)
Drechslerella Subram. (ca. 7)
Dwayaangam Subram. (8)
Helicoon Morgan (ca. 15)
Hyalorbilia Baral & G. Marson (40)
Orbilia Fr. (ca. 400)
Pseudotripoconidium Z.F. Yu & K.Q. Zhang (1)
Pseudorbilia Y. Zhang, Z.F. Yu, Baral & K-Q Zhang (1)
Retiarius D.L. Olivier (4)
Vermispora Deighton & Piroz. (7)

Orbiliales genus incertae sedis


Microdochiella Hern.-Restr. & Crous (1)

Orbiliomycetes genus incertae sedis


Mycoceros D. Magyar & Z. Merényi (1)*

Pezizomycetes O.E. Erikss. & Winka


Pezizales J. Schröt.
Ascobolaceae Boud. ex Sacc.
Ascobolus Pers. (ca. 70)
Cleistoiodophanus J.L. Bezerra & Kimbr. (1)
Cubonia Sacc. (ca. 7)
Saccobolus Boud. (33)
Thecotheus Boud. (23)*

Ascodesmidaceae J. Schröt.
Ascodesmis Tiegh. (~10)
Cephaliophora Thaxt. (2)*
Chalazion Dissing & Sivertsen (3)
Coprotiella Jeng & J.C. Krug (1)
Dictyocoprotus J.C. Krug & R.S. Khan (1)
Eleutherascus Arx (4)
Lasiobolus Sacc. (11)
Luciotrichus R. Galán & Raitv. (1)
Ochotrichobolus Kimbr. & Korf (1)
Trichobolus (Sacc.) Kimbr. & Cain (6)

1193
Caloscyphaceae Harmaja
Caloscypha Boud. (2)

Chorioactidaceae Pfister
Chorioactis Kupfer ex Eckblad (1)
Desmazierella Lib. (2)
Neournula Paden & Tylutki (2)
Pseudosarcosoma M. Carbone, Agnello & P. Alvarado (1)
Trichaleurina Rehm (3)
Wolfina Seaver ex Eckblad (2)

Discinaceae Benedix
Discina (Fr.) Fr. (20)
Gymnohydnotrya B.C. Zhang & Minter (3)
Gyromitra Fr. (25)
Hydnotrya Berk. & Broome (11)
Pseudorhizina Jacz. (3)

Glaziellaceae J.L. Gibson


Glaziella Berk. (1)

Helvellaceae Fr.
Balsamia Vittad. (21)
Barssia Gilkey (8)
Helvella L. (ca. 80)
Underwoodia Peck (2)
Wynnella Boud. (3)

Kallistoskyphaceae Ekanayaka, K.D. Hyde, Q. Zhao & E.B.G. Jones


Kallistoskypha Pfister, Agnello, Lantieri & LoBuglio (1)

Karstenellaceae Harmaja
Karstenella Harmaja (1)

Morchellaceae H.G.L. Reichenbach


Disciotis Boud. (3)
Fischerula Mattir. (2)
Imaia Trappe & Kovács (1)
Kalapuya M.J. Trappe, Trappe & Bonito (1)
Leucangium Quél. (1)
Morchella Dill. ex Pers. (~60)
Verpa Sw. (4)

Pezizaceae Dumort. (= Carbomycetaceae Trappe)


Adelphella Pfister, Matočec & I. Kušan (1)
Amylascus Trappe (1)
Antrelloides P.S. Catches. & D.E.A. Catches. (1)
Aquapeziza D.M. Hu, L. Cai & K.D. Hyde (1)
Boudiera Cooke (10)
Calongea Healy, Bonito & Trappe (1)*
Carbomyces Gilkey (3)
Cazia Trappe (2)

1194
Delastria Tul. & C. Tul. (6)
Elderia McLennan (1)
Eremiomyces Trappe & Kagan-Zur (3)
Galactinia (Cooke) Boud. (ca. 5)
Hapsidomyces J.C. Krug & Jeng (1)
Hydnobolites Tul. & C. Tul. (ca. 6)
Hydnotryopsis Gilkey (4)
Iodophanus Korf (15)
Iodowynnea Medel, Guzmán & S. Chacón (1)
Kalaharituber Trappe & Kagan-Zur (1)
Lepidotia Boud. (1)
Luteoamylascus Cabero, P. Alvarado & G. Moreno (1)
Marcelleina Brumm., Korf & Rifai (11)
Mattirolomyces E. Fisch. (5)
Mycoclelandia Trappe & G.W. Beaton (2)
Pachyella Boud. (12)
Pachyphlodes Zobel (ca. 10)*
Peziza Dill. ex Fr. (ca. 120)
Plicaria Fuckel (10)
Plicariella (Sacc.) Rehm (2)*
Rhodopeziza Hohmeyer & Moravec (1)
Ruhlandiella P. Henn. (7)
Sarcopeziza Loizides, Agnello & P. Alvarado (1)*
Sarcosphaera Auersw. (4)
Sphaerozone Zobel (1)
Stouffera Kovács & Trappe (1)
Temperantia K. Hansen, Healy & Kovács (1)
Terfezia (Tul. & C. Tul.) Tul. & C. Tul. (19)
Tirmania Chatin (3)
Ulurua Trappe, Claridge & Kovács (1)

Pseudombrophilaceae Ekanayaka, K.D. Hyde, Q. Zhao & E.B.G. Jones


Heydenia Fresen. (3)
Lasiobolidium Malloch & Cain (7)
Orbicula Cooke (1)
Pseudombrophila Boud. (37)

Pulvinulaceae Ekanayaka, K.D. Hyde, Q. Zhao & E.B.G. Jones


Lazuardia Rifai (1)
Pseudoboubovia U. Lindem., M. Vega, B. Perić & R. Tena (1)
Pulvinula Boud. (~30)

Pyronemataceae Corda (= Otideaceae Eckblad)


Acervus Kanouse (9)
Aleuria Fuckel (ca. 10)
Aleurina Massee (ca. 10)
Anthracobia Boud. (ca. 10)
Arpinia Berthet (4)
Ascosparassis Kobayasi (1)
Byssonectria P. Karst. (7)*
Chaetothiersia B.A. Perry & Pfister (1)
Cheilymenia Boud. (67)

1195
Cupulina Dougoud, Van Vooren & M. Vega (2)
Diehliomyces Gilkey (1)
Eoaleurina Korf & W.Y. Zhuang (1)
Galeoscypha Svrček & J. Moravec (1)
Genabea Tul. & C. Tul. (4)
Genea Vittad. (ca. 40)
Geneosperma Rifai (2)
Geopora Harkn. (ca. 20)
Gilkeya M.E. Sm., Trappe & Rizzo (1)
Hoffmannoscypha Stielow, Göker & Klenk (1)*
Humaria Fuckel (ca. 10)
Jafnea Korf (2)
Lamprospora De Not. (ca. 50)
Lasiocupulina Van Vooren & M. Vega (1)
Lathraeodiscus Dissing & Sivertsen (1)
Lotinia Pérez-Butrón Fern.-Vic. & P. Alvarado (1)*
Melastiza Boud. (ca. 10)
Micronematobotrys Xiang Sun & L.D. Guo (1)
Miladina Svrček (1)
Monascella Guarro & Arx (1)
Myrmecocystis Harkn. (7)*
Neottiella (Cooke) Sacc. (ca. 5)
Octospora Hedw. (ca. 50)
Octosporopsis U. Lindem. & M. Vega (2)
Otidea (Pers.) Bonord. (ca. 52)
Oviascoma Y.J. Yao & Spooner (1)
Parascutellinia Svrček (6)
Paratricharina Van Vooren, U. Lindemann, M. Vega, Ribes, Illescas & Matočec (1)
Paratrichophaea Trigaux (5)
Petchiomyces E. Fisch. & Mattir. (1)
Picoa Vittad. (2)*
Planamyces Crous & Decock (1)
Pseudaleuria Lusk (2)
Pseudotricharina Van Vooren, Tello & M. Vega (3)
Pyronema Carus (3)
Pyropyxis Egger (1)
Ramsbottomia W.D. Buckley (3)
Rhizoblepharia Rifai (2)
Scutellinia (Cooke) Lambotte (70)
Selenaspora R. Heim & Le Gal (1)
Sepultariella Van Vooren, U. Lindemmann & Healy (2)*
Smardaea Svrček (9)
Smarodsia Raitv. & Vimba (1)
Sowerbyella Nannf. (17)
Sphaerosporella (Svrček) Svrček & Kubička (3)
Sphaerosporium Schwein. sensu stricto (1)*
Spooneromyces T. Schumach. & J. Moravec (5)
Tricharina Eckblad (=Ascorhizoctonia Chin S. Yang & Korf) (12)
Trichophaea Boud. (26)
Trichophaeopsis Korf & Erb (4)
Warcupia Paden & J.V. Cameron (1)
Wenyingia Zheng Wang & Pfister (1)

1196
Wilcoxina Chin S. Yang & Korf (5)

Rhizinaceae Bonord.
Phymatotrichopsis Hennebert (1)
Psilopezia Berk. (7)
Rhizina Fr. (1)

Sarcoscyphaceae LeGal ex Eckblad


Aurophora Rifai (1)
Cookeina Kuntze (11)
Geodina Denison (1)
Kompsoscypha Pfister (4)
Microstoma Bernstein (7)
Nanoscypha Denison (8)
Phillipsia Berk. (~20)
Pithya Fuckel (2)
Pseudopithyella Seaver (2)
Sarcoscypha (Fr.) Boud. (18)
Thindia Korf & Waraitch (1)
Wynnea Berk. & M.A. Curtis (7)

Sarcosomataceae Kobayasi
Conoplea Pers. (11)
Donadinia Bellem. & Mel.-Howell (5)
Galiella Nannf. & Korf (9)
Korfiella D.C. Pant & V.P. Tewari (1)
Plectania Fuckel (ca. 20)
Pseudoplectania Fuckel (4)
Sarcosoma Casp. (5)
Strumella Fr. (8)
Urnula Fr. (9)

Strobiloscyphaceae Ekanayaka, K.D. Hyde, Q. Zhao & E.B.G. Jones


Strobiloscypha N.S. Weber & Denison (2)

Tarzettaceae Ekanayaka, K.D. Hyde, Q. Zhao & E.B.G. Jones


Densocarpa Gilkey (2)*
Geopyxis (Pers.) Sacc. (7)
Hydnocystis Tul. (= Stephensia Tul. & C. Tul.) (5)*
Hypotarzetta Donadini (1)
Paurocotylis Berk. (4)
Tarzetta (Cooke) Lambotte (ca. 10)

Tuberaceae Dumort.
Choiromyces Vittad. (5)
Dingleya Trappe (6)
Labyrinthomyces Boedijn (1)
Nothojafnea Rifai (2)
Paradoxa Mattir. (2)
Reddellomyces Trappe, Castellano & Malajczuk (4)
Tuber P. Micheli ex F.H. Wigg. (ca. 120)

1197
Pezizales genera incertae sedis
Aparaphysaria Speg. (1)
Ascocalathium Eidam ex J. Schröt. (1)
Boubovia Svrček (5)*
Boudierella Sacc. (1)
Cidaris Fr. (1)
Coprotus Korf ex Korf & Kimbr. (33)*
Dennisiopsis Subram. & Chandras. (2)
Filicupula Y.J. Yao & Spooner (1)
Heydenia Fresen.
Hiemsia Svrček (2)
Leucoscypha Boud (4)
Microeurotium Ghatak (1)
Moravecia Benkert, Caillet & Moyne (2)
Mycoarctium K.P. Jain & Cain (2)
Mycogalopsis Gjurašin (1)
Octosporella Döbbeler (9)
Orcadia G.K. Sutherl. (1)
Sphaerosoma Klotzsch (3)

Sordariomycetes O.E. Erikss. & Winka


Diaporthomycetidae Senan., Maharachch. & K.D. Hyde
Annulatascales M.J. D'souza, Maharachch. & K.D. Hyde
Annulatascaceae S.W. Wong, K.D. Hyde & E.B.G. Jones
Annulatascus K.D. Hyde (18)
Annulusmagnus J. Campb. & Shearer (1)
Aqualignicola Ranghoo, C.K.M. Tsui & K.D. Hyde (2)
Ascitendus J. Campb. & Shearer (2)
Ayria Fryar & K.D. Hyde (2)
Cataractispora K.D. Hyde, S.W. Wong & E.B.G. Jones (5)
Chaetorostrum Zelski, Raja, A.N. Mill. & Shearer (1)
Longicollum Zelski, F.R. Barbosa, Raja, A.N. Mill. & Shearer (1)
Submersisphaeria K.D. Hyde (5)
Vertexicola K.D. Hyde, Ranghoo & S.W. Wong (3)

Annulatascales genus incertae sedis


Clohiesia K.D. Hyde (3)

Atractosporales H. Zhang, K.D. Hyde & Maharachch.


Atractosporaceae H. Zhang, K.D. Hyde & Maharachch.
Atractospora Réblová & J. Fourn. (5)
Rubellisphaeria Réblová & J. Fourn. (1)

Conlariaceae H. Zhang, K.D. Hyde & Maharachch.


Conlarium F. Liu & L. Cai (3)
Riomyces A. Ferrer, A.N. Mill., Sarmiento & Shearer (1)

Pseudoproboscisporaceae H. Zhang, K.D. Hyde & Maharachch.


Diluviicola K.D. Hyde, S.W. Wong & E.B.G. Jones (2)
Pseudoproboscispora Punith. (3)

Calosphaeriales M.E. Barr

1198
Calosphaeriaceae Munk
Calosphaeria Tul. & C Tul. (114)
Flabellascus Réblová (1)
Jattaea Berl (27)
Togniniella Réblová, L. Mostert, W. Gams & Crous (1)

Pleurostomataceae Réblová, L. Mostert, W. Gams & Crous


Pleurostoma Tul. & C. Tul. (7)

Calosphaeriales genera incertae sedis


Calosphaeriopsis Petr. (1)
Enchnoa Fr. (21)
Kacosphaeria Speg. (1)
Sulcatistroma A.W. Ramaley (1)

Diaporthales Nannf.
Apiosporopsidaceae Senan., Maharachch. & K.D. Hyde
Apiosporopsis (Traverso) Mariani. (3)

Apoharknessiaceae Senan., Maharachch. & K.D. Hyde


Apoharknessia Crous & S.J. Lee (3)
Lasmenia Speg. (5)

Asterosporiaceae Senan., Maharachch. & K.D. Hyde


Asterosporium Kunze (5)

Auratiopycnidiellaceae Senan., Maharachch. & K.D. Hyde


Auratiopycnidiella Crous & Summerell (1)

Coryneaceae Corda (=Pseudovalsaceae M.E. Barr)


Coryneum Nees (30)

Cryphonectriaceae Gryzenh. & M.J. Wingf.


Amphilogia Gryzenh., H.F. Glen & M.J. Wingf. (2)
Aurantioporthe G. Beier & R.A. Blanchette (1)
Aurantiosacculus Dyko & B. Sutton (3)
Aurapex Gryzenh. & M.J. Wingf. (1)
Aurifilum Begoude, Gryzenh. & Jol. Roux (1)
Capillaureum M.E.S. Oliveira (1)
Celoporthe Nakab., Gryzenh., Jol. Roux & M.J. Wingf. (2)
Chromendothia Lar.N. Vassiljeva (2)
Chrysofolia Crous & M.J. Wingf. (1)
Chrysomorbus S.F. Chen (1)
Chrysoporthe Gryzenh. & M.J. Wingf. (9)
Corticimorbus S.F. Chen & M.J. Wingf. (1)
Cryphonectria (Sacc.) Sacc. & D. Sacc. (1)
Cryptometrion Gryzenh. & M.J. Wingf. (1)
Diversimorbus S.F. Chen & J. Roux (1)
Endothia Fr. (2)
Eriocamporesia R.H. Perera, Samarak. & K.D. Hyde (1)
Foliocryphia Cheew. & Crous (2)
Holocryphia Gryzenh. & M.J. Wingf. (1)

1199
Immersiporthe S.F. Chen, M.J. Wingf. & Jol. Roux (1)
Latruncellus M. Verm., Gryzenh. & Jol. Roux (1)
Luteocirrhus C.F. Crane & T.I. Burgess (1)
Mastigosporella Höhn. (=Wuestneiopsis J. Reid & Dowsett) (5)
Microthia Gryzenh. & M.J. Wingf. (2)
Myrtonectria Marinc., D.B. Ali & J. Roux (1)
Rostraureum Gryzenh. & M.J. Wingf. (2)
Ursicollum Gryzenh. & M.J. Wingf. (1)
Wuestneia Auersw. ex Fuckel (13)

Cytosporaceae Fr. (= Valsaceae Tul. & C. Tul.)


Cryptascoma Ananthap. (2)
Cytospora Ehrenb. (123)
Pachytrype Berl. ex M.E. Barr, J.D. Rogers & Y.M. Ju (1)
Paravalsa Ananthap. (1)
Waydora B. Sutton (1)
Xenotypa Petr. (1)

Diaporthaceae Höhn. ex Wehm.


Apioporthella Petr. (1)
Apiosphaeria Höhn. (5)
Chaetoconis Clem. (1)
Chiangraiomyces Senan. & K.D. Hyde (1)
Diaporthe Nitschke (= Allantoporthe Petr.; = Clypeoporthella Petr.) (173)
Hyaliappendispora Senan., Camporesi & K.D. Hyde (1)
Leucodiaporthe M.E. Barr & Lar.N. Vassiljeva (1)
Massariothea Syd. (10)
Mazzantia Mont. (4)
Ophiodiaporthe Y.M. Ju, H.M. Hsieh, C.H. Fu, Chi Y. Chen & T.T. Chang (1)
Paradiaporthe Senan., Camporesi & K.D. Hyde (1)
Phaeocytostroma Petr. (4)
Phaeodiaporthe Petr. (2)
Pustulomyces D.Q. Dai, Bhat & K.D. Hyde (1)
Stenocarpella Syd. & P. Syd. (2)

Diaporthosporellaceae C.M. Tian & Q. Yang*


Diaporthosporella C.M. Tian & Q. Yang (1)

Diaporthostomataceae X.L. Fan & C.M. Tian*


Diaporthostoma X.L. Fan & C.M. Tian (1)*

Dwiroopaceae K.V. Xavier, A.N. KC, J.Z. Groenew., Vallad & Crous
Dwiroopa Subram. & Muthumary (2)

Erythrogloeaceae Senan., Maharachch. & K.D. Hyde


Chrysocrypta Crous & Summerell (1)
Dendrostoma X.L. Fan & C.M. Tian (4)*
Disculoides Crous, Pascoe, I.J. Porter & Jacq. Edwards (2)
Erythrogloeum Petr. (2)

Gnomoniaceae G. Winter
Alnecium Voglmayr & Jaklitsch (2)

1200
Ambarignomonia Sogonov (1)
Amphiporthe Petr. (=Amphicytostroma Petr.) (2)
Anisomyces Theiss. & Syd. (5)
Apiognomonia Höhn. (=Discula Sacc.) (28)
Apioplagiostoma M.E. Barr (3)
Asteroma DC. (54)
Bagcheea E. Müll. & R. Menon (2)
Chadefaudiomyces Kamat (1)
Clypeoporthe Höhn. (5)
Cryptosporella Sacc. (ca. 26)
Dictyoporthe Petr. (4)
Diplacella Syd. (2)
Ditopella De Not. (16)
Ditopellopsis J. Reid & C. Booth (4)
Flavignomonia C.M. Tian, Qin Yang & N. Jiang (1)*
Gloeosporidina Petr. (6)
Gnomonia Ces. & De Not. (ca. 70)
Gnomoniella Sacc. (=Cylindrosporella Höhn.) (ca. 70)
Gnomoniopsis Berl. (25)
Maculatipalma J. Fröhlich & K.D. Hyde (1)
Mamianiella Höhn. (=Anisogramma Theiss. & Syd.; = Mamiania Ces & De Not.) (2)
Marsupiomyces Senan. & K.D. Hyde (2)
Millerburtonia Cif. (1)
Occultocarpon L.C. Mejía & Zhu L. Yang (1)
Ophiognomonia (Sacc.) Sacc. (49)
Phragmoporthe Petr. (1)
Phylloporthe Syd. (2)
Plagiostoma Fuckel (52)
Pleuroceras Riess. (12)
Sirococcus Preuss (5)
Spataporthe Bronson, Klymiuk, Stockey & Tomescu (1)
Tenuignomonia Minosh., D.M. Walker & Hirooka (1)
Uleoporthe Petr. (1)
Valsalnicola D.M. Walker & Rossman (1)
Vismaya V.V. Sarma & K.D. Hyde (1)

Harknessiaceae Crous
Harknessia Cooke (ca. 50)
Mebarria J. Reid & C. Booth (1)

Juglanconidaceae Voglmayr & Jaklitsch (=Melanosporellaceae C.M. Tian & Z. Du)


Juglanconis Voglmayr & Jaklitsch (4)

Lamproconiaceae Norph., T.C. Wen & K.D. Hyde


Hercospora Fr. (= Rabenhorstia Fr.) (1)
Lamproconium (Grove) Grove (1)

Macrohilaceae Crous
Macrohilum H.J. Swart (1)

Melanconidaceae G. Winter
Melanconis Tul. & C. Tul. (1)

1201
Melanconiellaceae Senan., Maharachch. & K.D. Hyde
Dicarpella Syd. & P. Syd. (7)
Greeneria Scribn. & Viala (3)
Massariovalsa Sacc. (=Melanconiopsis Ellis & Everh.) (4)
Melanconiella Sacc. (2)
Microascospora Senan. & K.D. Hyde (2)
Septomelanconiella Samarak. & K.D. Hyde (1)*
Sheathospora X.L. Fan (1)*
Sphaeronaemella P. Karst. sensu lato (10)

Neomelanconiellaceae Crous
Neomelanconiella Crous (1)*

Phaeoappendicosporaceae Crous & M.J. Wingf.


Phaeoappendicospora Senan., Q.R. Li & K.D. Hyde (1)
Neophaeoappendicospora Crous & M.J. Wingf. (1)

Prosopidicolaceae Senan. & K.D. Hyde


Prosopidicola Crous & C.L. Lennox (2)

Pseudomelanconidaceae C.M. Tian & X.L. Fan*


Pseudomelanconis C.M. Tian & X.L. Fan (1)*
Neopseudomelanconis C.M. Tian & N. Jiang (1)

Pseudoplagiostomataceae Cheew., M.J. Wingf. & Crous


Pseudoplagiostoma Cheew., M.J. Wingf. & Crous (7)

Schizoparmaceae Rossman
Coniella Höhn. (34)

Stilbosporaceae Link
Crinitospora B. Sutton & Alcorn (1)
Natarajania Pratibha & Bhat (1)
Stegonsporium Corda (8)
Stilbospora Pers. (20)

Sydowiellaceae Lar.N. Vassiljeva


Alborbis Senan. & K.D. Hyde (1)
Breviappendix Senan. & K.D. Hyde (3)
Cainiella E. Müll. (2)
Calosporella J. Schröt (1)
Caudospora Starbäck (2)
Chapeckia M.E. Barr (2)
Hapalocystis Auersw. ex Fuckel (9)
Italiomyces Senan., Camporesi & K.D. Hyde (1)
Lambro Racib. (3)
Paragnomonia Senan. & K.D. Hyde (1)
Ranulospora Senan., Camporesi & K.D. Hyde (1)
Rossmania Lar.N. Vassiljeva (2)
Sillia P. Karst. (9)
Sydowiella Petr. (11)
Tenuiappendicula Senan., Camporesi & K.D. Hyde (1)

1202
Tortilispora Senan. & K.D. Hyde (3)

Synnemasporellaceae X.L. Fan & J.D.P. Bezerra*


Synnemasporella X.L. Fan & J.D.P. Bezerra (2)*

Tubakiaceae U. Braun, J.Z. Groenew. & Crous*


Apiognomonioides U. Braun, J.Z. Groenew. & Crous (1)
Involutscutellula U. Braun & C. Nakash. (1)
Oblongisporothyrium U. Braun & C. Nakash. (1)
Paratubakia U. Braun & C. Nakash. (2)
Racheliella Crous & U. Braun (2)
Saprothyrium U. Braun, Crous & J.Z. Groenew. (1)
Sphaerosporithyrium U. Braun, Crous, O. Moreno-Rico & Marm. (1)
Tubakia B. Sutton (25)

Diaporthales genera incertae sedis


Ceratoporthe Petr. (1)
Cryptoleptosphaeria Petr. (1)
Cryptonectriella (Höhn.) Weese (2)
Cryptonectriopsis (Höhn.) Weese (1)
Cytomelanconis Naumov (1)
Diaporthella Petr. (5)
Diatrypoidiella Manohar., Kunwar & D.K. Agarwa (1)
Ditopellina J. Reid & C. Booth (1)
Durispora K.D. Hyde (2)
Exormatostoma Gray (10 epithets in Index Fungorum 2020)
Fremineavia Nieuwl. (1)
Gibellia Sacc. (1)
Gyrostroma Naumov (3)
Hyalorostratum Raja & Shearer (1)
Hypophloeda K.D. Hyde & E.B.G. Jones (1)
Hypospilina (Sacc) Traverso (4)
Kapooria J. Reid & C. Booth (1)
Keinstirschia J. Reid & C. Booth (1)
Kensinjia J. Reid & C. Booth (1)
Lollipopaia Inderb. (1)
Macrodiaporthe Petr. (1)
Melanamphora Lafl. (1)
Phragmodiaporthe Wehm. (3)
Phruensis Pinruan (1)
Plagiophiale Petr. (2)
Plagiostigme Syd. (1)
Prostratus Sivan., W.H. Hsieh & Chi Y. Chen (1)
Pseudocryptosporella J. Reid & C. Booth (1)
Pseudothis Theiss. & Syd. (12)
Pseudovalsella Höhn. (2)
Savulescua Petr. (1)
Skottsbergiella Petr. (1)
Sphaerognomoniella Naumov & Kusnezowa (1)
Stioclettia Dennis (1)
Trematovalsa Jacobesco (1)
Wehmeyera J. Reid & C. Booth (1)

1203
Distoseptisporales Z.L. Luo, K.D. Hyde & H.Y. Su
Distoseptisporaceae K.D. Hyde & McKenzie
Distoseptispora K.D. Hyde, McKenzie & Maharachch. (18)

Magnaporthales Thongk., Vijaykr. & K.D. Hyde


Ceratosphaeriaceae Z.L. Luo, H.Y. Su & K.D. Hyde
Ceratosphaeria Niessl. (24)

Magnaporthaceae P.F. Cannon


Bifusisporella R.M.F. Silva, R.J.V. Oliveira, J.D.P. Bezerra, Souza-Motta & G.A. Silva (1)*
Budhanggurabania P. Wong, Khemmuk & R.G. Shivas (1)
Buergenerula Syd. (1)
Bussabanomyces Klaubauf, M.-H. Lebrun & Crous (1)
Ceratosphaerella Huhndorf, Greif, Mugambi & A.N. Mill. (2)
Clasterosphaeria Sivan. (2)
Clasterosporium Schwein (41)
Clavatisporella K.D. Hyde (1)
Falciphora J. Luo & N. Zhang (1)
Falciphoriella M. Hern.-Restr. & Crous (1)
Gaeumannomycella M. Hern.-Restr. & Crous (2)
Gaeumannomyces Arx & D.L. Olivier (20)
Herbampulla Scheuer & Nograsek (1)
Kohlmeyeriopsis Klaubauf, M.-H. Lebrun & Crous (1)
Magnaporthiopsis J. Luo & N. Zhang (7)
Muraeriata Huhndorf, Greif, Mugambi & A.N. Mill. (2)
Nakataea Hara (8)
Neogaeumannomyces D.Q. Dai & K.D. Hyde (1)
Omnidemptus P.F. Cannon & Alcorn (3)
Plagiosphaera Petr. (1)*
Pseudophialophora J. Luo & N. Zhang (9)
Pyriculariopsis M.B. Ellis (9)
Slopeiomyces Klaubauf, M.-H. Lebrun & Crous (1)

Ophioceraceae Klaubauf, E.G. LeBrun & Crous


Ophioceras Sacc. (50)

Pseudohalonectriaceae Hongsanan & K.D. Hyde


Pseudohalonectria Minoura & T. Muroi (16)

Pyriculariaceae Klaubauf, E.G. LeBrun & Crous


Bambusicularia Klaubauf, M.-H. Lebrun & Crous (1)
Barretomyces Klaubauf, M.-H. Lebrun & Crous (1)
Deightoniella S. Hughes (20)
Macgarvieomyces Klaubauf, M.-H. Lebrun & Crous (3)
Neocordana Hern.-Rest. & Crous (6)
Neopyricularia Klaubauf, M.-H. Lebrun & Crous (1)
Proxipyricularia Klaubauf, M.-H. Lebrun & Crous (2)
Pseudopyricularia Klaubauf, M.-H. Lebrun & Crous (7)
Pyricularia Sacc. (84)
Pyriculariomyces Y. Marín, M.J. Wingf. & Crous (1)
Xenopyricularia Klaubauf, M.-H. Lebrun & Crous (1)

1204
Myrmecridiales Crous
Myrmecridiaceae Crous
Myrmecridium Arzanlou, W. Gams & Crous (14)
Neomyrmecridium Crous (2)

Xenodactylariaceae Crous
Xenodactylaria Crous (1)*

Ophiostomatales Benny & Kimbr.


Kathistaceae Malloch & M. Blackw.
Kathistes Malloch & M. Blackw. (3)
Mattirolella S. Colla (2)
Termitariopsis M. Blackw., Samson & Kimbr. (1)

Ophiostomataceae Nannf.
Afroraffaele C.C. Bateman, Y.T. Huang & D.R. Simmons (1)
Aureovirg J.A. van der Linde, Z.W. de Beer & Jol. Roux (1)
Ceratocystiopsis H.P. Upadhyay & W.B. Kendr. (5)
Fragosphaeria Shear (2)
Graphilbum H.P. Upadhyay & W.B. Kendr. (13)
Hawksworthiomyces Z.W. de Beer, Marinc. & M.J. Wingf. (4)
Klasterskya Petr. (3)
Leptographium Lagerb. & Melin (= Grosmannia Gold.) (74)
Ophiostoma Syd. & P. Syd. (= Hyalorhinocladiella H.P. Upadhyay & W.B. Kendr.; =
Pesotum J.L. Crane & Schokn.) (134)
Raffaelea Arx & Hennebert (33)
Sporothrix Hektoen & C.F. Perkins (79)
Spumatoria Massee & E.S. Salmon (1)
Subbaromyces Hesselt. (2)

Phomatosporales Senan., Maharachch. & K.D. Hyde


Phomatosporaceae Senan. & K.D. Hyde
Lanspora K.D. Hyde & E.B.G. Jones (2)
Phomatospora Sacc. (ca. 100)
Tenuimurus Senan., Camporesi & K.D. Hyde (1)

Sporidesmiales Crous
Sporidesmiaceae Fr.
Sporidesmium Link (ca. 330)

Tirisporellales Suetrong, E.B.G. Jones & K.L. Pang


Tirisporellaceae Suetrong, E.B.G. Jones & K.L. Pang
Bacusphaeria Norlail., Alias & Suetrong (1)
Thailandiomyces Pinruan, Sakay., K.D. Hyde & E.B.G. Jones (1)
Tirisporella E.B.G. Jones, K.D. Hyde & Alias (1)

Togniniales Senan., Maharachch. & K.D. Hyde


Togniniaceae Réblová, L. Mostert, W. Gams & Crous
Conidiotheca Réblová & L Mostert (1)
Phaeoacremonium W. Gams, Crous & M.J. Wingf. (65)

Xenospadicoidales Hern.-Restr., J. Mena & Gené

1205
Xenospadicoidaceae Hern.-Restr., J. Mena & Gené (= Lentomitellaceae H. Zhang, K.D. Hyde &
Maharachch)*
Calyptosphaeria Réblová & A.N. Mill. (4)
Lentomitella Höhn. (13)
Neospadicoides Z.L. Luo (3)
Spadicoides S. Hughes (=Xenospadicoides Hern.-Restr., J. Mena & Gené;
Pseudodiplococcium Hern.-Restr., J. Mena & Gené) (45)*
Torrentispora K.D. Hyde, W.H. Ho, E.B.G. Jones (= Fusoidispora Vijaykr., Jeewon & K.D.
Hyde; = Pseudoannulatascus Z.L. Luo, Maharachch. & K.D. Hyde) (9)*

Diaporthomycetidae families incertae sedis


Barbatosphaeriaceae H. Zhang, K.D. Hyde & Maharachch.*
Barbatosphaeria Réblová (9)
Ceratostomella Sacc. (18)
Xylomelasma Réblová (4)

Papulosaceae Winka & O.E. Erikss.


Brunneosporella V.M. Ranghoo & K.D. Hyde (1)
Fluminicola S.W. Wong, K.D. Hyde & E.B.G. Jones (4)
Papulosa Kohlm & Volkm-Kohlm (1)
Wongia Khemmuk, Geering & R.G. Shivas (3)

Rhamphoriaceae Réblová*
Rhamphoria Niessl (15)*
Rhamphoriopsis Réblová & Gardiennet (1)*
Rhodoveronaea Arzanlou, W. Gams & Crous (1)*
Xylolentia Réblová (1)*

Thyridiaceae O.E. Erikss & J.Z. Yue


Pleurocytospora Petr. (3)
Thyridium Nitschke (34)

Trichosphaeriaceae G. Winter
Brachysporium Sacc. (25)
Collematospora Jeng & Cain (1)
Coniobrevicolla Réblová (1)
Eriosphaeria Sacc. (24)
Koorchaloma Subram. (= Kananascus Nag Raj) (11)
Rizalia Syd. & P. Syd. (6)
Schweinitziella Speg. (4)
Setocampanula Sivan. & W.H. Hsieh (1)
Trichosphaeria Fuckel (20)
Unisetosphaeria Pinnoi, E.B.G. Jones, McKenzie & K.D. Hyde (1)

Woswasiaceae H. Zhang, K.D. Hyde & Maharachch.


Cyanoannulus Raja, J. Campb. & Shearer (1)
Woswasia Jaklitsch, Réblová & Voglmayr (1)
Xylochrysis Réblová (1)

Diaporthomycetidae genera incertae sedis


Aquapteridospora Jiao Yang, K.D. Hyde & Maharachch. (1)
Aquaticola W.H. Ho, C.K.M. Tsui, Hodgkiss & K.D. Hyde (5)

1206
Aquimonospora J. Yang & K.D. Hyde (1)*
Fusoidispora D. Vijaykr., Jeewon & K.D. Hyde (1)
Platytrachelon Réblová (1)
Proliferophorum G.N. Wang, H. Zhang & Senan. (1)*
Pseudoconlarium N.G. Liu, K.D. Hyde & J.K. Liu (1)
Pseudostanjehughesia J. Yang & K.D. Hyde (1)

Hypocreomycetidae O.E. Erikss. & Winka


Coronophorales Nannf. (= Melanosporales N. Zhang & M. Blackw.)
Bertiaceae Smyk
Bertia De Not. (48)
Gaillardiella Pat. (6)

Ceratostomataceae G. Winter
Arxiomyces P.F. Cannon & D. Hawksw. (3)
Dactylidispora Y. Marín, Stchigel, Guarro & Cano (3)
Echinusitheca Y. Marín, Stchigel, Dania García, Guarro, A.N. Mill. & Cano (1)
Erythrocarpon Zukal (1)
Harzia Costantin (10)
Melanospora Corda (=Gonatobotrys Corda) (69)
Microthecium Corda (=Pteridiosperma J.C. Krug & Jeng) (ca. 20)
Pseudomicrothecium Y. Marín, Stchigel, Guarro & Cano (1)
Pustulipora P.F. Cannon (1)
Rhytidospora Jeng & Cain (5)
Scopinella Lév. (9)
Setiferotheca Matsush. (1)
Syspastospora P.F. Cannon & D. Hawksw. (4)
Vittatispora P. Chaudhary, J. Campb., D. Hawksw. & K.N. Sastry (1)

Chaetosphaerellaceae Huhndorf, A.N. Mill. & F.A. Fernández


Chaetosphaerella E. Müll. & C. Booth (4)
Crassochaeta Réblová (2)
Spinulosphaeria Sivan. (2)

Coronophoraceae Höhn.
Coronophora Fuckel (2)

Nitschkiaceae (Fitzp.) Nannf.


Acanthonitschkea Speg. (10)
Biciliosporina Subram. & Sekar (1)
Botryola Bat. & J.L. Bezerra (1)
Fracchiaea Sacc. (35)
Groenhiella Jørg. Koch, E.B.G. Jones & S.T. Moss (1)
Janannfeldtia Subram. & Sekar (1)
Lasiosphaeriopsis D. Hawksw. & Sivan. (7)
Loranitschkia Lar.N. Vassiljeva (1)
Neochaetosphaerella Lar.N. Vassiljeva, S.L. Stephenson & Chernyshev (4)
Neotrotteria Sacc. (1)
Nitschkia G.H. Otth ex P. Karst. (66)
Rhagadostoma Körb. (7)
Rhagadostomella Etayo (1)
Tortulomyces Lar.N. Vassiljeva, S.L. Stephenson, Chernyshev & K.D. Hyde (1)

1207
Scortechiniaceae Huhndorf, A.N. Mill. & F.A. Fernández
Biciliospora Petr. (1)
Coronophorella Höhn. (1)
Cryptosphaerella Sacc. (20)
Euacanthe Theiss. (2)
Neofracchiaea Teng (1)
Pseudocatenomycopsis Crous & L.A. Shuttlew. (1)
Scortechinia Sacc. (9)
Scortechiniella Arx & E. Müll. (1)
Scortechiniellopsis Sivan. (1)
Tympanopsis Starbäck (1)

Coronophorales genera incertae sedis


Papulaspora Preuss (33)
Sphaerodes Clem. (9)

Falcocladiales R.H. Perera, Maharachch., Somrith., Suetrong & K.D. Hyde


Falcocladiaceae Somrith., E.B.G. Jones & K.L. Pang
Falcocladium S.F. Silveira, Alfenas, Crous & M.J. Wingf. (5)

Glomerellales Chadef. ex Réblová, W. Gams & Seifert


Australiascaceae Réblová & W. Gams
Monilochaetes Halst. ex Harter (8)

Glomerellaceae Locq. ex Seifert & W. Gams


Colletotrichum Corda (ca. 895)

Malaysiascaceae Tibpromma & K.D. Hyde


Malaysiasca Crous & M.J. Wingf. (1)

Plectosphaerellaceae W. Gams, Summerb. & Zare


Acremoniisimulans Tibpromma & K.D. Hyde (1)
Acrostalagmus Corda (13)
Brunneochlamydosporium Giraldo López & Crous (4)
Brunneomyces A. Giraldo, Gené & Guarro (3)
Chlamydosporiella Giraldo López & Crous (1)
Chordomyces Bilanenko, Georgieva & Grum-Grzhim. (2)
Furcasterigmium Giraldo López & Crous (1)
Fuscohypha Giraldo López & Crous (1)
Gibellulopsis Bat. & H. Maia (3)
Lectera P.F. Cannon (6)
Longitudinalis Tibpromma & K.D. Hyde (1)
Musicillium Zare & W Gams (2)
Musidium Giraldo López & Crous (1)
Nigrocephalum Giraldo López & Crous (1)
Paragibellulopsis Giraldo López & Crous (1)
Paramusicillium Giraldo López & Crous (1)
Phialoparvum Giraldo López & Crous (1)
Plectosphaerella Kleb. (17)
Sayamraella Giraldo López & Crous (1)
Sodiomyces A.A. Grum-Grzhim., Debets & Bilanenko (5)
Stachylidium Link (7)

1208
Summerbellia Giraldo López & Crous (1)
Theobromium Giraldo López & Crous (1)
Verticillium Nees (81)

Reticulascaceae Réblová & W. Gams


Blastophorum Matsush. (5)
Cylindrotrichum Bonord. (23)
Kylindria DiCosmo, S.M. Berch & W.B. Kendr. (11)
Sporoschismopsis Hol-Jech. & Hennebert (8)

Glomerellales genus incertae sedis


Ascocodinaea Samuels, Cand. & Magni (2)

Hypocreales Lindau
Bionectriaceae Samuels & Rossman
Acremonium Link (ca. 150)
Anthonectria Döbbeler (1)
Aphanotria Döbbeler (1)
Battarrina (Sacc.) Clem. & Shear (1)
Bryocentria Döbbeler (15)
Bryotria Döbbeler & P.G. Davison (2)
Bullanockia Crous (1)
Chrysonectria Lechat & J. Fourn. (1)*
Clibanites (P. Karst.) P. Karst. (1)
Clonostachys Corda (78)
Dimerosporiella Speg. (8)
Fusariella Sacc. (17)
Geonectria Lechat & J. Fourn. (1)*
Geosmithia J. Pitt (24)
Gliomastix Guég. (24)
Globonectria Etayo (1)
Gracilistilbella Seifert (4)
Halonectria E.B.G. Jones (1)
Heleococcum P.M. Jørg. (5)
Hydropisphaera Dumort (29)
Ijuhya Starbäck (22)
Kallichroma Kohlm. & Volkm.-Kohlm. (4)
Laniatria Döbbeler & P.G. Davison (1)
Lasionectria (Sacc.) Cooke (23)
Lasionectriella Lechat & J. Fourn. (2)*
Mycoarachis Malloch & Cain (2)
Mycocitrus Möller (3)
Nectriella Nitschke ex Fuckel (84)
Nectriopsis Maire (70)
Nigrosabulum Malloch & Cain (1)
Ochronectria Rossman & Samuels (3)
Ovicuculispora Etayo (2)
Paracylindrocarpon Crous, Roets & L. Lombard (4)
Paranectria Sacc. (4)
Periantria Döbbeler & P.G. Davison (2)
Peristomialis (W. Phillips) Boud. (6)
Pronectria Clem. (44)

1209
Protocreopsis Yoshim Doi (12)
Roumegueriella Speg. (4)
Selinia P. Karst. (6)
Stephanonectria Schroers & Samuels (1)
Stilbocrea Pat. (7)
Stromatonectria Jaklitsch & H. Voglmayr (1)
Synnemellisia N.K. Rao, Manohar. & Goos (2)
Trichonectria Kirschst. (19)
Verrucostoma Hirooka, Tak. Kobay. & P. Chaverri (2)
Xanthonectria Lechat, J. Fourn. & P.-A. Moreau (1)*

Calcarisporiaceae Jing Z. Sun, X.Z. Liu & K.D. Hyde


Calcarisporium Preuss (8)

Clavicipitaceae (Lindau) Earle ex Rogerson


Aciculosporium I. Miyake (=Neoclaviceps J.F. White, Bills, S.C. Alderman & Spatafora) (4)
Aschersonia Mont. (=Hypocrella Sacc. fide Hyde et al. 2020) (170+)
Atkinsonella Diehl (2)
Balansia Speg. (49)
Cavimalum Yoshim. Doi, Dargan & K.S. Thind (2)
Claviceps Tul. (111)
Collarina A. Giraldo, Gené & Guarro (1)
Conoideocrella D. Johnson, G.H. Sung, Hywel-Jones & Spatafora (3)
Corallocytostroma Y.N. Yu & Z.Y. Zhang (2)
Dussiella Pat. (3)
Ephelis Fr. (4)
Epichloë (Fr.) Tul. & C. Tul. (75)
Epicrea Petr. (1)
Helicocollum Luangsa-ard (3)
Helminthascus Tranzschel (1)
Heteroepichloë E. Tanaka, C. Tanaka, Gafur & Tsuda (2)
Konradia Racib. (2)
Loculistroma F. Patt & Charles (1)
Metapochonia Kepler, S.A. Rehner & Humber (6)
Metarhiziopsis D.W. Li, R.S. Cowles & C.R. Vossbrinck (1)
Metarhizium Sorokīn (= Chamaeleomyces Sigler; = Metacordyceps G.H. Sung, J.M. Sung,
Hywel-Jones & Spatafora; = Nomuraea Maubl.; = Stereocrea Syd. & P. Syd.) (78)
Moelleriella Bres. (57)
Mycomalus A. Möller (1)
Mycophilomyces Crous & M.J. Wingf. (1)
Myriogenospora G.F. Atk. (4)
Neobarya Lowen (12)
Neocordyceps Kobayasi (1)
Nigelia Luangsa-ard (2)
Nigrocornus Ryley & Langdon (1)
Orbiocrella D. Johnson, G.H. Sung, Hywel-Jones & Spatafora (1)
Parepichloë J.F. White & P.V. Reddy (4)
Periglandula U. Steiner, E. Leistner & Leuchtm. (2)
Pochonia Bat. & O.M. Fonseca (4)
Pseudomeria G.L. Barron (1)
Regiocrella Chaverri & K.T. Hodge (2)
Romanoa Thirum. (1)

1210
Rotiferophthora G.L. Barron (27)
Samuelsia Chaverri & K.T. Hodge (6)
Shimizuomyces Kobayasi (2)
Sphaerocordyceps Kobayasi (2)
Tyrannicordyceps Kepler & Spatafora (5)
Ustilaginoidea Bref. (19)

Cocoonihabitaceae W.Y. Zhuang & Z.Q. Zeng


Cocoonihabitus W.Y. Zhuang & Z.Q. Zeng (1)

Cordycipitaceae Kreisel ex G.H. Sung, J.M. Sung, Hywel-Jones & Spatafora


Akanthomyces Lebert (= Torrubiella Boud., = Lecanicillium W. Gams & Zare) (21)*
Amphichorda Fr. (1)
Ascopolyporus Möller (7)
Beauveria Vuill. (54)
Beejasamuha Subram. & Chandrash. (1)
Blackwellomyces Spatafora & Luangsa-ard (2)
Cordyceps (Fr.) Link (= Isaria Pers.; = Microhilum H.Y. Yip & A.C. Rath) (498)
Coremiopsis Sizova & Suprun (2)
Engyodontium de Hoog (5)
Gibellula Cavara (=Granulomanus de Hoog & Samson) (29)
Hevansia Luangsa-ard, Hywel-Jones & Spatafora (8)
Hyperdermium J.F. White, R.F. Sullivan, Bills & Hywel-Jones (3)
Leptobacillium Zare & W. Gams (1)
Parengyodontium C.C. Tsang, J.F.W. Chan, W.M. Pong, J.H.K. Chen, A.H.Y. Ngan, M.
Cheung, C.K.C. Lai, D.N.C. Tsang, S.K.P. Lau & P.C.Y. Woo (1)
Pseudogibellula Samson & H.C. Evans (1)
Samsoniella Mongkols., Noisrip., Thanakitp., Spatafora & Luangsa-ard (3)
Simplicillium W. Gams & Zare (12)

Flammocladiellaceae Crous, L. Lombard & R.K. Schumach.


Flammocladiella Crous, L. Lombard & R.K. Schumach. (2)

Hypocreaceae De Not.
Arachnocrea Z. Moravec. (3)
Dialhypocrea Speg. (1)
Escovopsioides H.C. Evans & J.O. Augustin (1)
Escovopsis J.J. Muchovej & Della Lucia (14)
Hypocreopsis P. Karst. (14)
Hypomyces (Fr.) Tul. & C. Tul. (ca. 150)
Kiflimonium Summerb., J.A. Scott, Guarro & Crous (1)
Lichenobarya Etayo, Diederich & Lawrey (1)
Mycogone Link (28)
Protocrea Petch (6)
Rogersonia Samuels & Lodge (1)
Sepedonium Link (13)
Sphaerostilbella (Henn.) Sacc. & D. Sacc (13)
Sporophagomyces K. Põldmaa & Samuels (3)
Stephanoma Wallr. (?6)
Trichoderma Pers. (400+)
Verticimonosporium Matsush. (3)

1211
Myrotheciomycetaceae Crous
Emericellopsis J.F.H. Beyma (23)
Leucosphaerina Arx (2)
Myrotheciomyces Crous (1)
Trichothecium Link (9)

Nectriaceae Tul. & C. Tul.


Albonectria Rossman & Samuels (1)
Allantonectria Earle (1)
Allonectella Petr. (2)
Aphanocladium W. Gams (4)
Aquanectria L. Lombard & Crous (3)
Atractium Link (3)*
Baipadisphaeria Pinruan (1)
Bisifusarium L. Lombard, Crous & W. Gams (7)
Calonectria De Not. (400)
Calostilbe Sacc. & Syd. (4)
Campylocarpon Halleen, Schroers & Crous (3)
Chaetonectrioides Matsush. (1)
Chaetopsina Rambelli (19)
Coccinonectria Lombard & Crous (2)
Corallomycetella Henn. (4)
Corallonectria C. Herrera & P. Chaverri (1)
Corinectria C. González & P. Chaverri (3)
Cosmospora Rabenh. (50)
Cosmosporella S.K. Huang, R. Jeewon & K.D. Hyde (1)
Curvicladiella Decock & Crous (1)
Cyanochyta Höhn. (1)
Cyanonectria Samuels & Chaverri (2)
Cyanophomella Höhn. (1)
Cylindrocladiella Boesew. (45)
Cylindrodendrum Bonord. (4)
Dacryoma Samuels (2)
Dactylonectria L. Lombard & Crous (14)
Dematiocladium Allegr., Aramb., Cazau & Crous (2)
Fusarium Link (ca. 120)
Fusicolla Bonord (18)
Geejayessia Schroers, Gräfenhan & Seifert (7)
Gliocephalotrichum J.J. Ellis & Hesselt. (13)
Gliocladiopsis S.B. Saksena (15)
Ilyonectria P. Chaverri & C. Salgado (23)
Macroconia (Wollenw.) Gräfenhan, Seifert & Schroers (5)
Mariannaea G. Arnaud ex Samson (22)
Microcera Desm. (4)
Murinectria M. Niranjan & V.V. Sarma (4)
Nalanthamala Subram. (6)
Nectria (Fr.) Fr. (29)
Neocosmospora E.F. Sm. (84)
Neonectria Wollenw. (30)
Neothyronectria Crous & Thangavel (2)
Ophionectria Sacc. (39)
Pandanaceomyces Tibpromma & K.D. Hyde (1)

1212
Paracremonium L. Lombard & Crous (5)
Payosphaeria W.F. Leong (1)
Penicillifer Emden (7)
Persiciospora P.F. Cannon & D. Hawksw. (4)
Pleiocarpon L. Lombard & D. Aiello (3)
Pleogibberella Sacc. (3)
Pleurocolla Petr. (1)
Pseudoachroiostachys Tibpromma & K.D. Hyde (1)
Pseudocosmospora C. Herrera & P. Chaverri (13)
Pseudonectria Seaver (17)
Rectifusarium L. Lombard, Crous & W. Gams (2)
Rugonectria P. Chaverri & Samuels (5)
Sarcopodium Ehrenb. (22)
Stylonectria Höhn. (5)
Thelonectria P. Chaverri & C.G. Salgado (46)
Thyronectria Sacc. (41)
Varicosporella Lechat & J. Fourn. (1)
Varicosporellopsis Lechat & J. Fourn. (1)*
Volutella Fr. (127)
Xenoacremonium Lombard & Crous (2)
Xenocylindrocladium Decock, Hennebert & Crous (3)
Xenogliocladiopsis Crous & W.B. Kendr. (2)
Xenoleptographium Marinc., T.A. Duong, Z.W. de Beer & M.J. Wingf. (1)
Xenonectriella Weese (18)

Niessliaceae Kirschst.
Atronectria Etayo (1)
Circinoniesslia Samuels & M.E. Barr (1)
Cryptoniesslia Scheuer (1)
Eucasphaeria Crous (2)
Hyaloseta A.W. Ramaley (1)
Malmeomyces Starb. (1)
Melchioria Penz. & Sacc. (6)
Miyakeomyces Hara (1)
Myrmaeciella Lindau (2)
Myrtacremonium Crous (1)
Neoeucasphaeria Crous (1)
Niesslia Auersw. (43)
Paraniesslia K.M. Tsui, K.D. Hyde & Hodgkiss (2)
Pseudohyaloseta Tibpromma & K.D. Hyde (1)
Pseudonectriella Petr. (1)
Pseudorhynchia Höhn. (2)
Rosasphaeria Jaklitsch & Voglmayr (1)
Taiwanascus Sivan & H.S. Chang (2)
Trichosphaerella E. Bommer, M. Rousseau & Sacc. (=Neorehmia Höhn.; = Oplothecium
Syd.) (4)
Valetoniella Höhn. (3)
Valetoniellopsis Samuels & M.E. Barr (1)

Ophiocordycipitaceae G.H. Sung, J.M. Sung, Hywel-Jones & Spatafora


Drechmeria W. Gams & H.B. Jansson (12)
Harposporium Lohde (37)

1213
Hirsutella Pat. (50+)
Hymenostilbe Petch (12)
Ophiocordyceps Petch (263)
Paraisaria Samson & B.L. Brady (11)
Perennicordyceps Matočec & I. Kušan (4)*
Polycephalomyces Kobayasi (18)*
Purpureocillium Luangsa-ard, Hywel-Jones, Houbraken & Samson (5)
Tolypocladium W. Gams (47)

Sarocladiaceae L. Lombard
Parasarocladium Summerb., J.A. Scott, Guarro & Crous (4)
Sarocladium W. Gams & D. Hawksw. (22)

Stachybotryaceae L. Lombard & Crous


Achroiostachys L. Lombard & Crous (6)
Albifimbria L. Lombard & Crous (5)
Albosynnema E.F. Morris (2)
Alfaria Crous, Montaño-Mata & García-Jim. (13)
Alfariacladiella Crous & R.K. Schumach. (1)
Brevistachys L. Lombard & Crous (5)
Capitofimbria L. Lombard & Crous (1)
Cymostachys L. Lombard & Crous (3)
Didymostilbe Henn. (14)
Digitiseta Gordillo & Decock (4)*
Dimorphiseta L. Lombard & Crous (1)
Globobotrys L. Lombard & Crous (1)
Grandibotrys L. Lombard & Crous (3)
Gregatothecium L. Lombard & Crous (1)
Hyalinostachys C.G. Lin & K.D. Hyde (1)
Inaequalispora L. Lombard & Crous (3)
Kastanostachys L. Lombard & Crous (1)
Koorchalomella Chona, Munjal & J.N. Kapoor (2)
Melanopsamma Niessl (ca. 5)
Memnoniella Höhn. (9)
Myrothecium Tode (2)
Myxospora L. Lombard & Crous (6)
Neomyrothecium L. Lombard & Crous (1)
Paramyrothecium L. Lombard & Crous (14)
Parasarcopodium Melnik, S.J. Lee & Crous (3)
Parvothecium L. Lombard & Crous (2)
Peethambara Subram. & Bhat (1)
Pseudoornatispora Tibpromma & K.D. Hyde (1)
Septomyrothecium Matsush. (4)
Sirastachys L. Lombard & Crous (9)
Smaragdiniseta L. Lombard & Crous (1)
Stachybotrys Corda (12 phylogenetically studied, 81 epithets remain be studied)
Striatibotrys L. Lombard & Crous (7)
Striaticonidium L. Lombard & Crous (5)
Tangerinosporium L. Lombard & Crous (1)
Virgatospora Finley (2)
Xenomyrothecium L. Lombard & Crous (1)
Xepicula Nag Raj (4)

1214
Xepiculopsis Nag Raj (2)

Tilachlidiaceae Lombard & Crous


Psychronectria J. Pawłowska, Istel, Wrzosek, D. Hawksw. (47)
Septofusidium W. Gams (5)
Tilachlidium Preuss (1)

Hypocreales genera incertae sedis


Acremoniopsis A. Giraldo, Gené & Guarro (1)
Berkelella (Sacc.) Sacc. (2)
Bulbithecium Udagawa & T Muroi (1)
Cephalosporiopsis Peyronel (10)
Chondronectria Etayo, Flakus & Kukwa (1)
Cylindronectria Etayo (1)
Diploöspora Grove (ca. 7)
Gynonectria Döbbeler (1)
Hapsidospora Malloch & Cain (2)
Haptospora G.L. Barron (3)
Illosporiopsis D. Hawksw. (1)
Illosporium Mart. (17)
Leptobarya Etayo (2)
Lichenopenicillus Etayo (1)
Metadothella Henn. (1)
Munkia Speg. (4)
Neomunkia Petr. (1)
Peloronectria Möller (3)
Pseudoacremonium Crous (1)
Pseudoidriella Crous & R.G. Shivas (1)
Pseudomeliola Speg. (10)
Rodentomyces Doveri, Pecchia, Sarrocco & Vannacci (1)
Roselliniella Vain (19)
Saksenamyces A.N. Rai & P.N. Singh (1)
Sedecimiella K.L. Pang, Alias & E.B.G. Jones (1)
Stanjemonium W. Gams, O'Donnell, Schroers & M. Chr. (4)
Stilbella Lindau (61)
Ticonectria Döbbeler (3)
Tilakidium Vaidya, C.D. Naik & Rathod (1)

Jobellisiales M.J. D’souza & K.D. Hyde


Jobellisiaceae Réblová
Jobellisia M.E. Barr (8)

Microascales Luttr. ex Benny & Kimbr.


Ceratocystidaceae Locq. ex Réblová, W. Gams & Seifert
Ambrosiella Brader ex Arx & Hennebert (10)
Berkeleyomyces W.J. Nel, Z.W. de Beer, T.A. Duong & M.J. Wingf. (2)
Bretziella Z.W. de Beer, Marinc., T.A. Duong & M.J. Wingf. (1)
Ceratocystis Ellis & Halst. (105)
Chalaropsis Peyronel (3)
Davidsoniella Z.W. de Beer, T.A. Duong & M.J. Wingf. (4)
Endoconidiophora Münch (9)
Huntiella Z.W. de Beer, T.A. Duong & M.J. Wingf. (29)

1215
Meredithiella McNew, C. Mayers & T.C. Harr. (3)
Phialophoropsis L.R. Batra emend. T.C. Harr. (2)
Thielaviopsis Went. (7)

Chadefaudiellaceae Faurel & Schotter ex Benny & Kimbr.


Chadefaudiella Faurel & Schotter (2)
Faurelina Locq-Lin. (4)

Gondwanamycetaceae Réblová, W. Gams & Seifert


Custingophora Stolk (1)
Knoxdaviesia M.J. Wingf., P.S. van Wyk & Marasas. (5)

Graphiaceae De Beer
Graphium Corda (20)

Halosphaeriaceae E. Müll & Arx ex Kohlm.


Alisea J. Dupont & E.B.G. Jones (1)
Amphitrite S. Tibell (1)
Aniptodera Shearer & M. Miller (21)
Aniptosporopsis (K.D. Hyde) K.L. Pang (1)
Anisostagma K.R.L. Petersen & Jørg. Koch (1)
Antennospora Meyers (2)
Appendichordella R.G. Johnson, E.B.G. Jones & S.T. Moss (1)
Arenariomyces Höhnk (5)
Ascosacculus J. Campbell, J.L. Anderson & Shearer (1)
Bathyascus Kohlm. (5)
Carbosphaerella I. Schmidt (2)
Ceriosporopsis Linder (9)
Chadefaudia Feldm.-Maz. (6)
Corallicola Volkm.-Kohlm. & Kohlm. (1)
Corollospora Werderm (= Cirrenalia Meyers & R.T. Moore; = Sigmoidea J.L. Crane) (25)
Cucullosporella K.D. Hyde & E.B.G. Jones (1)
Ebullia K.L. Pang (1)
Fluviatispora K.D. Hyde (3)
Gesasha Abdel-Wahab & Nagah. (3)
Haiyanga K.L. Pang & E.B.G. Jones (1)
Haligena Kohlm. (1)
Halosarpheia Kohlm. & E. Kohlm. (8)
Halosphaeria Linder (1)
Halosphaeriopsis T.W. Johnson (1)
Havispora K.L. Pang & Vrijmoed (1)
Iwilsoniella E.B.G.Jones (1)
Kitesporella Jheng & K.L. Pang (1)
Kochiella Sakay., K.L. Pang & E.B.G. Jones (1)
Lautisporopsis E.B.G. Jones, Yusoff & S.T. Moss (1)
Lignincola Höhnk (2)
Limacospora Jørg. Koch & E.B.G. Jones (1)
Luttrellia Shearer (4)
Magnisphaera J. Campb., J.L. Anderson & Shearer (2)
Marinospora A.R. Caval. (2)
Moana Kohlm. & Volkm.-Kohlm. (1)
Morakotiella Sakay. (1)

1216
Naïs Kohlm. (3)
Natantispora J. Campb., J.L. Anderson & Shearer (3)
Nautosphaeria E.B.G. Jones (1)
Neptunella K.L. Pang & E.B.G. Jones (1)
Nereiospora E.B.G. Jones, R.G. Johnson & S.T. Moss. (2)
Nimbospora Jørg. Koch (1)
Nohea Kohlm. & Volkm.-Kohlm. (3)
Oceanitis Kohlm. (4)
Ocostaspora E.B.G. Jones, R.G. Johnson & S.T. Moss (1)
Okeanomyces K.L. Pang & E.B.G. Jones (1)
Ondiniella E.B.G. Jones, R.G. Johnson & S.T. Moss (1)
Ophiodeira Kohlm. & Volkm.-Kohlm. (1)
Paraaniptodera K.L. Pang, C.L. Lu, W.T. Ju & E.B.G. Jones (1)
Phaeonectriella R.A. Eaton & E.B.G. Jones (1)
Praelongicaulis E.B.G. Jones, Abdel-Wahab & K.L. Pang (1)
Panorbis J. Campb., J.L. Anderson & Shearer (1)
Pileomyces K.L. Pang & Jheng (1)
Pseudolignincola Chatmala & E.B.G. Jones (1)
Remispora Linder (5)
Saagaromyces K.L. Pang & E.B.G. Jones (3)
Sablicola E B.G. Jones, K.L. Pang & Vrijmoed (1)
Thalassogena Kohlm. & Volkm.-Kohlm. (1)
Thalespora Chatmala & E.B.G. Jones (1)
Tinhaudeus K.L. Pang, S.Y. Guo & E.B.G. Jones (1)
Tirispora E.B.G. Jones & Vrijmoed (1)
Toriella Sakay., K.L. Pang & E.B.G. Jones (1)
Trailia G.K. Sutherl. (1)
Trichomaris Hibbits, G.C. Hughes & Sparks (1)
Tubakiella Sakay., K.L. Pang & E.B.G. Jones (1)
Tunicatispora K.D. Hyde (1)

Microascaceae Luttr. ex Malloch


Acaulium Sopp (4)
Brachyconidiellopsis Decock, R.F. Castañeda & Adhikari (1)
Canariomyces Arx (3)
Cephalotrichum Link (37)
Doratomyces Corda (3)
Echinobotryum Corda (2)
Enterocarpus Locq.-Lin. (2)
Fairmania Sacc. (1)
Gamsia M. Morelet (5)
Kernia Nieuwl. (14)
Lomentospora Hennebert & B.G. Desai (1)
Lophotrichus R.K. Benj. (8)
Microascus Zukal (60)
Parascedosporium Gilgado, Gené, Cano & Guarro (2)
Petriella Curzi (8)
Pseudallescheria Negroni & I. Fisch. (8)
Pseudoscopulariopsis Sand.-Den., Gené & Guarro (2)
Rhinocladium Sacc. & Marchal (11)
Scedosporium Sacc. ex Castell. & Chalm. (12)
Scopulariopsis Bainier (87)

1217
Wardomyces F.T. Brooks & Hansf. (11)
Wardomycopsis Udagawa & Furuya (5)
Yunnania H.Z. Kong (3)

Triadelphiaceae Y.Z. Lu, J.K. Liu, Z.L. Luo & K.D. Hyde
Synnematotriadelphia Chuaseehar., Somrith., Nuankaew & Boonyuen (2)
Triadelphia Shearer & J.L. Crane (18)*

Microascales genera incertae sedis


Bisporostilbella Brandsb. & E.F. Morris (1)
Cephalotrichiella Crous (1)
Cornuvesica C.D. Viljoen, M.J. Wingf. & K. Jacobs (4)
Gabarnaudia Samson & W. Gams (2)
Sporendocladia G. Arnaud ex Nag Raj & W.B. Kendr. (7)

Pararamichloridiales Crous
Pararamichloridiaceae Crous
Pararamichloridium Crous (2)

Torpedosporales E.B.G. Jones, Abdel-Wahab & K.L. Pang


Etheirophoraceae Rungjind., Somrith. & Suetrong
Etheirophora Kohlm. & Volkm.-Kohlm. (3)
Swampomyces Kohlm. & Volkm. (2)

Juncigenaceae E.B.G. Jones, Abdel-Wahab & K.L. Pang


Elbamycella A. Poli, E. Bovio, V. Prigione & G.C. Varese (1)
Fulvocentrum E.B.G. Jones & Abdel-Wahab (3)
Juncigena Kohlm Kohlm., Volkm.-Kohlm. & O.E. Erikss. (2)
Khaleijomyces Abdel-Wahab (1)*
Marinokulati E.B.G. Jones & K.L. Pang (1)
Moheitospora Abdel-Wahab, Abdel-Aziz & Nagah. (2)

Torpedosporaceae E.B.G. Jones & K.L. Pang


Torpedospora Meyers (3)

Hypocreomycetidae genera incertae sedis


Campylospora Ranzoni (5)
Dendroclathra Voglmayr & G. Delgado (2)

Lulworthiomycetidae Dayar., E.B.G. Jones & K.D. Hyde


Koralionastetales Kohlm., Volkm.-Kohlm., J. Campb. & Inderb.
Koralionastetaceae Kohlm. & Volkm.-Kohlm.
Koralionastes Kohlm. & Volkm.-Kohlm. (5)
Pontogeneia Kohlm. (8)

Lulworthiales Kohlm., Spatafora & Volkm.-Kohlm.


Lulworthiaceae Kohlm., Spatafora & Volkm.-Kohlm.
Cumulospora I. Schmidt (2)
Halazoon Abdel-Aziz, Abdel-Wahab & Nagah. (2)
Haloguignardia A. Cribb & J. Cribb (1)
Hydea K.L. Pang & E.B.G Jones (1)
Kohlmeyeriella E.B.G. Jones, R.G. Johnson & S.T. Moss (2)

1218
Lindra I. Wilson (2)
Lulwoana Kohlm., Volkm.-Kohlm., J. Campb., Spatafora & Gräfenhan (1)
Lulwoidea Kohlm., Volkm.-Kohlm., J. Campb., Spatafora & Gräfenhan (1)
Lulworthia G.K. Sutherl (32)
Matsusporium E.B.G. Jones & K.L. Pang (1)
Moleospora Abdel-Wahab, Abdel-Aziz & Nagah (1)
Moromyces Abdel-Wahab, K.L. Pang, Nagah., Abdel-Aziz & E.B.G. Jones (1)
Orbimyces Linder (1)
Rostrupiella Jørg Koch, K.L. Pang & E.B.G. Jones. (1)
Sammeyersia S.Y. Guo, E.B.G. Jones & K.L. Pang (1)

Pisorisporiomycetidae Bundhun, Maharachch. & K.D. Hyde


Pisorisporiales Réblová & J. Fourn.
Pisorisporiaceae Réblová & J. Fourn.
Achroceratosphaeria Réblová, Fourn., K.D. Hyde & Ranghoo (2)
Pisorisporium Réblová & J. Fourn. (2)

Savoryellomycetidae Hongsanan, K.D. Hyde & Maharachch.


Conioscyphales Réblová & Seifert
Conioscyphaceae Réblová & Seifert
Conioscypha Höhn. (16)

Fuscosporellales Jing Yang, Bhat & K.D. Hyde


Fuscosporellaceae Jing Yang, Bhat & K.D. Hyde
Bactrodesmiastrum Hol.-Jech. (5)
Fuscosporella Jing Yang (2)
Mucispora Jing Yang (2)
Parafuscosporella Jing Yang & K.D. Hyde (3)
Plagiascoma Réblová & J. Fourn. (1)
Pseudoascotaiwania Jing Yang, Bhat & K.D. Hyde (1)

Pleurotheciales Réblová & Seifert


Pleurotheciaceae Réblová & Seifert
Adelosphaeria Réblová (1)
Anapleurothecium Hern.-Restr., R.F. Castañeda & Gené (1)
Helicoön Morgan (28)
Melanotrigonum Réblová (1)
Monotosporella S. Hughes (4)
Neomonodictys Y.Z. Lu, C.G. Lin & K.D. Hyde (1)
Phaeoisaria Höhn. (23)
Phragmocephala E.W. Mason & S. Hughes (15)
Pleurotheciella Réblová (11)
Pleurothecium Höhn. (11)
Sterigmatobotrys Oudem. (6)

Savoryellales Boonyuen, Suetrong, Sivichai, K.L. Pang & E.B.G. Jones


Savoryellaceae Jaklitsch & Réblová
Ascotaiwania Sivan. & H.S. Chang (= Neoascotaiwania Hern.-Restr., R.F. Castañeda &
Guarro fide Dayarathne et al. 2019) (9)
Canalisporium Nawawi & Kuthub. (= Ascothailandia Sri-indr., Boonyuen, Sivichai &
E.B.G. Jones) (15)
Rhexoacrodictys W.A. Baker & Morgan-Jones (5)*

1219
Savoryella E.B.G. Jones & R.A. Eaton (11)

Sordariomycetidae O.E. Erikss & Winka (= Meliolomycetidae P.M. Kirk & K.D. Hyde)*
Boliniales P.F. Cannon
Boliniaceae Rick
Apiocamarops Samuels & J.D. Rogers (4)
Apiorhynchostoma Petr. (4)
Camaropella Lar.N. Vassiljeva (2)
Camarops P. Karst. (= Bolinia (Nitschke) Sacc.) (28)
Cornipulvina Huhndorf, A.N. Mill., F.A. Fernández & Lodge (1)
Endoxyla Fuckel (3)
Mollicamarops Lar.N. Vassiljeva (1)
Neohypodiscus J.D. Rogers, Y.M. Ju & Læssøe (3)
Pseudovalsaria Spooner (3)

Cephalothecales Maharachch. & K.D. Hyde


Cephalothecaceae Höhn.
Albertiniella Kirschst. (2)
Cephalotheca Fuckel (ca. 10)
Cryptendoxyla Malloch & Cain (2)
Phialemonium W. Gams & McGinnis (6)
Victoriomyces D. Davolos, B. Pietrangeli, A.M. Persiani & O. Maggi (1)

Chaetosphaeriales Huhndorf, A.N. Mill. & F.A. Fernández


Chaetosphaeriaceae Réblová, M.E. Barr & Samuels
Adautomilanezia Gusmão, S.S. Silva, Fiuza, L.A. Costa & T.A.B. Santos (1)
Anacacumisporium Y.R. Ma & X.G. Zhang (1)
Ascochalara Réblová (1)
Bahusutrabeeja Subram. & Bhat (6)
Brunneodinemasporium Crous & R.F. Castañeda (2)
Catenularia Grove (13)
Chaetosphaeria Tul. & C. Tul. (ca. 150)
Chloridium Link (= Melanopsammella Höhn.) (ca. 30)
Codinaea Maire (15)
Conicomyces R.C. Sinclair, Eicker & Morgan-Jones (4)
Craspedodidymum Hol.-Jech. (14)
Cryptophiale Piroz. (ca. 20)
Cryptophialoidea Kuthub. & Nawawi (5)
Dendrophoma Sacc. (ca. 100)
Dictyochaeta Speg. (84)
Dictyochaetopsis Aramb. & Cabello (14)
Dinemasporium Lév. (35)
Eucalyptostroma Crous & M.J. Wingf. (2)
Exserticlava S. Hughes (7)
Hemicorynespora M.B. Ellis (12)
Infundibulomyces Plaingam, Somrith. & E.B.G. Jones (2)
Kionochaeta P.M. Kirk & B. Sutton (13)
Lecythothecium Réblová & Winka (1)
Menispora Pers. (14)
Menisporopsis S. Hughes (ca. 10)
Miyoshiella Kawam. (3)
Morrisiella Saikia & A.K. Sarbhoy (1)

1220
Nawawia Marvanová (7)
Neopseudolachnella A. Hashim. & Kaz. Tanaka (3)
Paliphora Sivan. & B. Sutton (7)
Phialosporostilbe Mercado & J. Mena (5)
Polynema Lév. (13)
Pseudodinemasporium A. Hashim. & Kaz. Tanaka (1)
Pseudolachnea Ranoj. (5)
Pseudolachnella Teng (18)
Pyrigemmula D. Magyar & R. Shoemaker (1)
Rattania Prabhug. & Bhat (1)
Sporoschisma Berk. & Broome (15)
Striatosphaeria Samuels & E. Müll. (1)
Tainosphaeria F.A. Fernández & Huhndorf (3)
Thozetella Kuntze (22)
Umbrinosphaeria Réblová (1)
Verhulstia Hern.-Rest. (1)
Zanclospora S. Hughes & W.B. Kendr. (10)

Helminthosphaeriaceae Samuels, Cand. & Magni.


Echinosphaeria A.N. Mill. & Huhndorf (14)
Endophragmiella B. Sutton (ca. 80)
Helminthosphaeria Fuckel (ca. 20)
Hilberina Huhndorf & A.N. Mill. (ca. 20)
Ruzenia O. Hilber (1)
Synaptospora Cain (5)
Tengiomyces Réblová (1)

Leptosporellaceae Konta & K.D. Hyde


Leptosporella Penz. & Sacc. (17)

Linocarpaceae Konta & K.D. Hyde


Linocarpon Syd. & P. Syd. (42)
Neolinocarpon K.D. Hyde (13)

Chaetosphaeriales genera incertae sedis


Calvolachnella Marinc., T.A. Duong & M.J. Wingf. (1)
Caudatispora J. Fröhl. & K.D. Hyde (2)
Erythromada Huhndorf, A.N. Mill., F.A. Fernández & Lodge (1)
Lasiosphaeriella Sivan. (6)
Neoleptosporella Phukhams., Perera & K.D. Hyde (2)
Neonawawia Jing Yang, K.D. Hyde & J.K. Liu (1)
Rimaconus Huhndorf, F.A. Fernández, Joanne E. Taylor & K.D. Hyde (2)

Coniochaetales Huhndorf, A.N. Mill. & F.A. Fernández (= Cordanales M. Hern.-Rest. & Crous)
Coniochaetaceae Malloch & Cain
Barrina A.W. Ramaley (1)
Coniochaeta (Sacc.) Cooke (82)

Cordanaceae Nann.
Cordana Preuss (19)

Coniochaetales genera incertae sedis

1221
Cannonia J.E. Taylor & K.D. Hyde (1)
Pseudogliomastix W. Gams (1)

Meliolales Gäum. ex D. Hawksw. & O.E. Erikss.


Armatellaceae Hosag.
Armatella Theiss. & Syd. (19)

Meliolaceae G.W. Martin ex Hansf.


Amazonia Theiss. (60)
Appendiculella Höhn. (70)
Asteridiella McAlpine (2)
Cryptomeliola S. Hughes & Piroz. (3)
Endomeliola S. Hughes & Piroz. (1)
Irenopsis F. Stevens (150)
Meliola Fr. (1700)
Setameliola D.R. Reynolds (17)

Phyllachorales M.E. Barr


Phaeochoraceae K.D. Hyde, P.F. Cannon & M.E. Barr
Cocoicola K.D. Hyde (5)
Phaeochora Höhn. (4)
Phaeochoropsis K.D. Hyde & P.F. Cannon (4)
Serenomyces Petr. (4)

Phaeochorellaceae Guterres, Galvão-Elias & Dianese


Phaeochorella Theiss. & Syd. (6)

Phyllachoraceae Theiss. & H. Syd.


Ascovaginospora Fallah, Shearer & W.D. Chen (1)
Brobdingnagia K.D. Hyde & P.F. Cannon (4)
Camarotella Theiss. & Syd. (8)
Coccodiella Hara (27)
Cyclodomus Höhn. (5)
Deshpandiella Kamat & Ullasa (1)
Diachora Müll. Arg. (4)
Diatractium Syd. & P. Syd. (4)
Erikssonia Penz. & Sacc. (5)
Fremitomyces P.F. Cannon & H.C. Evans (2)
Geminispora Pat. (2)
Gibellina Pass. Ex Roum. (2)
Imazekia Tak. Kobay. & Y. Kawabe (1)
Isothea Fr. (4)
Lichenochora Hafellner (44)
Lindauella Rehm (1)
Linochora Höhn. (37)
Lohwagia Petr. (3)
Maculatifrondes K.D. Hyde (1)
Malthomyces K.D. Hyde & P.F. Cannon (2)
Muelleromyces Kamat & Anahosur (1)
Neoflageoletia J. Reid & C. Booth (1)
Neophyllachora Dayar. & K.D. Hyde (4)
Ophiodothis Sacc. (6)

1222
Ophiodothella (Henn.). Höhn. (31)
Orphnodactylis Malloch & Mallik (2)
Oxodeora K.D. Hyde & P.F. Cannon (1)
Parberya C.A. Pearce & K.D. Hyde (2)
Petrakiella Syd. (1)
Phycomelaina Kohlm. (1)
Phyllachora Nitschke ex Fuckel (1513)
Phylleutypa Petr. (3)
Phyllocrea Höhn. (3)
Pseudothiella Petr. (1)
Pseudothiopsella Petr. (1)
Pterosporidium W.H. Ho & K.D. Hyde (2)
Rehmiodothis Theiss. & Syd. (10)
Retroa P.F. Cannon (2)
Rhodosticta Woron. (2)
Rikatlia P.F. Cannon (1)
Schizochora Syd. & P. Syd. (3)
Sphaerodothella C.A. Pearce & K.D. Hyde (1)
Sphaerodothis (Sacc. & P. Syd.) Shear (43)
Stigmatula (Sacc.) Syd. & P. Syd. (10)
Stigmochora Theiss. & Syd. (12)
Stromaster Höhn. (1)
Tamsiniella S.W. Wong, K.D. Hyde, W.H. Ho & S.J. Stanley (1)
Telimenella Petr. (3)
Telimenochora Sivan. (1)
Trabutia Sacc. & Roum. (1)
Tribulatia J.E. Taylor, Hyde & E.B.G. Jones (1)
Uropolystigma Maubl. (1)
Vitreostroma P.F. Cannon (3)
Zimmermanniella Henn. (1)

Telimenaceae Mardones, T. Trampe & M. Piepenbr


Telimena Racib. (14)

Phyllachorales genus incertae sedis


Marinosphaera K.D. Hyde (1)

Pseudodactylariales Crous
Pseudodactylariaceae Crous
Pseudodactylaria Crous (3)

Sordariales Chad. ex D. Hawksw. & O.E. Erikss.


Chaetomiaceae G. Winter
Achaetomium J.N. Rai, Tewari & Mukerji (16)
Acrophialophora Edward (17)
Allobotryotrichum M. Raza & L. Cai (1)
Amesia X. Wei Wang, Samson & Crous (4)
Arcopilus X. Wei Wang, Samson & Crous (5)
Arxotrichum A. Nováková & M. Kolařik (2)
Botryotrichum Sacc. & Marchal (11)
Brachychaeta X. Wei Wang & Houbraken (1)
Carteria X. Wei Wang & Houbraken (1)

1223
Chaetomium Kunze (359)
Chrysanthotrichum X. Wei Wang & Houbraken (4)
Chrysocorona X. Wei Wang & Houbraken (1)
Collariella X. Wei Wang, Samson & Crous (9)
Condenascus X. Wei Wang & Houbraken (1)
Corynascella Arx & Hodges (1)
Crassicarpon Y. Marín, Stchigel, Guarro & Cano (3)
Dichotomopilus X. Wei Wang, Samson & Crous (12)
Floropilus X. Wei Wang & Houbraken (1)
Guanomyces M.C. Gonzáles, Hanlin & Ulloa (1)
Humicola Traaen (86)
Hyalosphaerella X. Wei Wang & Houbraken (1)
Madurella Brumpt (15)
Melanocarpus Arx (5)
Microthielavia X. Wei Wang & Houbraken (1)
Myceliophthora Costantin (4)
Ovatospora X. Wei Wang, Samson & Crous (6)
Parathielavia X. Wei Wang & Houbraken (3)
Pseudothielavia X. Wei Wang & Houbraken (4)
Remersonia Samson & Seifert (2)*
Retroconis de Hoog & Bat. Vegte (1)*
Staphylotrichum J.A. Mey. & Nicot (8)
Stolonocarpus X. Wei Wang & Houbraken (1)
Subramaniula Arx (9)
Thermothelomyces Y. Marín, Stchigel, Guarro & Cano (4)
Thermothielavioides X. Wei Wang & Houbraken (1)
Thielavia Zopf (47)
Trichocladium Harz (44)

Lasiosphaeriaceae Nannf.
Anopodium Lundq. (2)
Apiosordaria Arx & W. Gams (31)
Apodospora Cain & J.H. Mirza (6)
Apodus Malloch & Cain (2)
Arnium Nitschke ex G. Winter (34)
Bellojisia Réblová (1)
Biconiosporella Schaumann (1)
Bombardia (Fr.) P. Karst. (43)
Bombardioidea C. Moreau ex N. Lundqv. (5)
Camptosphaeria Fuckel (4)
Cercophora Fuckel (77)
Corylomyces Stchigel, M. Calduch & Guarro (1)
Diffractella Guarro, P.F. Cannon & Aa (1)
Diplogelasinospora Cain (4)
Emblemospora Jeng & J.C. Krug (2)
Eosphaeria Höhn. (2)
Episternus Górz & Boroń (1)
Fimetariella N. Lundq. (9)
Immersiella A.N. Mill. & Huhndorf (2)
Jugulospora N. Lundq. (1)
Lasiosphaeria Ces. & De Not. (229)
Mammaria Ces. ex Rabenh. (2)

1224
Periamphispora J.C. Krug (1)
Ramophialophora M. Calduch, Stchigel, Gené & Guarro (4)*
Rinaldiella Deanna A. Sutton, Y. Marín, Guarro & E.H. Thomps (1)
Schizothecium Corda (31)
Strattonia Cif. (11)
Thaxteria Sacc. (8)
Tripterosporella Subram. & Lodha (5)
Zopfiella G. Winter (22)
Zygopleurage Boedijn (3)
Zygospermella Cain (3)

Podosporaceae X. Wei Wang & Houbraken


Cladorrhinum Sacc. & Marchal (13)*
Triangularia Boedijn (7)
Podospora Ces. (92)

Sordariaceae G. Winter
Copromyces N. Lundq. (1)
Effetia Bartoli, Maggi & Persiani (1)
Guilliermondia Boud. (1)
Neurospora Shear & B.O. Dodge (= Gelasinospora Dowding) (60)
Pseudoneurospora Dania García, Stchigel & Guarro (2)
Sordaria Ces. & De Not. (37)
Stellatospora T. Ito & A. Nakagiri (1)

Sordariales genera incertae sedis


Abyssomyces Kohlm (1)
Acanthotheciella Höhn. (3)
Ascolacicola Ranghoo & K.D. Hyde (1)
Bombardiella Höhn. (1)
Coronatomyces Dania García, Stchigel & Guarro (1)
Cuspidatispora Shearer & Bartolata (1)
Globosphaeria D. Hawksw. (1)
Isia D. Hawksw & Manohar (2)
Lasiosphaeris Clem. (3)
Lunulospora Ingold (2)
Lockerbia K.D. Hyde (2)
Nitschkiopsis Nannf. & R. Sant. (1)
Onygenopsis Henn. (1)
Phaeosporis Clem. (2)
Reconditella Matzer & Hafellner (1)
Rhexodenticula W.A. Baker & Morgan-Jones (4)
Rhexosporium Udagawa & Furuya (1)
Roselliniomyces Matzer & Hafellner (7)
Roselliniopsis Matzer & Hafellner (7)
Stromatographium Höhn. (= Fluviostroma Samuels & E. Müll.) (2)
Utriascus Réblová (1)
Ypsilonia Lév. (3)

Sordariomycetidae family incertae sedis


Batistiaceae Samuels & K.F. Rodrigues
Batistia Cif. (1)

1225
Sordariomycetidae genera incertae sedis
Arecacicola Joanne E. Taylor, J. Fröhl. & K.D. Hyde (1)
Bullimyces A. Ferrer, A.N. Mill., Sarmiento & Shearer (3)
Cancellidium Tubaki (2)
Ceratolenta Réblová (1)
Chaetosphaerides Matsush. (1)
Cryptophyllachora L. Kiss, Kovács & R.G. Shivas (2)*
Hanliniomyces Raja & Shearer (1)
Hydromelitis A. Ferrer, A.N. Mill., Sarmiento & Shearer (1)
Merugia Rogerson & Samuels (1)
Mycomedusiospora G.C. Carroll & Munk (1)
Myxocephala G. Weber, Spaaij & Oberw. (1)
Nigromammilla K.D. Hyde & J. Fröhl. (1)
Phaeotrichosphaeria Sivan. (4)
Phragmodiscus Hansf. (2)
Pseudobotrytis Krzemien. & Badura (2)

Xylariomycetidae O.E. Erikss & Winka


Amphisphaeriales D. Hawksw. & O.E. Erikss.
Amphisphaeriaceae G. Winter
Amphisphaeria Ces. & De Not. (66)
Griphosphaerioma Höhn. (2)
Lepteutypa Petr. (14)

Apiosporaceae K.D. Hyde, J. Fröhl., Joanne E. Taylor & M.E. Barr


Appendicospora K.D. Hyde (2)
Arthrinium Kunze (74)
Dictyoarthrinium S. Hughes (6)
Endocalyx Berk. & Broome (8)
Nigrospora Zimm. (ca. 20)

Beltraniaceae Nann.
Beltrania Penz. (17)
Beltraniella Subram. (25)
Beltraniopsis Bat. & J.L. Bezerra (11)
Hemibeltrania Piroz. (13)
Parapleurotheciopsis P.M. Kirk (5)
Porobeltraniella Gusmão (2)
Pseudobeltrania Henn. (9)
Subramaniomyces Varghese & V.G. Rao (3)
Subsessila C.G. Lin & K.D. Hyde (1)

Clypeophysalosporaceae Giraldo & Crous


Bagadiella Cheew. & Crous (4)
Clypeophysalospora H.J. Swart (1)
Neophysalospora Crous & M.J. Wingf. (1)
Plectosphaera Theiss. (28)

Cylindriaceae Crous & L. Lombard*


Cylindrium Bonord (6)

Hansfordiaceae Crous

1226
Hansfordia S. Hughes (7)

Hyponectriaceae Petr.
Apiothyrium Petr. (2)
Arecomyces K.D. Hyde (10)
Arwidssonia B. Erikss. (2)
Cesatiella Sacc. (3)
Chamaeascus L. Holm, K. Holm & M.E. Barr (1)
Discosphaerina Höhn. (21)
Exarmidium P. Karst. (14)
Frondicola K.D. Hyde (1)
Hyponectria Sacc. (30)
Lichenoverruculina Etayo (1)
Micronectria Speg. (4)
Papilionovela Aptroot (1)
Pellucida Dulym., Sivan., P.F. Cannon & Peerally (1)
Phragmitensis M.K.M. Wong, Poon & K.D. Hyde (2)
Physalospora Niessl (37)
Rachidicola K.D. Hyde & J. Fröhl. (1)
Xenothecium Höhn. (1)

Iodosphaeriaceae O. Hilber
Iodosphaeria Samuels (8)

Melogrammataceae G. Winter
Melogramma Fr. (20)

Phlogicylindriaceae Senan. & K.D. Hyde


Ciferriascosea Senan., Bhat, Camporesi & K.D. Hyde (2)
Idriellomyces Crous (1)
Phlogicylindrium Crous, Summerb. & Summerell (5)

Pseudomassariaceae Senan. & K.D. Hyde


Leiosphaerella Höhn. (15)
Pseudapiospora Petr. (3)
Pseudomassaria Jacz. (24)
Pseudomassariella Petr (1)

Pseudotruncatellaceae Crous
Pseudotruncatella R.H. Perera, Camporesi, Maharachch. & K.D. Hyde (2)

Sporocadaceae Corda*
Allelochaeta Petr. (50)
Annellolacinia B. Sutton (2)
Bartalinia Tassi (19)
Broomella Sacc. (2)
Ciliochorella Syd. (4)
Dilophospora Desm. (ca. 2 + few orphaned names)
Diploceras (Sacc.) Died (2)
Disaeta Bonar (1)
Discosia Lib. (ca. 17)
Distononappendiculata F. Liu, L. Cai & Crous (3)

1227
Diversimediispora F. Liu, L. Cai & Crous (1)
Doliomyces Steyaert (3)
Heterotruncatella F. Liu, L. Cai & Crous (17)
Hyalotiella Papendorf (6)
Hymenopleella Munk (= Dyrithiopsis L. Cai, Jeewon & K.D. Hyde; = Neotruncatella
Hyang B. Lee & T.T.T. Nguyen) (7)
Immersidiscosia Kaz. Tanaka, Okane & Hosoya (1)
Monochaetia (Sacc.) Allesch. (ca. 30)
Morinia Berl. & Bres. (=Zetiasplozna Nag Raj) (2)
Neopestalotiopsis Maharachch., K.D. Hyde & Crous (33)
Nonappendiculata F. Liu, L. Cai & Crous (1)
Parabartalinia F. Liu, L. Cai & Crous (1)
Pestalotiopsis Steyaert (ca. 100)
Pseudopestalotiopsis Maharachch., K.D. Hyde & Crous (12)
Pseudosarcostroma F. Liu, L. Cai & Crous (1)
Robillarda Sacc. (ca. 15)
Sarcostroma Cooke (28)
Seimatosporium Corda (ca. 100)
Seiridium Nees (20)
Sporocadus Corda (49)
Strickeria Körb. (10)
Synnemapestaloides T. Handa & Y. Harada (2)
Truncatella Steyaert (13)
Xenoseimatosporium F. Liu, L. Cai & Crous (1)

Vialaeaceae P.F. Cannon


Vialaea Sacc. (50)

Amphisphaeriales genus incertae sedis


Chitonospora E. Bommer, M. Rousseau & Sacc. (1)

Delonicicolales R.H. Perera, Maharachch. & K.D. Hyde


Delonicicolaceae R.H. Perera, Maharachch. & K.D. Hyde
Delonicicola R.H. Perera, Maharachch. & K.D. Hyde (1)
Furfurella Voglmayr & Jaklitsch (3)

Xylariales Nannf.
Anungitiomycetaceae Crous
Anungitiomyces Crous (1)

Barrmaeliaceae Voglmayr & Jaklitsch*


Barrmaelia Rappaz. (8)
Entosordaria (Sacc.) Höhn. (ca. 18)

Castanediellaceae Hern.-Restr., Guarro & Crous


Castanediella Hern.-Restr., Crous & M.J. Wingf. (12)

Clypeosphaeriaceae G. Winter
Aquasphaeria K.D. Hyde (1)
Apioclypea K.D. Hyde (7)
Brunneiapiospora K.D. Hyde, J. Fröhl. & Joanne E. Taylor (9)
Clypeosphaeria Fuckel (37)

1228
Crassoascus Checa, Barrasa & A.T. Martínez (3)
Palmaria K.D. Hyde, J. Fröhl. & Joanne E. Taylor (1)

Coniocessiaceae Asgari & Zare


Coniocessia Dania García, Stchigel, D. Hawksw. & Guarro (5)
Paraxylaria Wanas., E.B.G. Jones, Gafforov & K.D. Hyde (1)

Diatrypaceae Nitschke
Allocryptovalsa Senwanna, Phook. & K.D. Hyde (2)
Anthostoma Nitschke (ca. 101)
Cryptosphaeria Ces & De Not. (48)
Cryptovalsa Ces. & De Not. ex Fuckel (43)
Diatrypasimilis J.J. Zhou & Kohlm. (1)
Diatrype Fr. (ca. 244)
Diatrypella (Ces. & De Not.) De Not. (ca. 115)
Echinomyces Rappaz (2)
Endoxylina Romell (16)
Eutypa Tul. & C. Tul. (ca. 131)
Eutypella (Nitschke) Sacc. (ca. 196)
Halodiatrype Dayar. & K.D. Hyde (3)
Halocryptovalsa Dayar. & K.D. Hyde (2)
Leptoperidia Rappaz (4)
Libertella Desm. (ca. 72)
Monosporascus Pollack & Uecker (4)
Neoeutypella M. Raza, Q.J. Shang, Phook. & L. Cai (1)*
Pedumispora K.D. Hyde & E.B.G. Jones (1)
Peroneutypa Berl. (30)
Quaternaria Tul. & C. Tul. (14)

Fasciatisporaceae S.N. Zhang, K.D. Hyde & J.K. Liu


Fasciatispora K.D. Hyde (11)

Graphostromataceae M.E. Barr, J.D. Rogers & Y.M. Ju


Biscogniauxia Kuntze (ca. 76)
Camillea Fr. (50)
Graphostroma Piroz. (1)
Obolarina Pouzar (2)
Vivantia J.D. Rogers, Y.M. Ju & Cand. (1)

Hypoxylaceae DC.
Annulohypoxylon Y.M. Ju, J.D. Rogers & H.M. Hsieh (ca. 60)
Anthocanalis Daranag., Camporesi & K.D. Hyde (1)
Chlorostroma A.N. Mill., Lar.N. Vassiljeva & J.D. Rogers (3)
Daldinia Ces. & De Not. (ca. 67)
Durotheca Læssøe, Srikit., Luangsa-ard & M. Stadler (4)*
Entonaema Möller (10)
Hypomontagnella Sir, L. Wendt & C. Lambert (4)*
Hypoxylon Bull. (141)
Jackrogersella L. Wendt, Kuhnert & M. Stadler (6)
Natonodosa Heredia (1)
Phylacia Lév. (12)
Pyrenomyxa Morgan (3)

1229
Pyrenopolyporus Lloyd (5)
Rhopalostroma D. Hawksw. (11)
Rostrohypoxylon J. Fourn. & M. Stadler (1)
Ruwenzoria J. Fourn., M. Stadler, Læssøe & Decock (1)
Thamnomyces Ehrenb. (11)
Theissenia Maubl. (8)
Thuemenella Penz. & Sacc. (10)

Induratiaceae Samarak., Thongbai, K.D. Hyde & M. Stadler


Emarcea Duong, Jeewon & K.D. Hyde (3)
Induratia Samuels, E. Müll. & Petrini (26)

Leptosilliaceae Voglmayr & Jaklitsch


Leptosillia Höhn. (=Cresporhaphis M.B. Aguirre) (9)*

Lopadostomataceae Daranag. & K.D. Hyde


Creosphaeria Theiss. (3)
Jumillera J.D. Rogers, Y.M. Ju & F. San Martín (8)
Lopadostoma (Nitschke) Traverso (27)
Whalleya J.D. Rogers, Y.M. Ju & F. San Martín (2)

Microdochiaceae Hern.-Restr., Crous & J.Z. Groenew.


Idriella P.E. Nelson & S. Wilh. (=Monographella Petr.) (24)
Microdochium Syd. (38)
Selenodriella R.F. Castañeda & W.B. Kendr. (7)

Myelospermataceae K.D. Hyde & S.W. Wong


Myelosperma Syd. & P. Syd. (5)

Nothodactylariaceae Crous
Nothodactylaria Crous (1)

Oxydothidaceae Konta & K.D. Hyde


Oxydothis Penz. & Sacc. (79)

Polystigmataceae Höhn. ex Nannf.*


Polystigma DC. (23)

Pseudosporidesmiaceae Crous
Pseudosporidesmium K.D. Hyde & McKenzie (2)

Requienellaceae Boise*
Acrocordiella O.E. Erikss. (2)
Lacrymospora Aptroot (1)
Parapyrenis Aptroot (8)
Requienella Fabre (8)

Xyladictyochaetaceae Crous & Hern.-Restr*


Xyladictyochaeta Hern.-Restr., R.F. Castañeda & Gené (1)

Xylariaceae Tul. & C. Tul.


Abieticola Hyang B. Lee (1)

1230
Amphirosellinia Y.M. Ju, J.D. Rogers, H.M. Hsieh & Lar.N. Vassiljeva (6)
Anthostomella Sacc. (ca. 100)
Anthostomelloides Tibpromma & K.D. Hyde (5)
Ascotricha Berk. (27)
Astrocystis Berk. & Broome (24)
Brunneiperidium Daranag., Camporesi & K.D. Hyde (2)
Collodiscula I. Hino & Katum. (5)
Coniolariella Dania García, Stchigel & Guarro (5)
Engleromyces Henn. (2)
Entalbostroma J.D. Rogers & P.R. Johnst. (1)
Entoleuca Syd. (3)
Euepixylon Füisting (2)
Halorosellinia Whalley, E.B.G. Jones, K.D. Hyde & Læssøe (3)
Helicogermslita Lodha & D. Hawksw. (9)
Hypocopra (Fr) J. Kickx f . (58)
Hypocreodendron Henn. (1)
Kretzschmaria Fr. (ca. 57)
Kretzschmariella Viégas (2)
Leprieuria Læssøe, J.D. Rogers & Whalley (1)
Lunatiannulus Daranag., Camporesi & K.D. Hyde (1)
Nemania Gray (57)
Podosordaria Ellis & Holw. (35)
Poronia Willd. (ca. 24)
Rosellinia De Not. (ca. 359)
Sarcoxylon Cooke (6)
Squamotubera Henn. (1)
Stilbohypoxylon Henn. (12)
Vamsapriya Gawas & Bhat (8)
Virgaria Nees (11)
Wawelia Namysl. (5)
Xylaria Hill ex Schrank (ca. 571)

Zygosporiaceae J.F. Li, Phook. & K.D. Hyde.


Zygosporium Mont. (25)

Xylariales genera incertae sedis


Adomia S. Schatz (1)
Alloanthostomella Daranag., Camporesi & K.D. Hyde (1)
Anungitea B. Sutton (22)
Ascotrichella Valldos. & Guarro (1)
Basifimbria Subram. & Lodha (1)
Biporispora J.D. Rogers, Y.M. Ju & Cand. (1)
Botryohypoxylon Samuels & J.D. Rogers (1)
Castellaniomyces Senan., Camporesi & K.D. Hyde (1)
Chaenocarpus Rebent. (4)
Circinotrichum Nees (15)
Cryptostroma P.H. Greg. & S. Waller (1)
Cyanopulvis J. Fröhl. & K.D. Hyde (1)
Diamantinia A.N. Mill., Læssøe & Huhndorf (1)
Gigantospora B.S. Lu & K.D. Hyde (1)
Guestia G.J.D. Sm. & K.D. Hyde (1)
Gyrothrix (Corda) Corda (22)

1231
Hadrotrichum Fuckel (22)
Idriellopsis Hern.-Restr. & Crous (1)
Kirstenboschia Quaedvl., Verkley & Crous (1)
Lanceispora Nakagiri, Okane, Tad. Ito & Katum. (2)
Lasiobertia Sivan. (2)
Leptomassaria Petr. (4)
Melanographium Sacc. (11)
Neoanthostomella D.Q. Dai & K.D. Hyde (2)
Neoidriella Hern.-Restr. & Crous (1)
Nipicola K.D. Hyde (4)
Occultitheca J.D. Rogers & Y.M. Ju (1)
Ophiorosellinia J.D. Rogers, A. Hidalgo, F.A. Fernández & Huhndorf (1)
Palmicola K.D. Hyde (4)
Pandanicola K.D. Hyde (2)
Paraidriella Hern.-Restr. & Crous (1)
Paramphisphaeria F.A. Fernández, J.D. Rogers, Y.M. Ju, Huhndorf & L. Umaña (1)
Paraphysalospora Crous (1)
Paucithecium Lloyd (1)
Pidoplitchkoviella Kiril. (1)
Polyancora Voglmayr & Yule (1)
Polyscytalum Riess (28)
Poroleprieuria M.C. González, Hanlin, Ulloa & Elv. Aguirre (1)
Pseudoanthostomella Daranag., Camporesi & K.D. Hyde (5)
Pseudophloeospora Crous & R.G. Shivas (2)
Pseudosubramaniomyces Crous (1)
Pulmosphaeria Joanne E. Taylor, K.D. Hyde & E.B.G. Jones (1)
Pyriformiascoma Daranag., Camporesi & K.D. Hyde (1)
Roselymyces Fiuza, C.R. Silva, R.F. Castañeda & Gusmão (1)*
Sabalicola K.D. Hyde (1)
Spirodecospora B.S. Lu, K.D. Hyde & W.H. Ho (2)
Sporidesmina Subram. & Bhat (1)
Striatodecospora D.Q. Zhou, K.D. Hyde & B.S. Lu (1)
Stromatoneurospora S.C. Jong & E.E. Davis (2)
Surculiseries Okane (1)
Synnemadiella Crous & M.J. Wingf. (1)
Tristratiperidium Daranag., Camporesi & K.D. Hyde (1)
Xenoanthostomella Mapook & K.D. Hyde (1)
Xylocrea Möller (2)
Xylotumulus J.D. Rogers, Y.M. Ju & Cand. (1)
Yuea O.E. Erikss. (1)

Xylariomycetidae family incertae sedis


Cainiaceae J.C. Krug
Alishanica A. Karunarathna, C.H. Kuo & K. D. Hyde (1)
Amphibambusa D.Q. Dai & K.D. Hyde (1)
Arecophila K.D. Hyde (14)
Atrotorquata Kohlm. & Volkm.-Kohlm. (2)
Cainia Arx & E. Müll. (6)
Seynesia Sacc. (ca. 46)

Xylariomycetidae genus incertae sedis


Calceomyces Udagawa & S. Ueda (1)

1232
Sordariomycetes orders incertae sedis
Amplistromatales M.J. D'souza, Maharachch. & K.D. Hyde
Amplistromataceae Huhndorf, A.N. Mill., Greif & Samuels*
Acidothrix Hujslová & M. Kolařík (1)
Amplistroma Huhndorf, A.N. Mill., Greif & Samuels (9)
Wallrothiella Sacc. (ca. 10)

Catabotryaceae Petr. ex M.E. Barr


Catabotrys Theiss. & Syd. (3)

Parasympodiellales Hern.-Restr., Gené, R.F. Castañeda & Crous


Parasympodiellaceae Hern.-Restr., Gené, Guarro & Crous
Parasympodiella Ponnappa (10)

Spathulosporales Kohlm.
Hispidicarpomycetaceae Nakagiri
Hispidicarpomyces Nakagiri (1)

Spathulosporaceae Kohlm.
Retrostium Nakagiri & Tad Ito (1)
Spathulospora A.R. Caval. & T.W. Johnson (4)

Tracyllalales Crous
Tracyllaceae Crous
Tracylla (Sacc.) Tassi (3)

Vermiculariopsiellales Hern.-Restr., J. Mena, Gené & Crous


Vermiculariopsiellaceae Hern.-Restr, J. Mena, Gené & Crous
Vermiculariopsiella Bender (22)

Sordariomycetes families incertae sedis


Acrodictyaceae J.W. Xia & X.G. Zhang*
Acrodictys M.B. Ellis (25)

Junewangiaceae J.W. Xia & X.G. Zhang*


Dictyosporella Abdel-Aziz (2)
Junewangia W.A. Baker & Morgan-Jones (6)

Lautosporaceae Kohlm., Volkm.-Kohlm. & O.E. Erikss


Lautospora K. D. Hyde & E.B.G. Jones (2)

Obryzaceae Körb.
Obryzum Wallr. (3)

Sordariomycetes genera incertae sedis


Acerbiella Sacc. (4)
Acrospermoides Miller & G.E. Thomps. (2)
Ameromassaria Hara (1)
Amphisphaerellula Gucevič (1)
Amphisphaerina Höhn. (3 epithets in Index Fungorum 2020)
Amphorulopsis Petr. (1)
Amylis Speg. (1)

1233
Anisomycopsis I. Hino & Katum. (1)
Antennopsis R. Heim (1)*
Anthostomaria (Sacc.) Theiss. & Syd. (1)
Anthostomellina L.A. Kantsch. (2)
Apodothina Petr. (1)
Apogaeumannomyces Matsush. (1)
Aquadulciospora Fallah & Shearer (1)
Areolospora S.C. Jong & E.E. Davis (2 epithets in Index Fungorum 2020)
Aropsiclus Kohlm. & Volkm.-Kohlm. (1)
Ascorhiza Lecht.-Trinka (1)
Ascoyunnania L. Cai & K.D. Hyde (1)
Atrogeniculata J.S. Monteiro, Gusmão & R.F. Castañeda (1)
Aulospora Speg. (1)
Azbukinia Lar.N. Vassiljeva (1)
Bactrosphaeria Penz. & Sacc. (1)
Basidiobotrys Höhn. (1)
Biciliopsis Diederich (2)
Bombardiastrum Pat. (1)
Boothiella Lodhi & Mirza (1)
Botryosporium Corda (11)
Brenesiella Syd. (1)
Byrsomyces Cavalc. (1)
Byssotheciella Petr. (2)
Caleutypa Petr. (1)
Caproniella Berl. (1)
Chaetoamphisphaeria Hara (1)
Charonectria Sacc. (3)
Ciliofusospora Bat. & J.L. Bezerra (1)
Clypeoceriospora Sousa da Câmara (1)
Clypeosphaerulina Sousa da Câmara (1)
Cryptoascus Petri (2)
Cryptomycella Höhn. (2)
Cryptomycina Höhn. (2)
Cucurbitopsis Bat. & Cif. (1)
Curvatispora V.V. Sarma & K.D. Hyde (1)
Dasysphaeria Speg. (1)
Delpinoëlla Sacc. (1)
Diacrochordon Petr. (1)
Didymobotryum Sacc. (6)
Duradens Samuels & Rogerson (1)
Ellisembia Subram. (ca. 60)
Esfandiariomyces Ershad (1)
Fantasmomyces Dong Hyeon Lee, Marinc., Z.W. de Beer & M.J. Wingf. (1)
Farrowia D. Hawksw. (3)
Fassia Dennis (1)
Flammispora Pinruan, Sakay., K.D. Hyde & E.B.G. Jones (2)
Frondisphaeria K.D. Hyde (2)
Hapsidascus Kohlm. & Volkm.-Kohlm. (1)
Hassea Zahlbr. (1)
Heliastrum Petr. (1)
Hyaloderma Speg. (1)
Hyalotiopsis Punith. (1)

1234
Hydronectria Kirschst. (1)
Immersisphaeria Jaklitsch (1)
Iraniella Petr. (1)
Konenia Hara (1)
Kravtzevia Schwartzman (1)
Kurssanovia Kravtzev (1)
Lecythiomyces Doweld (1)
Leptosacca Syd. (1)
Leptosphaerella Speg. (14 epithets in Index Fungorum 2020)
Mangrovispora K.D. Hyde & Nageire (1)
Marisolaris Jørg. Koch & E.B.G. Jones (1)
Microcyclephaeria Bat. (1)
Mirannulata Huhndorf, F.A. Fernández, A.N. Mill. & Lodge (2)
Mycothermus D.O. Natvig, J.W. Taylor, A. Tsang, M.I. Hutch. & A.J. Powell ex X. Wei
Wang, Houbraken & D.O. Natvig (2)
Natantiella Réblová (1)
Naumovela Kravtzev (2)
Neocryptospora Petr. (1)
Neoeriomycopsis Crous & M.J. Wingf. (1)
Neolamya Theiss. & Syd. (3)
Neothyridaria Petr. (1)
Ophiomassaria Jacz. (1)
Ophiomeliola Starbäck (3)
Paoayensis Cabanela, Jeewon & K.D. Hyde (2)
Paradiplococcium Hern.-Restr., J. Mena & Gené (1)
Paramicrodochium Hern.-Restr. & Crous (1)
Pareutypella Y.M. Ju & J.D. Rogers (2)
Phialemoniopsis Perdomo, Dania García, Gené, Cano & Guarro (6)
Phragmeriella Hansf. (1)
Phyllocelis Syd. (2)
Pleocryptospora J. Reid & C. Booth (1)
Pleosphaeria Speg. (24)
Pleurophragmium Costantin (22)
Protocucurbitaria Naumov (1)
Pulvinaria Bon. (2)
Pumilus Viala & Marsais (1)
Rehmiomycella E. Müll. (1)
Rhamphosphaeria Kirschst. (1)
Rhizophila K.D. Hyde & E.B.G. Jones (1)
Rhopographella (Henn.) Sacc. & Trotter (2)
Rhynchosphaeria (Sacc.) Berl. (5)
Rivulicola K.D. Hyde (3)
Romellina Petr. (1)
Saccardoëlla Speg. (15)
Sartorya Vuill. (9 epithets in Index Fungorum 2020)
Scharifia Petr. (1)
Scoliocarpon Nyl. (1 epithets in Index Fungorum 2020)
Scotiosphaeria Sivan. (1)
Selenosporella G. Arnaud ex MacGarvie (12)
Servazziella J. Reid & C. Booth (1)
Sporoctomorpha J.V. Almeida & Sousa da Câmara (1)
Stanjehughesia Subram. (16)

1235
Stearophora L. Mangin & Viala (1)
Steganopycnis Syd. & P. Syd. (1)
Stegophorella Petr. (1)
Stellosetifera Matsush. (1)
Stereosphaeria Kirschst. (1)
Stomatogenella Petr. (1)
Sungaiicola Fryar & K.D. Hyde (1)
Synsphaeria Bon. (4 epithets in Index Fungorum 2020)
Teracosphaeria Réblová & Seifert (1)
Thelidiella Fink (1)
Thyridella (Sacc.) Sacc. (3)
Thyrotheca Kirschst. (1 epithets in Index Fungorum 2020)
Trichospermella Speg. (2)
Trichosphaeropsis Bat. & Nasc. (1)
Tulipispora Révay & Gönczöl (1)
Tunstallia Agnihothr. (3 epithets in Index Fungorum 2020)
Urosporella G.F. Atk. (5)
Urupe Viégas (1)
Vleugelia J. Reid & C. Booth (1)
Xenodium Syd. (1)
Zalerion R.T. Moore & Meyers (6)

Xylonomycetes Gazis & P. Chaverri


Symbiotaphrinales Baral & E. Weber
Symbiotaphrinaceae Baral & E. Weber
Symbiotaphrina Kühlw. & Jurzitza ex W. Gams & Arx (17)*

Xylonales Gazis & P. Chaverri


Xylonaceae Gazis & P. Chaverri
Trinosporium Crous & Decock (1)
Xylona Gazis & P. Chaverri (1)

Xylobotryomycetes Voglmayr & Jaklitsch


Xylobotryales Voglmayr & Jaklitsch
Cirrosporiaceae Voglmayr & Jaklitsch
Cirrosporium S. Hughes (1)

Xylobotryaceae Voglmayr & Jaklitsch


Xylobotryum Pat. (2)

Pezizomycotina orders incertae sedis


Thelocarpales Lücking & Lumbsch
Thelocarpaceae Zukal
Sarcosagium A. Massal. (1)
Thelocarpon Nyl (25)

Vezdaeales Lumbsch & Lücking


Vezdaeaceae Poelt & Vězda ex J.C. David & D. Hawksw.
Vezdaea Tsch.-Woess & Poelt (12)

Pezizomycotina family incertae sedis


Harpidiaceae Vězda ex Hafellner

1236
Euopsis Nyl. (2)
Harpidium Körb. (3)

Pezizomycotina genera incertae sedis


Angatia Syd. (5)
Biatoridium J. Lahm ex Körb. (3)
Cyanoporina Groenh. (1)
Melanophloea P. James & Vězda (1)
Milospium D. Hawksw. (4)
Oevstedalia Ertz & Diederich (1)
Psammina Sacc. & M. Rousseau ex E. Bommer & M. Rousseau (8)
Pygmaeosphaera Etayo & Diederich (3)
Pyrenocollema Reinke (1)
Solanella Vaňha (1)*
Wadeana Coppins & P. James (2)

Saccharomycotina O.E. Erikss. & Winka


Saccharomycetes O.E. Erikss. & Winka*
Saccharomycetales Kudrjanzev
Alloascoideaceae Kurtzman & Robnett
Alloascoidea Kurtzman & Robnett (2)

Ascoideaceae J. Schröter
Ascoidea Bref. (4)

Cephaloascaceae L.R. Batra


Cephaloascus Hanawa (2)

Debaryomycetaceae Kurtzman & M. Suzuki


Babjeviella Kurtzman & M. Suzuki (1)
Debaryomyces Lodder & Kreger-van Rij (15)
Hemisphaericaspora Hui, Ren, Chen, Li, Zhan & Niu (2)
Kurtzmaniella M.A. Lachance & W.T. Starmer (5)
Lodderomyces Van der Walt (2)*
Meyerozyma Kurtzman & M. Suzuki (8)
Millerozyma Kurtzman & M. Suzuki (5)
Priceomyces Kurtzman & M. Suzuki (8)
Scheffersomyces Kurtzman & M. Suzuki (18)
Schwanniomyces Klöcker emend. M. Suzuki & Kurtzman (7)
Spathaspora N.H. Nguyen, S.O. Suh & M. Blackwell (11)
Suhomyces M. Blackwell & Kurtzman (26)
Yamadazyma Billon-Grand (23)

Dipodascaceae Engl. & E. Gilg


Dipodascus Lagerh. (14)
Galactomyces Redhead & Malloch (5)
Geotrichum Link (8)
Magnusiomyces Zender (7)
Saprochaete Coker & Shanor ex D.T.S. Wagner & Dawes (10)*

Lipomycetaceae E.K. Novák & Zsolt


Dipodascopsis Batra & P. Millner emend. Kurtzman, Albertyn & Basehoar-Powers (3)

1237
Kockiozyma Jindam., Yukphan & Y. Yamada (1)
Limtongia Jindam., Am-in, Yukphan & Y. Yamada (1)
Lipomyces Lodder & Kreger (16)
Myxozyma Van der Walt, Weijman & von Arx (12)

Metschnikowiaceae T. Kamienski*
Clavispora Rodr. Mir. (4)
Kodamaea Y. Yamada, T. Suzuki, Matsuda & Mikata emend. Rosa, Lachance, Starmer,
Barker, Bowles & Schlag-Edler (8)
Metschnikowia T. Kamienski (64)

Phaffomycetaceae Y. Yamada, H. Kawas., Nagats., Mikata & Tats. Seki


Barnettozyma Kurtzman, Robnett & Basehoar-Powers (9)
Cyberlindnera Minter (27)
Phaffomyces Y. Yamada (4)
Starmera Y. Yamada, Higashi, Ando & Mikata (7)
Wickerhamomyces Kurtzman, Robnett & Basehoar-Powers (31)

Pichiaceae Zender
Brettanomyces Kufferath & van Laer (3)
Dekkera Van der Walt (2)*
Komagataella Y. Yamada, Matsuda, Maeda & Mikata (6)
Kregervanrija Kurtzman (3)
Kuraishia Y. Yamada, Maeda & Mikata (9)
Martiniozyma Kurtzman (2)
Ogataea Y. Yamada, K. Maeda & Mikata (46)
Pachysolen Boidin & Adzet (1)
Pichia E.C. Hansen (27)
Saturnispora Z.W. Liu & Kurtzman (21)

Saccharomycetaceae G. Winter
Citeromyces Santa Maria (4)
Cyniclomyces Van der Walt & D.B. Scott (1)
Eremothecium Borzi emend. Kurtzman (5)
Hagleromyces Sousa, Morais, Lachance & Rosa (1)
Kazachstania Zubcova (44)
Kluyveromyces Van der Walt (6)
Lachancea Kurtzman (10)
Nakaseomyces Kurtzman (2)
Naumovozyma Kurtzman (3)
Saccharomyces Meyen (10)
Tetrapisispora Ueda-Nishimura & K. Mikata emend. Kurtzman (9)
Torulaspora Lindner (8)
Vanderwaltozyma Kurtzman (4)
Yueomyces Q.M. Wang, L. Wang, M. Groenewald & T. Boekhout (1)
Zygosaccharomyces B.T.P. Barker (11)
Zygotorulaspora Kurtzman (4)

Saccharomycodaceae Kudrjanzev
Hanseniaspora Zikes (17)*
Saccharomycodes E.C. Hansen (2)

1238
Saccharomycopsidaceae Arx & Van der Walt
Ambrosiozyma Van der Walt (14)
Saccharomycopsis Schiønning (19)

Trichomonascaceae Kurtzman & Robnett*


Blastobotrys Klopotek (23)
Diddensiella Péter, Dlauchy & Kurtzman (1)
Groenewaldozyma Kurtzman (3)
Spencermartinsiella Péter, Dlauchy, Tornai-Lehoczki, M. Suzuki & Kurtzman (4)
Starmerella Rosa & Lachance (44)
Sugiyamaella Kurtzman & Robnett (27)
Trichomonascus H.S. Jackson emend. Kurtzman & Robnett (6)*
Wickerhamiella Van der Walt (39)
Zygoascus M.T. Sm. (8)

Trigonopsidaceae M.A. Lachance & C.P. Kurtzman


Botryozyma Shann & M.T. Sm. emend. Lachance & Kurtzman (4)*
Tortispora Lachance & Kurtzman (8)
Trigonopsis Schachner emend. Kurtzman & Robnett (4)

Saccharomycetales genera incertae sedis


Aciculoconidium D.S. King & S.C. Jong (1)
Candida Berkhout (316)*
Coccidiascus Chatton (1)
Conidiascus Holterm. (1)
Danielozyma Kurtzman & Robnett (2)
Deakozyma Kurtzman & Robnett (2)
Diutina Khunnamwong, Lertwattanasakul, Jindam., Limtong & Lachance (10)
Endomyces Reess (4)
Hyphopichia von Arx & van der Walt (12)
Macrorhabdus Tomaszewski, Logan, Snowden, Kurtzman & Phalen. (1)
Metahyphopichia Sipiczki & Pfliegler (1)
Middelhovenomyces Kurtzman & Robnett (2)
Nadsonia Syd. (3)
Nakazawaea Y. Yamada, Maeda & Mikata (13)
Oscarbrefeldia Holterm. (1)*
Peterozyma Kurtzman & Robnett (2)
Phialoascus Redhead & Malloch (1)
Sporopachydermia Rodr. Mir. (3)
Teunomyces Kurtzman & M. Blackwell (12)
Wickerhamia Soneda (1)
Yarrowia Van der Walt & Arx (12)

TAPHRINOMYCOTINA O.E. Erikss. & Winka


Archaeorhizomycetes Rosling & T.Y. James
Archaeorhizomycetales Rosling & T.Y. James
Archaeorhizomycetaceae Rosling & T.Y. James
Archaeorhizomyces Rosling & T.Y. James (2)

Neolectomycetes O.E. Erikss. & Winka


Neolectales Landvik, O.E. Erikss., Gargas & P. Gust.
Neolectaceae Redhead

1239
Neolecta Speg. (3)

Pneumocystomycetes O.E. Erikss. & Winka


Pneumocystidales O.E. Erikss.
Pneumocystidaceae O.E. Erikss.
Pneumocystis P. Delanoë & Delanoë (5)

Schizosaccharomycetes O.E. Erikss. & Winka


Schizosaccharomycetales O.E. Erikss.
Schizosaccharomycetaceae Beij. ex Klöcker
Schizosaccharomyces Lindner (4)

Taphrinomycetes O.E. Erikss. & Winka


Taphrinales Gäum. & C.W. Dodge
Protomycetaceae Gray
Buerenia M.S. Reddy & C.L. Kramer (4)
Protomyces Unger (ca. 10)
Protomycopsis Magnus (5)
Saitoella Goto, Sugiy., Hamam. & Komag. (2)
Taphridium Lagerh. & Juel ex Juel (2)
Volkartia Maire (1)

Taphrinaceae Gäum.
Taphrina Fr. (ca. 95)

Ascomycota families incertae sedis


Aphanopsidaceae Printzen & Rambold
Aphanopsis Nyl. ex Syd. (1)
Steinia Körb. (3)

Diporothecaceae R.K. Mibey & D. Hawksw.


Diporotheca C.C. Gordon & C.G. Shaw (4)

Eoterfeziaceae G.F. Atk.


Acanthogymnomyces Udagawa & Uchiyama (1)
Eoterfezia G.F. Atk. (2)

Mucomassariaceae Petr. & Cif.


Mucomassaria Petr. (1)

Saccardiaceae Höhn.
Ascolectus Samuels & Rogerson (1)
Cyanodiscus E. Müll. & M.L. Farr (2)
Henningsiella Rehm (2)
Phillipsiella Cooke (7)
Pseudodiscus Arx & E. Müll. (1)
Saccardia Cooke (3)
Schenckiella P. Henn. (1)

Seuratiaceae Vuill. ex M.E. Barr


Seuratia Pat. (5)
Seuratiopsis Woron. (1)

1240
Strangosporaceae S. Stenroos, Miądl. & Lutzoni
Strangospora Körb. (ca. 11)

Ascomycota genera incertae sedis


Abropelta B. Sutton (1)
Acarellina Bat. & H. Maia (1)
Acaroconium Kocourk. & D. Hawksw. (1)
Acarocybe Syd. (3)
Acarocybella M.B. Ellis (1)
Acarocybellina Subram. (1)
Acarocybiopsis J. Mena, A. Hern.-Gut. & Mercado (1)
Acaropeltis Petr. (1)
Achoropeltis Syd. (1)
Acleistia Bayl. Ell. (1)
Acontium Morgan (4)
Acrodictyella W.A. Baker & Partr. (1)
Acrodictyopsis P.M. Kirk (1)
Acrodontiella U. Braun & Scheuer (1)
Acrophragmis Kiffer & Reisinger (4)
Acrospeira Berk. & Broome (1)
Acrostaurus Deighton & Piroz. (1)
Actinocladium Ehrenb. (6)
Actinotexis Arx (1)
Actinothecium Ces. (5)
Actinothyrium Kunze (10)
Acumispora Matsush. (5)
Agaricodochium X.J. Liu (1)
Agarwalomyces R.K. Verma & Kamal (1)
Agrabeeja Subram. (1)
Agyriella Sacc. (2)
Agyriellopsis Höhn. (3)
Ahmadia Syd. (1)
Ajrekarella Kamat & Kalani (1)
Alatosessilispora K. Ando & Tubaki (1)
Alciphila Harmaja (1)
Algonquinia R.F. Castañeda & W.B. Kendr. (1)
Allantophomoides S.L. Wei & T.Y. Zhang (1)
Alloneottiosporina Nag Raj (2)
Allophoron Nádv. (1)
Allothyriella Bat., Cif. & Nascim. (3)
Allothyrina Bat. & J.L. Bezerra (1)
Allothyriopsis Bat., Cif. & H. Maia (1)
Alpakesa Subram. & K. Ramakr. (4)
Alpakesiopsis Abbas, B. Sutton, Ghaffar & A. Abbas (1)
Alveariospora Meir. Silva, R.F. Castañeda, O.L. Pereira & R.W. Barreto (1)
Alveophoma Alcalde (1)
Alysidiopsis B. Sutton (5)
Amallospora Penz. (1)
Amblyosporium Fresen. (4)
Ameroconium U. Braun & Zhurb. (1)
Amerodiscosiella M.L. Farr (1)
Amerodiscosiellina Bat. & Cavalc. (1)

1241
Amerosporiopsis Petr. (1)
Amerosympodula Matsush. (1)
Amoenodochium Peláez & R.F. Castañeda (1)
Amoenomyces R.F. Castañeda, Saikawa & Hennebert (1)
Amphichaetella Höhn. (1)
Amphophialis R.F. Castañeda, W.B. Kendr. & Guarro (1)
Amphoropycnium Bat. (1)
Ampullicephala R.F. Castañeda, Minter & M. Stadler (1)
Ampulliferina B. Sutton (2)
Amylogalla Suija, Motiej. & Kantvilas (1)
Anabahusakala Carmo, J.S. Monteiro, Gusmão & R.F. Castañeda (1)
Anacraspedodidymum C.R. Silva, R.F. Castañeda & Gusmão (2)
Anaexserticlava Santa Izabel, R.F. Castañeda & Gusmão (1)
Anaphysmene Bubák (2)
Anarhyma M.H. Pei & Z.W. Yuan (1)
Anaselenosporella Heredia, R.F. Castañeda & R.M. Arias (2)
Anaseptoidium R.F. Castañeda, Heredia & R.M. Arias (1)
Anaverticicladus P.O. Costa, Malosso & R.F. Castañeda (1)
Ancoraspora Mig. Rodr. (1)
Ancorasporella J. Mena, Mercado & Heredia (1)
Angiopomopsis Höhn. (1)
Angulimaya Subram. & Lodha (1)
Angulospora Sv. Nilsson (1)
Annellodentimyces Matsush. (1)
Annellodochium Deighton (1)
Annellophorella Subram. (5)
Annellospermosporella P.R. Johnst. (1)
Antennatula Fr. ex F. Strauss (10)
Anthracoderma Speg. (3)
Antimanoa Syd. (1)
Antromyces Fresen. (4)
Anulohypha Cif. (1)
Anungitopsis R.F. Castañeda & W.B. Kendr. (7)
Aoria Cif. (1)
Aphanofalx B. Sutton (2)
Apiocarpella Syd. & P. Syd. (8)
Apiotypa Petr. (1)
Apogloeum Petr. (1)
Apomelasmia Grove (8)
Aporellula B. Sutton (2)
Aposporella Thaxt. (1)
Apostrasseria Nag Raj (2)
Arachnophora Hennebert (11)
Arachnospora R.F. Castañeda, Minter & Camino (1)
Arborillus Munt.-Cvetk. & Gómez-Bolea (1)
Arborispora K. Ando (4)
Arcuadendron Sigler & J.W. Carmich. (2)
Ardhachandra Subram. & Sudha (3)
Argentinomyces Peña & Arambarri (1)
Argopericonia B. Sutton & Pascoe (2)
Aristastoma Tehon (1)
Arthrobotryum Ces. (5)

1242
Arthrocristula Sigler, M.T. Dunn & J.W. Carmich. (1)
Arthromoniliphora S.S. Silva, Gusmão & R.F. Castañeda (1)
Arthrosporium Sacc. (2)
Arthrowallemia R.F. Castañeda, Dania García & Guarro (2)
Articulophora C.J.K. Wang & B. Sutton (1)
Artocarpomyces Subram. (1)
Ascochytopsis Henn. (5)
Ascochytulina Petr. (3)
Ascofascicula Matsush. (6)
Ascomauritiana V.M. Ranghoo & K.D. Hyde (1)
Ascosubramania Rajendran (1)
Ashtaangam Subram. (1)
Aspilaima Bat. & H. Maia (1)
Astelechia Cif. (2)
Asterinothyriella Bat. & Cif. (3)
Asterinothyrium Bat., Cif. & H. Maia (1)
Asteroconium Syd. & P. Syd. (2)
Asteroglobulus Brackel (2)
Asteromyces F. Moreau & V. Moreau (1)
Asterophoma D. Hawksw. (1)
Asteroscutula Petr. (1)
Asterostomopora Bat. & H. Maia (1)
Asterostomopsis Bat., Cif. & H. Maia (1)
Asterostomula Theiss. (4)
Asterostomulina Bat., J.L. Bezerra & H. Maia (1)
Astomella Thirum. (1)
Astronatelia Bat. & H. Maia (1)
Atractilina Dearn. & Barthol. (2)
Atractobolus Tode (1)
Atrosetaphiale Matsush. (1)
Atrosynnema J.W. Xia, X.G. Zhang & Z. Li (1)
Aurosphaeria Sun J. Lee, Strobel, Eisenman, Geary, P.N. Vargas & S.A. Strobel (1)
Avesicladiella W.P. Wu, B. Sutton & Gange (2)
Avettaea Petr. & Syd. (3)
Bacillopeltis Bat. (1)
Bactridium Kunze (15)
Bactrodesmiella M.B. Ellis (2)
Baculospora Zukal (1)
Badarisama Kunwar, J.B. Manandhar & J.B. Sinclair (1)
Bahuchashaka Subram. (1)
Bahugada K.A. Reddy & Vasant Rao (2)
Bahukalasa Subram. & Chandrash. (1)
Balaniopsis P.M. Kirk (4)
Balanium Wallr. (1)
Barbarosporina Ķirulis (1)
Barnettella D. Rao & P. Rag. Rao (1)
Basauxia Subram. (1)
Batistina Peres (1)
Batistospora J.L. Bezerra & M.M.P. Herrera (1)
Beauveriphora Matsush. (1)
Beccopycnidium F. Stevens (1)
Beejadwaya Subram. (1)

1243
Belemnospora P.M. Kirk (7)
Bellulicauda B. Sutton (2)
Beltramono Rashmi Dubey, A.K. Pandey bis & Manohar. (1)
Beltraniomyces Manohar., D.K. Agarwal & Rao (1)
Beniowskia Racib. (4)
Benjpalia Subram. & Bhat (1)
Berggrenia Cooke (2)
Bhadradriella Nagaraju, Kunwar & Manohar. (1)
Bhadradriomyces Sureshk., Manohar. & Kunwar (1)
Bharatheeya D'Souza & Bhat (3)
Bhatia W.A. Baker & Morgan-Jones (2)
Bibanasiella R.F. Castañeda & W.B. Kendr. (1)
Bicoloromyces Heuchert, U. Braun & D. Hawksw. (1)
Biflagellospora Matsush. (1)
Biflagellosporella Matsush. (1)
Biflua Jørgen Koch & E.B.G. Jones (1)
Bimeris Petr. (1)
Bioconiosporium Bat. & J.L. Bezerra (2)
Biophomopsis Petr. (3)
Bisbyopeltis Bat. & A.F. Vital (1)
Bispora Corda (31)
Bisseomyces R.F. Castañeda (1)
Blastocatena Subram. & Bhat (2)
Blastodictys M.B. Ellis (1)
Blastofusarioides Matsush. (1)
Blastophorella Boedijn (1)
Blastophragma Subram. (4)
Blennoria Moug. & Fr. (4)
Blennoriopsis Petr. (1)
Bleptosporium Steyaert (4)
Blodgettia Harv. (2)
Bostrichonema Ces. (4)
Botryoderma Papendorf & H.P. Upadhyay (4)
Botryodiplodina Dias & Sousa da Câmara (1)
Botryomonilia Goos & Piroz. (1)
Botryostroma Höhn. (2)
Brachycephala J.S. Monteiro, Gusmão & R.F. Castañeda (1)
Brachydesmiella G. Arnaud ex S. Hughes (8)
Brachysporiellina Subram. & Bhat (2)
Brachysporiopsis Yanna, W.H. Ho & K.D. Hyde (1)
Braunomyces V.A. Melnik & Crous (1)
Brefeldiopycnis Petr. & Cif. (1)
Brencklea Petrak (1)
Brevicatenospora R.F. Castañeda, Minter & Saikawa (1)
Briosia Cavara (6)
Brycekendrickia Nag Raj (1)
Bryophytomyces Cif. (1)
Bulbilopycnis Matsush. (1)
Bulbocatenospora R.F. Castañeda & Iturr. (1)
Bullaserpens Bat., J.L. Bezerra & Cavalc. (1)
Cacumisporium Preuss (9)

1244
Caeruleoconidia Zhurb. & Pino-Bodas (= Caeruleoconidia Zhurb. & Diederich 2015 nom.
inv.) (2)
Calcarispora Marvanová & Marvan (1)
Calceispora Matsush. (2)
Callistospora Petr. (1)
Calocline Syd. (1)
Calongeomyces D. Hawksw. & Etayo (1)
Camaroglobulus Speer (1)
Camaropycnis E.K. Cash (1)
Camarosporellum Tassi (1)
Camarosporiopsis Abbas, B. Sutton & Ghaffar (1)
Camposporidium Nawawi & Kuthub. (3)
Candelabrum Beverw. (7)
Candelosynnema K.D. Hyde & Seifert (1)
Capitorostrum Bat. (1)
Capnocheirides J.L. Crane & S. Hughes (1)
Capnofrasera S. Hughes (1)
Capsicumyces Gamundí et al. (1)
Carnegieispora Etayo & F. Berger (1)
Carnia Bat. (1)
Carrismyces R.F. Castañeda & Heredia (1)
Casaresia Gonz. Frag. (1)
Castanedaea W.A. Baker & Partr. (1)
Catenocuneiphora Matsush. (1)
Catenophora Luttr. (3)
Catenophoropsis Nag Raj & W.B. Kendr. (1)
Catenosubulispora Matsush. (1)
Catenosynnema Kodsueb, K.D. Hyde & W.H. Ho (1)
Catenulaster Bat. & C.A.A. Costa (1)
Catinopeltis Bat. & C.A.A. Costa (1)
Cecidiomyces U. Braun & Zhurb. (1)
Ceeveesubramaniomyces J. Pratibha, K.D. Hyde & Bhat (1)
Ceratocladium Corda (2)
Ceratophorum Sacc. (2)
Ceratopycnis Höhn. (2)
Ceratosporella Höhn. (19)
Ceratosporium Schwein. (11)
Ceuthodiplospora Died. (1)
Ceuthosira Petr. (1)
Ceuthosporella Petr. & Syd. (1)
Chaetendophragmia Matsush. (7)
Chaetoblastophorum Morgan-Jones (1)
Chaetochalara B. Sutton & Piroz. (7)
Chaetocytostroma Petr. (1)
Chaetodiplis Clem. (1)
Chaetodiplodina Speg. (2)
Chaetopeltaster Katum. (1)
Chaetophiophoma Speg. (1)
Chaetoplaca Syd. & P. Syd. (1)
Chaetopsis Grev. (7)
Chaetopyrena Pass. (2)
Chaetoseptoria Tehon. (1)

1245
Chalarodendron C.J.K. Wang & B. Sutton (1)
Chalarodes McKenzie (2)
Chantransiopsis Thaxt. (3)
Characonidia Bat. & Cavalc. (1)
Charomyces Seifert (2)
Chasakopama Manohar., Bagyan., N.K. Rao & Kunwar (1)
Cheilaria Lib. (1)
Cheiroidea W.A. Baker & Morgan-Jones (1)
Cheiromycella Höhn. (3)
Cheiromyceopsis Mercado & J. Mena (1)
Cheiromyces Berk. & M.A. Curtis (6)
Cheiropolyschema Matsush. (2)
Chiastospora Riess (1)
Chithramia Nag Raj (1)
Chlamydopsis Hol.-Jech. & R.F. Castañeda (1)
Choanatiara DiCosmo (2)
Choreospora Constant. & R. Sant. (1)
Chrysachne Cif. (2)
Chrysalidopsis Steyaert (1)
Chryseidea Onofri (1)
Ciferria Gonz. Frag. (1)
Ciferrina Petr. (1)
Ciferriopeltis Bat. & H. Maia (1)
Ciferrioxyphium Bat. & H. Maia (2)
Ciliochora Höhn. (2)
Ciliophora Petr. (2)
Ciliophorella Petr. (2)
Ciliosporella Petr. (2)
Circinoconiopsis A. Hern.-Gut. (1)
Circinoconis Boedijn (1)
Cissococcomyces Brain (1)
Civisubramaniania Vittal & Dorai (2)
Cladoconidium Bandoni & Tubaki (1)
Cladoniicola Diederich, van den Boom & Aptroot (2)
Cladosphaera Dumort. (1)
Cladosporiopsis S.C. Ren & X.G. Zhang (1)
Clasteropycnis Bat. & Cavalc. (1)
Clathroconium Samson & H.C. Evans (2)
Clauzadeomyces Diederich (1)
Clavariana Nawawi (1)
Cleistocystis Sousa da Câmara (1)
Cleistonium Speer (1)
Cleistophoma Petr. & Syd. (2)
Clypeochorella Petr. (1)
Clypeolum Speg. (8)
Clypeopatella Petr. (1)
Clypeophialophora Bat. & Peres (1)
Clypeopycnis Petr. (3)
Clypeoseptoria F. Stevens & P.A. Young (3)
Clypeostagonospora Punith. (1)
Coccogloeum Petr. (1)
Codonmyces Calat. & Etayo (1)

1246
Colemaniella Agnihothr. (1)
Coleodictyospora Charles (2)
Coleoseptoria Petr. (1)
Colispora Marvanová (3)
Colletoconis de Hoog & Aa (1)
Colletosporium Link (1)
Collostroma Petr. (1)
Columnodomus Petr. (1)
Columnothyrium Bubák (1)
Comatospora Piroz. & Shoemaker (1)
Comocephalum Syd. (1)
Complexipes C. Walker (2)
Condylospora Nawawi (4)
Coniambigua Etayo & Diederich (1)
Conioscyphopsis Goh & K.D. Hyde (1)
Coniothyrina Syd. (1)
Conjunctospora Udagawa & Uchiy. (1)
Conostoma Bat. & J.L. Bezerra (2)
Conostroma Moesz (3)
Consetiella Hol.-Jech. & Mercado (1)
Coremiella Bubák & K. Krieg. (1)
Cornucopiella Höhn. (2)
Cornutispora Piroz. (9)
Cornutostilbe Seifert (1)
Coronospora M.B. Ellis (4)
Corynecercospora V.K. Pal, M. Akhtar, N. Ahmad, Kamal & D.K. Agarwal (1)
Coryneliella Har. & P. Karst. (1)
Corynesporella Munjal & H.S. Gill (11)
Corynesporina Subram. (1)
Corynesporopsis P.M. Kirk (16)
Costanetoa Bat. & J.L. Bezerra (1)
Crandallia Ellis & Sacc. (4)
Craneomyces Morgan-Jones, R.C. Sinclair & Eicker (1)
Craspedodidimella F.R. Barbosa, R.F. Castañeda & Gusmão (1)*
Creodiplodina Petr. (1)
Creonecte Petr. (1)
Creoseptoria Petr. (1)
Creothyriella Bat. & C.A.A. Costa (1)
Cribropeltis Tehon (1)
Crinigera I. Schmidt (1)
Crousobrauniella Sh. Kumar, Raghv. Singh, D.P. Singh & Kamal (1)
Crustodiplodina Punith. (1)
Cryptoceuthospora Petr. (2)
Cryptocoryneopsis B. Sutton (1)
Cryptosporium Kunze (25)
Cryptumbellata Udagawa & Uchiy. (1)
Ctenosporium R. Kirschner (1)
Cubasina R.F. Castañeda (2)
Culicidospora R.H. Petersen (2)
Culicinomyces Couch, Romney & B. Rao (3)
Curucispora Matsush. (3)
Curvulariopsis M.B. Ellis (1)

1247
Cyanopatella Petr. (1)
Cyanopyrenia Harada (1)
Cyclomarsonina Petr. (1)
Cylindrogloeum Petr. (1)
Cylindromyces Manohar., D.K. Agarwal & N.K. Rao (1)
Cylindrothyrium Maire (1)
Cylindroxyphium Bat. & Cif. (1)
Cyrtidium Vain (1)
Cyrtidula Minks (ca. 5)
Cyrtopsis Vain. (1)
Cystodium Fée (1)
Cystotricha Berk. & Broome (1)
Cytodiscula Petr. (1)
Cytogloeum Petr. (1)
Cytonaema Höhn. (2)
Cytoplacosphaeria Petr. (2)
Cytosphaera Died. (2)
Cytosporella Sacc. (32)
Cyttariella Palm (1)
Dactylifera Alcorn (1)
Dactylosporium Harz (2)
Dasysticta Speg. (2)
Davisiella Petr. (2)
Dearnessia Bubák (1)
Deichmannia Alstrup & D. Hawksw. (1)
Delortia Pat. & Gaillard (3)
Dendrodomus Bubák (1)
Dendrographiella Agnihothr. (1)
Dendrographium Massee (8)
Dendrospora Ingold (10)
Dendrosporium Plakidas & Edgerton ex J.L. Crane (2)
Dendryphiosphaera Lunghini & Rambelli (4)
Dennisographium Rifai (2)
Denticularia Deighton (7)
Dentocircinomyces R.F. Castañeda & W.B. Kendr. (1)
Descalsia A. Roldán & Honrubia (1)
Desertella Mouch. (2)
Desmidiospora Thaxt. (3)
Dexhowardia J.J. Taylor (1)
Diaboliumbilicus I. Hino & Katum. (1)
Diademospora B.E. Söderstr. & Bååth (1)
Diarimella B. Sutton (3)
Dichelostroma Bat. & Peres (1)
Dicholobodigitus G.P. White & Illman (1)
Dichotomophthoropsis M.B. Ellis (2)
Dichotophora Whitton, K.D. Hyde & McKenzie (2)
Dictyoceratosporella Y.R. Ma & X.G. Zhang (3)*
Dictyodesmium S. Hughes (4)
Dictyophrynella Bat. & Cavalc. (1)
Dictyopolyschema M.B. Ellis (1)
Dictyorostrella U. Braun (1)
Dictyospiropes M.B. Ellis (1)

1248
Dictyotrichocladium Fiuza, Gusmão & R.F. Castañeda (1)
Didymochaetina Bat. & J.L. Bezerra (1)
Didymochora Höhn. (1)
Didymopsis Sacc. & Marchal (5)
Didymosporina Höhn. (1)
Diedickea Syd. & P. Syd. (3)
Digicatenosporium S.M. Leão, Gusmão & R.F. Castañeda (1)
Digitodochium Tubaki & Kubono (1)
Digitopodium U. Braun et al. (1)
Digitoramispora R.F. Castañeda & W.B. Kendr. (4)
Dimastigosporium Faurel & Schotter (2)
Diplocladiella G. Arnaud ex M.B. Ellis (8)
Diplodinis Clem. (1)
Diplodinula Tassi (1)
Diploplenodomus Died. (2)
Diplosporonema Höhn. (1)
Diplozythiella Died. (1)
Dipyrgis Clem. (1)
Discogloeum Petr. (1)
Discomycetoidea Matsush. (1)
Discosiellina Subram. & K.R.C. Reddy (1)
Discosporina Höhn. (1)
Discotheciella Syd. & P. Syd (1)
Discozythia Petr. (1)
Dissitimurus E.G. Simmons, McGinnis & Rinaldi (1)
Distophragmia R.F. Castañeda, S.M. Leão & Gusmão (1)
Ditangifibula G.C. Adams (1)
Domingoella Petr. & Cif. (4)
Dothideodiplodia Murashk. (1)
Dothioropsis Riedl (1)
Drepanospora Berk. & M.A. Curtis (1)
Drudeola Kuntze (1)
Drumopama Subram. (1)
Dryosphaera Jørg. Koch & E.B.G. Jones (3)
Dualomyces Matsush. (2)
Dwayabeeja Subram. (3)
Dwayaloma Subram. (1)
Dwayalomella Brisson, Piroz. & Pauzé (1)
Dwibahubeeja N. Srivast., A.K. Srivast. & Kamal (1)
Dwibeeja Subram. (1)
Dwiroopella Subram. & Muthumary (1)
Ebollia Minter & Caine (1)
Echinocatena R. Campb. & B. Sutton (1)
Echinochondrium Samson & Aa (1)
Echinoconidiophorum Pereira-Carv. & Dianese (1)
Eiona Kohlm. (1)
Elachopeltella Bat. & Cavalc. (2)
Elattopycnis Bat. & Cavalc. (1)
Elegantimyces Goh, C.K.M. Tsui & K.D. Hyde (1)
Elletevera Deighton (2)
Ellisembiopsis T.S. Santa Izabel & Gusmão (2)
Ellismarsporium R.F. Castañeda & X.G. Zhang (7)

1249
Elotespora R.F. Castañeda & Heredia (1)
Embryonispora G.Z. Zhao (1)
Enantioptera Descals (2)
Endobotrya Berk. & M.A. Curtis (1)
Endobotryella Höhn. (1)
Endocolium Syd. (1)
Endoconospora Gjaerum (2)
Endocoryneum Petr. (3)
Endogenospora R.F. Castañeda, O. Morillo & Minter (1)
Endomelanconium Petr. (4)
Endophragmiopsis M.B. Ellis (2)
Endoplacodium Petr. (1)
Endoramularia Petr. (1)
Endosporoideus W.H. Ho, Yanna, K.D. Hyde & Goh (1)
Endozythia Petr. (1)
Enerthidium Syd. (1)
Engelhardtiella A. Funk (1)
Enridescalsia R.F. Castañeda & Guarro (1)
Enthallopycnidium F. Stevens (1)
Entoderma Hanula, Andreadis & M. Blackw. (1)
Epaphroconidia Calat. & V. Atienza (1)
Ephelidium C.W. Dodge & E.D. Rudolph (1)
Epiclinium Fr. (2)
Epicoccospora Budathoki & S.K. Singh (2)
Episporogoniella U. Braun (1)
Epistigme Syd. (2)
Epithyrium (Sacc.) Trotter (2)
Eriocercospora Deighton (3)
Eriocercosporella Rak. Kumar, A.N. Rai & Kamal ex U. Braun (2)
Eriospora Berk. & Broome (1)
Erispora Pat. (1)
Esteya J.Y. Liou, J.Y. Shih & Tzean (1)
Evanidomus Caball. (1)
Everhartia Sacc. & Ellis (6)
Everniicola D. Hawksw. (1)
Eversia J.L. Crane & Schokn. (2)
Excipularia Sacc. (2)
Exophoma Weedon (1)
Exosporella Höhn. (1)
Exosporodiella Ganie, Azam & A.H. Wani (1)
Fairmaniella Petr. & Syd. (1)
Farriolla Norman (1)
Favostroma B. Sutton & E.M. Davison (1)
Feltgeniomyces Diederich (4)
Fenestroconidia Calat. & Etayo (1)
Fissuricella Pore, D'Amatao & Ajello (1)
Flabellocladia Nawawi (2)
Flabellospora Alas. (6)
Flosculomyces B. Sutton (2)
Frigidispora K.D. Hyde & Goh (1)
Fujimyces Minter & Caine (2)
Fuligomyces Morgan-Jones & Kamal (4)

1250
Fumagopsis Speg. (2)
Furcaspora Bonar (2)
Fusamen (Sacc.) P. Karst. (2)
Fuscophialis B. Sutton (4)
Fusticeps J. Webster & R.A. Davey (5)
Gaeumanniella Petr. (1)
Gallaicolichen Serux. & Lücking (1)
Gampsonema Nag Raj (1)
Gangliophora Subram. (1)
Gangliostilbe Subram. & Vittal (5)
Garnaudia Borowska (3)
Gaubaea Petr. (2)
Gelatinocrinis Matsush. (1)
Gelatinopycnis Dyko & B. Sutton (1)
Geminoarcus K. Ando (3)
Gemmulina Descals & Marvanová (1)
Gilmaniella G.L. Barron (9)
Glaphyriopsis B. Sutton & Pascoe (2)
Glioannellodochium Matsush. (1)
Glioblastocladium Matsush. (1)
Globoconidiopsis G.F. Sepúlveda, Pereira-Carv. & Dianese (1)
Globoconidium G.F. Sepúlveda, Pereira-Carv. & Dianese (1)
Gloeocoryneum Weindlm. (3)
Gloeodes Colby (1)
Gloeosporiella Cavara (1)
Gloiosphaera Höhn. (2)
Glutinium Fr. (2)
Goidanichiella G.L. Barron ex W. Gams (5)
Gonatobotryum Sacc. (4)
Goniopila Marvanová & Descals (1)
Goosiella Morgan-Jones, Kamal & R.K. Verma (1)
Goosiomyces N.K. Rao & Manohar. (2)
Grallomyces F. Stevens (1)
Graphiothecium Fuckel (6)
Groveolopsis Boedijn (6)
Guarroa M. Calduch, Gené, Heredia & R.F. Castañeda (1)
Guedea Rambelli & Bartoli (3)
Guelichia Speg. (6)
Gymnoxyphium Cif., Bat. & I.J. Araújo (6)
Gyrophthorus Hafellner & Sancho (3)
Hadronema Syd. & P. Syd. (4)
Hadrosporium Syd. (2)
Halysiomyces E.G. Simmons (1)
Hansfordiopeltis Bat. & C.A.A. Costa (5)
Hansfordiopeltopsis M.L. Farr (1)
Hapalosphaeria Syd. (1)
Haplariopsis Oudem. (2)
Haplobasidion Erikss. (3)
Haplolepis Syd. (3)
Haptocara Drechsler (1)
Harmoniella V.N. Boriss. (2)*
Harpographium Sacc. (5)

1251
Harpostroma Höhn. (1)
Hawksworthiana U. Braun (4)
Heimiodora Nicot (1)
Helensiella Minter, R.F. Castañeda & Heredia (1)
Helhonia B. Sutton (1)
Helicofilia Matsush. (2)
Helicogoosia Hol.-Jech. (1)
Helicominopsis Deighton (2)
Helicorhoidion S. Hughes (6)
Helicosingula P.S. van Wyk, Marasas, Baard & Knox-Dav. (1)
Helicothyrium I. Hino & Katum. (1)
Helicoubisia Lunghini & Rambelli (1)
Heliscella Marvanová (2)
Heliscina Marvanová (2)
Helminthosporiomyces G.F. Sepúlveda, Pereira-Carv. & Dianese (1)
Helochora Sherwood (1)
Hemicorynesporella Subram. (1)
Hemidothis Syd. & P. Syd. (1)
Hemisphaeropsis Petr. (1)
Hendersoniella Tassi (1)
Hendersonina E.J. Butler (1)
Hendersoniopsis Höhn. (1)
Hendersonula Speg. (20)
Hendersonulina Petr. (1)
Henfellra Halici, D. Hawksw., Z. Kocak. & M. Kocak (1)
Henicospora P.M. Kirk & B. Sutton (6)
Herposira Syd. (1)
Herreromyces R.F. Castañeda & W.B. Kendr. (1)
Heterocephalum Thaxt. (2)
Heterosporiopsis Petr. (1)
Heuflera Bail (1)
Hexacladium D.L. Olivier (1)
Himantia Pers. (4)
Hinoa Hara & I. Hino (2)
Hirudinaria Ces. (2)
Hobsoniopsis D. Hawksw. (1)
Hoehneliella Bres. & Sacc. (2)
Holubovaea Mercado (2)
Homalopeltis Bat. & Valle (1)
Hoornsmania Crous (1)
Hormiactis Preuss (5)
Hormiscioideus M. Blackw. & Kimbr. (1)
Hormocephalum Syd. (1)
Hormographis Guarro, Punsola & Arx (1)
Hughesinia J.C. Lindq. & Gamundí (3)
Hyalobelemnospora Matsush. (1)
Hyalocamposporium Révay & J. Gönczöl (4)
Hyalocephalotrichum Nagaraju, Kunwar, Sureshk. & Manohar. (1)
Hyalocladium Mustafa (1)
Hyalocylindrophora J.L. Crane & Dumont (3)
Hyalodermella Speg. (1)
Hyalodictyum Woron. (1)

1252
Hyalohelicomina T. Yokoy. (1)
Hyalopleiochaeta R.F. Castañeda, Guarro & Cano (1)
Hyalopyrenia H. Harada (1)
Hyalosynnema Matsush. (1)
Hyalothyridium Tassi (1)
Hydrometrospora J. Gönczöl & Révay (1)
Hymenella Fr. (11)
Hymeniopeltis Bat. (3)
Hymenobactron (Sacc.) Höhn.
Hymenobia Nyl. (1)
Hymenopsis Sacc. (13)
Hyphodiscosia Lodha & K.R.C. Reddy (5)
Hyphodiscosioides Matsush. (1)
Hyphopolynema Nag Raj (6)
Hyphostereum Pat. (1)
Hyphothyrium B. Sutton & Pascoe (1)
Hyphozyma de Hoog & M.T. Sm. (4)
Hypnotheca Tommerup (1)
Hypocline Syd. (1)
Hypodermina Höhn. (1)
Hypogloeum Petr. (1)
Hypotrachynicola Etayo (1)
Hysteridium P. Karst. (1)
Hysterodiscula Petr. (1)
Hysteropycnis Hilitzer (1)
Ialomitzia Gruia (1)
Idiocercus B. Sutton (2)
Igneocumulus A.W. Ramaley (10)
Imicles Shoemaker & Hambl. (6)
Impudentia Vujanović (1)
Inesiosporium R.F. Castañeda & W. Gams (2)
Inifatiella R.F. Castañeda (1)
Intercalarispora J.L. Crane & Schokn. (1)
Intralichen D. Hawksw. & M.S. Cole (4)*
Ionophragmium Peres (1)
Irpicomyces Deighton (3)
Isariella Henn. (2)
Ischnostroma Syd. & P. Syd. (1)
Isthmoconidium Etayo & Fr. Berger (1)
Isthmolongispora Matsush. (11)
Isthmophragmospora Kuthub. & Nawawi (2)
Isthmotricladia Matsush. (3)
Ityorhoptrum P.M. Kirk (4)
Iyengarina Subram. (3)
Jahniella Petr. (3)
Javonarxia Subram. (2)
Jayarambhatia J. Pratibha (1)
Jerainum Nawawi & Kuthub. (1)
Jubispora B. Sutton & H.J. Swart (1)
Junctospora Minter & Hol.-Jech. (1)
Kalamarospora G. Delgado (1)
Kalchbrenneriella Diederich & M.S. Christ. (1)

1253
Kaleidosporium Van Warmelo & B. Sutton (1)
Kamatella Anahosur (1)
Kamatia V.G. Rao & Subhedar (1)
Kameshwaromyces Kamal, R.K. Verma & Morgan-Jones (2)
Katherinomyces Khodos. (1)
Keissleriomyces D. Hawksw. (1)
Kendrickiella K. Jacobs & M.J. Wingf. (1)
Ketubakia Kamat, Varghese & V.G. Rao (1)
Kiliophora Kuthub. & Nawawi (3)
Kionocephala P.M. Kirk (1)
Kmetia Bres. & Sacc. (1)
Kmetiopsis Bat. & Peres (1)
Knemiothyrium Bat. & J.L. Bezerra (1)
Kodonospora K. Ando (1)
Kolletes Kohlm. & Volkm.-Kohlm. (1)
Kontospora A. Roldán et al. (1)
Korunomyces Hodges & F.A. Ferreira (3)
Kostermansinda Rifai (4)
Kostermansindiopsis R.F. Castañeda (1)
Kramabeeja G.V. Rao & K.A. Reddy (1)
Kramasamuha Subram. & Vittal (1)
Kreiseliella Braun (1)
Kumanasamuha P. Rag. Rao & D. Rao (5)
Kutilakesa Subram. (2)
Kyphophora B. Sutton (1)
Lacellina Sacc. (3)
Lacellinopsis Subram. (3)
Laciniocladium Petri (1)
Lagenomyces Cavalc. & A.A. Silva (1)
Lambdasporium Matsush. (3)
Lambinonia Sérus. & Diederich (1)
Laocoön J.C. David (1)
Lappodochium Matsush. (1)
Lasiodiplodiella Zambett. (3)
Lasiothyrium Syd. & P. Syd. (1)
Lasmeniella Petr. & Syd. (13)
Latericonis G.V. Rao, K.A. Reddy & de Hoog (1)
Lateriramulosa Matsush. (5)
Laterispora Uecker, W.A. Ayers & P.B. Adams (1)
Lawalreea Diederich (1)
Lecaniocola Brain (1)
Lecanostictopsis B. Sutton & Crous (4)
Leeina Petr. (1)
Leightoniomyces D. Hawksw. & B. Sutton (2)
Lembuncula Cif. (1)
Lemkea Morgan-Jones & R.C. Sinclair (1)
Lepisticola W. Gams (1)
Leprieurinella Bat. & H. Maia (1)
Leptascospora Speg. (1)
Leptochlamys Died. (1)
Leptodermella Höhn. (1)
Leptophyllosticta I.E. Brezhnev (2)

1254
Leptostromella (Sacc.) Sacc. (2)
Leptothyrella Sacc. (10)
Leptothyrina Höhn. (1)
Leptothyrium Kunze (2)
Leucoconiella Bat., H. Maia & Peres (1)
Leucoconis Theiss. & Syd. (1)
Leucodochium Syd. & P. Syd. (1)
Leuliisinea Matsush. (2)
Lichenobactridium Diederich & Etayo (1)
Lichenohendersonia Calat. & Etayo (3)
Lichenopeziza Zukal (1)
Lichenophoma Keissl. (2)
Lichenopuccinia D. Hawksw. & Hafellner (1)
Lichenostella Calat. & Etayo (1)
Linkosia A. Hern. Gut. & B. Sutton (12)
Linochorella Syd. & P. Syd. (1)
Linodochium Höhn. (5)
Listeromyces Penz. & Sacc. (1)
Lithopythium Bornet & Flahault (3)
Lobatopedis P.M. Kirk (5)
Loliomyces Maire (1)
Lomaantha Subram. (3)
Lomachashaka Subram. (5)
Ludwigomyces Kirschst. (1)
Luxuriomyces R.F. Castañeda (1)
Luzfridiella R.F. Castañeda & W.B. Kendr. (1)
Lylea Morgan-Jones (6)
Lysotheca Cif. (6)
Mackenziella Yanna & K.D. Hyde (1)
Macroallantina Speer (1)
Macrodiplodia Sacc. (2)
Macrotrichum Grev. (2)
Magmopsis Nyl. (1)
Mahabalella B. Sutton & S.D. Patil (4)
Manginella Bat. & H. Maia (2)
Mapletonia B. Sutton (1)
Margarinomyces Laxa (1 fide Kirk et al. 2008)
Martinellisia V.G. Rao & Varghese (1)
Massalongina Bubák (2)
Matsushimiella R.F. Castañeda & Heredia (2)
Matsushimomyces V.G. Rao & Varghese (2)
Medusamyces G.L. Barron & Szijarto (1)
Megalodochium Deighton (4)
Megaloseptoria Naumov (1)
Melanocephala S. Hughes (5)
Melanophoma Papendorf & J.W. du Toit (1)
Melophia Sacc. (4)
Menidochium R.F. Castañeda & W.B. Kendr. (1)
Mercadomyces J. Mena (1)
Merismella Syd. (6)
Metadiplodia Syd. (40)
Metazythia Petr. (1)

1255
Metazythiopsis M. Morelet (1)
Microblastosporon Cif. (1)
Microclava F. Stevens (5)
Microdiscula Höhn. (2)
Microdothiorella C.A.A. Costa & Sousa da Câmara (1)
Microhendersonula Dias & Sousa da Câmara (1)
Micromastia Speg. (2)
Microperella Höhn. (1)
Micropustulomyces R.W. Barreto (1)
Microtyle Speg. (1)
Microxyphiella Speg. (15)
Microxyphiopsis Bat. (2)
Mindoa Petr. (2)
Minimidochium B. Sutton (8)
Minteriella Heredia, R.F. Castañeda & R.M. Arias (1)
Minutophoma D. Hawksw. (1)
Mirandina G. Arnaud ex Matsush. (ca. 10)
Miricatena Punith. & Spooner (2)
Mirimyces Nag Raj (1)
Monochaetiella E. Castell. (3)
Monochaetinula Muthumary, Abbas & B. Sutton (6)
Monochaetopsis Pat. (1)
Monodia Breton & Faurel (2)
Monodidymaria U. Braun (5)
Monodisma Alcorn (1)
Monostichella Höhn. (15)
Moorella P. Rag. Rao & D. Rao (3)
Moralesia Urries (1)
Morrisographium M. Morelet (8)
Mucosetospora M. Morelet (1)
Muiogone Thaxt. (2)
Muirella R. Sprague (1)
Murogenella Goos & E.F. Morris (3)
Mycelephas R.F. Castañeda (2)
Mycocentrodochium K. Matsush. & Matsush. (1)
Mycoënterolobium Goos (3)
Mycohypallage B. Sutton (2)
Mycopara Bat. & J.L. Bezerra (1)
Mycospraguea U. Braun & Rogerson (1)
Mycosticta Höhn. (1)
Mycosylva M.C. Tulloch (3)
Mycotodea Kirschst. (14)
Mycousteria M.L. Farr (2)
Myiocoprula Petr. (2)
Myriellina Höhn. (2)
Myrmecomyces Jouvenaz & Kimbr. (1)
Myrotheciastrum Abbas & B. Sutton (1)
Mystrosporiella Munjal & Kulshr. (4)
Myxoparaphysella Caball. (2)
Myxosporella Sacc. (1)
Myxosporidiella Negru (1)
Myxostomellina Syd. (1)

1256
Myxothyriopsis Bat. & A.F. Vital (1)
Myxothyrium Bubák & Kabát (1)
Naemosphaera P. Karst. (1)
Naemosphaerella Höhn. (2)
Nagrajia R.F. Castañeda & W.B. Kendr. (1)
Nagrajomyces Mel’nik (1)
Nakatopsis Whitton, McKenzie & K.D. Hyde (2)
Nanoschema B. Sutton (1)
Naothyrsium Bat. (1)
Necraphidium Cif. (1)
Nematogonum Desm. (1)
Nematographium Goid. (5)
Nemozythiella Höhn. (1)
Neoalpakesa Punith. (1)
Neoarbuscula B. Sutton (1)
Neobarclaya Sacc. (2)
Neodiplodina Petr. (1)
Neofuckelia Zeller & Goodd. (1)
Neoheteroceras Nag Raj (2)
Neojohnstonia B. Sutton (2)
Neoligniella Naumov (4)
Neomarssoniella U. Braun (1)
Neomelanconium Petr. (3)
Neoovularia U. Braun (6)
Neopeltis Syd. (3)
Neopericonia Kamal, A.N. Rai & Morgan-Jones (1)
Neophoma Petr. & Syd. (2)
Neoplaconema B. Sutton (2)
Neopodoconis Rifai (3)
Neoramularia U. Braun (9)
Neospegazzinia Petr. & Syd. (2)
Neottiospora Desm. (2)
Neozythia Petr. (1)
Neta Shearer & J.L. Crane (10)
Nidulispora Nawawi & Kuthub. (1)
Nigrolentilocus R.F. Castañeda & Heredia (6)
Nigromacula Etayo (1)
Nigropuncta D. Hawksw. (2)
Nosophloea Fr. (3)
Nothospora Peyronel (1)
Novozymia W.P. Wu (1)
Nummospora E. Müll. & Shoemaker (1)
Nusia Subram. (2)
Nyctalospora E.F. Morris (1)
Nypaella K.D. Hyde & B. Sutton (2)
Obeliospora Nawawi & Kuthub. (5)
Obstipipilus B. Sutton (1)
Octopodotus Kohlm. & Volkm.-Kohlm. (1)
Odontodictyospora Mercado (1)
Oedothea Syd. (1)
Ojibwaya B. Sutton (1)
Omega B. Sutton & Minter (1)

1257
Oncopodium Sacc. (12)
Oncospora Kalchbr. (8)
Oncosporella P. Karst. (1)
Oncostroma Bat. & Marasas (1)
Onychophora W. Gams, P.J. Fisher & J. Webster (1)
Oothyrium Syd. (1)
Ophiosira Petr. (1)
Orphanocoela Nag Raj (3)
Ostracoderma Fr. (3)
Ostracodermidium Mukerji (1)
Oswaldina Rangel (1)
Paathramaya Subram. (5)
Pachycladina Marvanová (3)
Palawaniopsis Bat., Cif. & Nascim. (1)
Papilionospora V.G. Rao & B. Sutton (1)
Pappimyces B. Sutton & Hodges (1)
Paraaoria R.K. Verma & Kamal (1)
Paraarthrocladium Matsush. (1)
Parablastocatena Y.D. Zhang & X.G. Zhang (1)
Paraceratocladium R.F. Castañeda (6)
Parachionomyces Thaung (1)
Paracostantinella Subram. & Sudha (1)
Paracryptophiale Kuthub. & Nawawi (2)
Paracytospora Petr. (1)
Paradendryphiopsis M.B. Ellis (5)
Paradidymobotryum C.J.K. Wang & B. Sutton (1)
Paradiplodia Speg. ex Trotter (6)
Paradischloridium Bhat & B. Sutton (1)
Paradiscula Petr. (1)
Paraëpicoccum Matsush. (1)
Parafulvia Kamal, A.N. Rai & Morgan-Jones (1)
Parahaplotrichum W.A. Baker & Partr. (1)
Paraharknessia Matsush. (1)
Parahyalotiopsis Nag Raj (1)
Paramassariothea Subram. & Muthumary (1)
Paramenisporopsis Matsush. (1)
Parapericonia M.B. Ellis (2)
Parapericoniella U. Braun, Heuchert & K. Schub. (1)
Paraphaeoisaria de Hoog & Morgan-Jones (1)
Parapithomyces Thaung (1)
Parapyricularia M.B. Ellis (4)
Pararobillarda Matsush. (1)
Parasphaeropsis Petr. (1)
Parastigmatellina Bat. & C.A.A. Costa (1)
Paratetraploa M.K.M. Wong & K.D. Hyde (1)
Paratomenticola M.B. Ellis (2)
Paratrichoconis Deighton & Piroz. (4)
Paraulocladium R.F. Castañeda (2)
Paspalomyces Linder (1)
Patriciomyces D. Hawksw. (1)
Pazschkeella Syd. & P. Syd.
Peethasthabeeja P. Rag. Rao (1)

1258
Pellionella (Sacc.) Sacc. (1)
Peltasterinostroma Punith. (1)
Peltasteropsis Bat. & H. Maia (7)
Peltistroma Henn. (1)
Peltistromella Höhn. (1)
Peltosoma Syd. (1)
Peltostromellina Bat. & A.F. Vital (1)
Peltostromopsis Bat. & A.F. Vital (1)
Penzigomyces Subram. (13)
Perelegamyces R.F. Castañeda & W.B. Kendr. (1)
Perizomella Syd. (1)
Pestalozziella Sacc. & Ellis ex Sacc. (4)
Petrakiopsis Subram. & K.R.C. Reddy (1)
Phacostroma Petr. (1)
Phacostromella Petr. (1)
Phaeoblastophora Partr. & Morgan-Jones (2)
Phaeocandelabrum R.F. Castañeda, Gusmão, Guarro & Iturr. (3)
Phaeodactylium Agnihothr. (7)
Phaeodiscula Cub. (1)
Phaeodomus Höhn. (3)
Phaeohiratsukaea Udagawa & Iwatsu (1)
Phaeoidiomyces Dorn.-Silva & Dianese (2)
Phaeolabrella Speg. (1)
Phaeomonilia R.F. Castañeda, Heredia & R.M. Arias (5)
Phaeomonostichella Keissl. ex Petr. (1)
Phaeophloeosporella Crous & B. Sutton (1)
Phaeophomopsis Höhn. (1)
Phaeoschizotrichum C.R. Silva, Gusmão & R.F. Castañeda (1)
Phaeostalagmus W. Gams (7)
Phaeostilbelloides Armando, Z.M. Chaves & Dianese (1)
Phaeothyrium Petr. (1)
Phaeotrichoconis Subram. (8)
Phaeoxyphiella Bat. & Cif. (7)
Phellostroma Syd. & P. Syd. (1)
Phialoarthrobotryum Matsush. (2)
Phialogeniculata Matsush. (4)
Phialophaeoisaria Matsush. (1)
Phialostele Deighton (1)
Phialotubus R.Y. Roy & Leelav. (1)
Phloeosporina Höhn. (1)
Phlyctaeniella Petr. (2)
Phomachora Petr. & Syd. (2)
Phomachorella Petr. (1)
Phomatosporella Tak. Kobay. & K. Sasaki (1)
Phomyces Clem. (1)
Phragmoconidium G.F. Sepúlveda, Pereira-Carv. & Dianese (1)
Phragmopeltis Henn. (5)
Phragmospathula Subram. & N.G. Nair (3)
Phragmospathulella J. Mena & Mercado (1)
Phthora d'Hérelle (1)
Phylloedium Fr. (1)
Phyllohendersonia Tassi (25)

1259
Physalidiella Rulamort (2)
Physalidiopsis R.F. Castañeda & W.B. Kendr. (1)
Piggotia Berk. & Broome (3)
Pinatubo J.B. Manandhar & Mew (1)
Piperivora Siboe, P.M. Kirk & P.F. Cannon (1)
Piricauda Bubák (8)
Piricaudilium Hol.-Jech. (2)
Piricaudiopsis J. Mena & Mercado (1)
Pirispora Faurel & Schotter (1)
Pirostomella Sacc. (2)
Pithosira Petr. (1)
Pittostroma Kowalski & T.N. Sieber (1)
Placella Syd. (1)
Placodiplodia Bubák (2)
Placonema (Sacc.) Petr. (3)
Placonemina Petr. (1)
Placosphaerina Maire (1)
Placothea Syd. (1)
Placothyrium Bubák (1)
Plagiostigmella Petr. (1)
Plasia Sherwood (1)
Plectonaemella Höhn. (1)
Plectopeltis Syd. (1)
Plectophomopsis Petr. (1)
Plectopycnis Bat. & A.F. Vital (4)
Plectosira Petr. (1)
Plectronidiopsis Nag Raj (1)
Plectronidium Nag Raj (4)
Plenocatenulis Bat. & Cif. (1)
Plenophysa Syd. & P. Syd. (1)
Plenotrichopsis Bat. (1)
Plenotrichum Syd. (2)
Plenozythia Syd. & P. Syd. (2)
Pleocouturea G. Arnaud (2)
Plesiospora Drechsler (1)
Pleurodesmospora Samson, W. Gams & H.C. Evans (1)
Pleurodiscula Höhn. (1)
Pleurodomus Petr. (1)
Pleuropedium Marvanová & S.H. Iqbal (3)
Pleurophomopsis Petr. (7)
Pleuroplaconema Petr. (2)
Pleuroplacosphaeria Syd. (1)
Pleurostromella Petr. (15)
Pleurotheciopsis B. Sutton (6)
Pleurothyriella Petr. & Syd. (1)
Pleurovularia R. Kirschner & U. Braun (1)
Pocillopycnis Dyko & B. Sutton (1)
Podoplaconema Petr. (1)
Podosporiella Ellis & Everh. (4)
Podosporiopsis Jian Ma, X.G. Zhang & R.F. Castañeda (2)
Podosporium Schwein. (67)
Poikilosperma Bat. & J.L. Bezerra (1)

1260
Polybulbophiale Goh & K.D. Hyde (1)
Polychaetella Speg. (3)
Polycladium Ingold (1)
Polydesmus Mont. (14)
Polyetron Bat. & Peres (1)
Polylobatispora Matsush. (3)
Polyrostrata T.P. Devi & N. Mathur (2)
Polystomellomyces Bat. (1)
Polystratorictus Matsush. (2)
Polytretophora Mercado (3)
Porocladium Descals (1)
Poropeltis Henn. (1)
Porophilomyces U. Braun (1)
Porosubramaniania Hol.-Jech. (2)
Porrectotheca Matsush. (1)
Potamomyces K.D. Hyde (1)
Proboscispora Punith. (1)
Protostegiomyces Bat. & A.F. Vital (1)
Protostroma Bat. (1)
Pseudoacrodictys W.A. Baker & Morgan-Jones (14)
Pseudoanguillospora S.H. Iqbal (3)
Pseudoaristastoma Suj. Singh (1)
Pseudoasperisporium U. Braun (3)
Pseudobasidiospora Dyko & B. Sutton (1)
Pseudocanalisporium R.F. Castañeda & W.B. Kendr. (1)
Pseudocenangium P. Karst. (1)
Pseudochuppia Kamal et al. (1)
Pseudoclathrosphaerina Voglmayr (2)
Pseudoconium Petr. (1)
Pseudocytoplacosphaeria Punith. & Spooner (1)
Pseudocytospora Petr. (1)
Pseudodichomera Höhn. (3)
Pseudodidymaria U. Braun (3)
Pseudodiplodia (P. Karst.) Sacc. (45)
Pseudodiscula Laubert (2)
Pseudofuscophialis Sivan. & H.S. Chang (1)
Pseudogaster Höhn. (1)
Pseudographiella E.F. Morris (3)
Pseudohepatica P.M. Jørg. (1)
Pseudomicrodochium B. Sutton (8)
Pseudoneottiospora Faurel & Schotter (2)
Pseudopatellina Höhn. (1)
Pseudopeltistroma Katum. (1)
Pseudoperitheca Elenkin (1)
Pseudopetrakia M.B. Ellis (2)
Pseudophloeosporella U. Braun (1)
Pseudophragmotrichum W.P. Wu, B. Sutton & Gange (1)
Pseudopolystigmina Murashk. (2)
Pseudoramularia Matsush. (2)
Pseudorhizopogon Kobayasi (1)
Pseudoschizothyra Punith. (1)
Pseudosigmoidea K. Ando & N. Nakam. (2)

1261
Pseudostegia Bubák (1)
Pseudothyrium Höhn. (1)
Pseudotorula Subram. (3)
Pseudotracylla B. Sutton & Hodges (2)
Pseudotrichoconis W.A. Baker & Morgan-Jones (1)
Pseudozythia Höhn. (1)
Psilosphaeria Cooke (1)
Pteromycula P. Cannon (1)
Pterulopsis Wakef. & Hansf. (1)
Pterygosporopsis P.M. Kirk (2)
Pucciniospora Speg. (1)
Pulchromyces Hennebert (1)
Pullospora Faurel & Schotter (2)
Pulvinella A.W. Ramaley (1)
Punctillina Toro (1)
Pycmaeosphaera Etayo & Diederich (3)
Pycnidioarxiella Punith. & N.D. Sharma (1)
Pycnidiopeltis Bat. & C.A.A. Costa (1)
Pycnis Bref. (1)
Pycnodactylus Bat., A.A. Silva & Cavalc. (1)
Pycnodallia Kohlm. & Volkm.-Kohlm. (1)
Pycnoharknessia Matsush. (1)
Pycnomma Syd. (1)
Pycnomoreletia Rulamort (2)
Pycnoseynesia Kuntze (1)
Pycnothera N.D. Sharma & G.P. Agarwal (1)
Pycnothyriella Bat. (2)
Pycnothyrium Diederich (6)
Pyramidospora Sv. Nilsson (9)
Pyrenyllium Clem. (2)
Pyrgostroma Petr. (2)
Pyripnomyces Cavalc. (1)
Quadracaea Lunghini, Pinzari & Zucconi (3)
Quadricladium Nawawi & Kuthub. (1)
Quasidiscus B. Sutton (1)
Quasiphloeospora B. Sutton, Crous & Shamoun (1)
Queenslandia Bat. & H. Maia (5)
Quezelia Faurel & Schotter (1)
Raciborskiomyces Siemaszko (4)
Radiatispora Matsush. (1)
Raizadenia S.L. Srivast. (1)
Ramakrishnanella Kamat & Ullasa ex Ullasa (1)
Ramicapitulum Whitton, K.D. Hyde & McKenzie (1)
Ramicephala Voglmayr & G. Delgado (1)
Ramoconidiifera B. Sutton, Carmarán & A.I. Romero (2)
Redbia Deighton & Piroz. (5)
Refractohilum D. Hawksw. (5)
Repetoblastiella R.F. Castañeda, Minter & M. Stadler (1)
Rhabdoclema Syd. (2)
Rhabdogloeopsis Petr. (2)
Rhabdostromella Höhn. (1)
Rhabdostromina Died. (3)

1262
Rhexoampullifera P.M. Kirk (3)
Rhexoprolifer Matsush. (1)
Rhinotrichella G. Arnaud ex de Hoog (4)
Rhipidocephalum Trail (2)
Rhizosphaerina B. Sutton (2)
Rhodesia Grove (2)
Rhodesiopsis B. Sutton & R. Campb. (2)
Rhodothallus Bat. & Cif. (2)
Rhombostilbella Zimm. (2)
Rhopalocladium Schroers, Samuels & W. Gams (1)
Rhynchodiplodia Briosi & Farneti (1)
Rhynchomyces Willk. (1)
Rhynchoseptoria Unamuno (1)
Rhynchosporina Arx (2)
Riclaretia Peyronel (1)
Rileya A. Funk (1)
Robakia Petr. (1)
Rogergoosiella A. Hern.-Gut. & J. Mena (1)
Roscoepoundia Kuntze (1)
Rosulomyces S. Marchand & Cabral (1)
Rota Bat., Cif. & Nascim. (1)
Ruggieria Cif. & Montemart. (1)
Saania Zhurb. (1)
Sadasivania Subram. (3)
Sanjuanomyces R.F. Castañeda & W.B. Kendr. (1)
Sarcinosporon D.S. King & S.C. Jong (1)
Sarcoexcipula Etayo (1)
Sarcophoma Höhn. (3)
Sarophorum Syd. & P. Syd. (1)
Satchmopsis B. Sutton & Hodges (1)
Sativumoides S.C. Ren, Jian Ma & X.G. Zhang (1)
Scaphidium Clem. (1)
Sceptrifera Deighton (1)
Schizothyra Bat. & C.A.A. Costa (1)
Schizothyrella Thüm. (1)
Schizothyropsis Bat. & A.F. Vital (1)
Schizotrichum McAlpine (1)
Schroeteria G. Winter (1)
Schwarzmannia Pisareva (1)
Scirrhophoma Petr. (1)
Sclerographiopsis Deighton (1)
Sclerographium Berk. (4)
Scleromeris Syd. (3)
Sclerophoma Höhn. (30)
Scleropycnis Syd. & P. Syd. (2)
Sclerozythia Petch (1)
Scolecobasidiella M.B. Ellis (2)
Scolecobeltrania Iturr., R.F. Castañeda & Rob. Fernández (1)
Scolecodochium K. Matsush. & Matsush. (1)
Scolecosporiella Petr. (6)
Scolecotheca Søchting & B. Sutton (1)
Scolecozythia Curzi (1)

1263
Scoliotidium Bat. & Cavalc. (1)
Scopaphoma Dearn. & House (1)
Scopulariella Gjaerum (1)
Scothelius Bat., J.L. Bezerra & Cavalc. (1)
Scutisporus K. Ando & Tubaki (1)
Scutopeltis Bat. & H. Maia (2)
Scutopycnis Bat. (2)
Seimatosporiopsis B. Sutton, Ghaffer & Abbas (2)
Selenosira Petr. (1)
Selenosporopsis R.F. Castañeda & W.B. Kendr. (1)
Semipseudocercospora J.M. Yen (2)
Septocytella Syd. (1)
Septogloeum Sacc. (2)
Septomyxella (Höhn.) Höhn. (1)
Septopatella Petr. (1)
Septosporiopsis W.A. Baker & Morgan-Jones (1)
Septosporium Corda (5)
Septotrullula Höhn. (2)
Sessiliospora D. Hawksw. (1)
Setolibertella Punith. & Spooner (1)
Setophiale Matsush. (1)
Setosporella Mustafa & Abdul-Wahid (1)
Seychellomyces Matsush. (1)
Seynesiopsis Henn. (1)
Shawiella Hansf. (1)
Sheariella Petr. (1)
Sheathnema Dubey & Moonambeth (2)
Shivomyces Hosag. (2)
Siamia V. Robert, Decock & R.F. Castañeda (1)
Sigmatomyces Sacc. & P. Syd. (1)
Simmonsiella J.L. Crane & A.N. Mill. (1)
Sirexcipula Bubák (1)
Sirocyphis Clem. (1)
Sirogloea Petr. (1)
Siroligniella Naumov (1)
Sirophoma Höhn. (3)
Siroplacodium Petr. (6)
Siropleura Petr. (1)
Siroscyphellina Petr. (2)
Sirosperma Syd. & P. Syd. (2)
Sirosphaera Syd. & P. Syd. (2)
Sirosporonaemella Naumov (1)
Sirothecium P. Karst. (3)
Sirothyriella Höhn. (2)
Sirothyrium Syd. & P. Syd. (1)
Sirozythia Höhn. (2)
Sirozythiella Höhn. (1)
Sitochora H.B.P. Upadhyay (1)
Slimacomyces Minter (2)
Soloacrospora W.B. Kendr. & R.F. Castañeda (2)
Solosympodiella Matsush. (8)
Soloterminospora Matsush. (1)

1264
Spermatoloncha Speg. (1)
Spermatoloncha Speg. (1)
Spermochaetella Cif. (1)
Spermospora R. Sprague (9)
Spermosporella Deighton (4)
Sphaeridium Fresen. (5)
Sphaeriostromella Bubák (1)
Sphaeriothyrium Bubák (2)
Sphaeromma H.B.P. Upadhyay (2)
Sphaeronaema Fr. (50)
Sphaerophoma Petr. (2)
Sphaerulomyces Marvanová (1)
Spinulospora Deighton (1)
Spiralum J.L. Mulder (2)
Spiropes Cif. (ca. 40)
Splanchospora Lar.N. Vassiljeva (1)
Spondylocladiella Linder (2)
Spondylocladiopsis M.B. Ellis (2)
Sporhaplus H.B.P. Upadhyay (1)
Sporidesmiopsis Subram. & Bhat (6)
Sporoglena Sacc. (1)
Sporophiala P. Rag. Rao (3)
Sporotretophora Whitton, McKenzie & K.D. Hyde (1)
Stachybotryella Ellis & Barthol. (3)
Stachybotryna Tubaki & T. Yokoy. (6)
Stagonopatella Petr. (1)
Stagonopsis Sacc. (4)
Stagonosporina Tassi (1)
Stagonostromella Petr. & Syd. (1)
Staheliella Emden (2)
Stalagmochaetia Cif. & Bat. (2)
Stanhughesiella R.F. Castañeda & D.W. Li (1)
Stauronema (Sacc.) Syd., P. Syd. & E.J. Butler (5)
Stauronematopsis Abbas, B. Sutton & Ghaffar (1)
Staurophoma Höhn. (1)
Stegonsporiopsis Van Warmelo & B. Sutton (1)
Stellifraga Alstrup & Olech (1)
Stellomyces Morgan-Jones, R.C. Sinclair & Eicker (2)
Stellopeltis Bat. & A.F. Vital (2)
Stellospora Alcorn & B. Sutton (2)
Stellothyriella Bat. & Cif. (2)
Stenocephalopsis Chamuris & C.J.K. Wang (1)
Stenocladiella Marvanová & Descals (1)
Stenospora Deighton (1)
Stephembruneria R.F. Castañeda (1)
Stevensonula Petr. (1)
Stictopatella Höhn. (1)
Stictosepta Petr. (1)
Stigmatellina Bat. & H. Maia (1)
Stigmea Fr. (1)
Stigmella Lév. (28)
Stigmopeltis Syd. (2)

1265
Stilbellula Boedijn (1)
Stilbodendron Syd. & P. Syd. (1)
Stilbophoma Petr. (1)
Strasseriopsis B. Sutton & Tak. Kobay. (1)
Stratiphoromyces Goh & K.D. Hyde (2)
Striosphaeropsis Verkley & Aa (1)
Stromatocrea W.B. Cooke (1)
Stromatopogon Zahlbr. (3)
Stromatopycnis A.F. Vital (1)
Stromatostysanus Höhn. (3)
Strongylothallus Bat. & Cif. (1)
Stygiomyces Coppins & S.Y. Kondr. (1)
Stylaspergillus B. Sutton, Alcorn & P.J. Fisher (1)
Subhysteropycnis Wedin & Hafellner (1)
Subicularium M.L. Farr & Goos (1)
Subulispora Tubaki (8)
Suttoniella S. Ahmad (3)
Suttonina H.C. Evans (1)
Syamithabeeja Subram. & Natarajan (1)
Sylviacollaea Cif. (1)
Symphysos Bat. & Cavalc. (1)
Sympodiella W.B. Kendr. (5)
Sympodiocladium Descals (1)
Sympodioclathra Voglmayr (1)
Sympodioplanus R.C. Sinclair & Boshoff (3)
Sympodiosynnema J.W. Xia & X.G. Zhang (1)
Synchronoblastia Uecker & F.L. Caruso (1)
Syncladium Rabenh. (1)
Synnemacrodictys W.A. Baker & Morgan-Jones (1)
Synnemaseimatoides K. Matsush. & Matsush. (1)
Synnematomyces Kobayasi (1)
Synostomina Petr. (1)
Syphosphaera Dumort. (1)
Systremmopsis Petr. (1)
Taeniolina M.B. Ellis (6)
Talekpea Lunghini & Rambelli (1)
Talpapellis Alstrup & M.S. Cole (4)
Tandonea M.D. Mehrotra (1)
Tarsodisporus Bat. & A.A. Silva (1)
Tectacervulus A.W. Ramaley (1)
Telioclipeum Viégas (1)
Temerariomyces B. Sutton (1)
Teratosperma Syd. & P. Syd. (11)
Termitaria Thaxt. (6)
Tetrabrachium Nawawi & Kuthub. (1)
Tetrabrunneospora Dyko (1)
Tetracoccosporium Szabó (4)
Tetrameronycha Speg. ex W. Rossi & M. Blackw. (1)
Tetranacriella Kohlm. & Volkm.-Kohlm. (1)
Tetranacrium H.J. Huds. & B. Sutton (1)
Tetraposporium S. Hughes (2)
Textotheca Matsush. (1)

1266
Thaptospora B. Sutton & Pascoe (3)
Thirumalacharia Rathaiah (1)
Tholomyces Matsush. (1)
Thoracella Oudem. (1)
Thrinacospora Petr. (1)
Thyriostromella Bat. & C.A.A. Costa (1)
Thyrostromella Höhn. (3)
Thyrsidiella Höhn. ex Höhn. (2)
Thyrsidina Höhn. (1)
Tiarosporellivora Punith. (1)
Ticogloea G. Weber et al. (2)
Ticosynnema R.F. Castañeda, Granados & Mardones (1)
Titaea Sacc. (23)
Titaeopsis B. Sutton & Deighton (1)
Titaeospora Bubák (2)
Tomenticola Deighton (1)
Tompetchia Subram. (1)
Toxosporiella B. Sutton (1)
Toxosporiopsis B. Sutton & Sellar (1)
Toxosporium Vuill. (2)
Trematophoma Petr. (2)
Tremellidium Petr. (1)
Tretendophragmia Subram. (1)
Tretocephala Subram. (1)
Tretolylea Cantillo, R.F. Castañeda & Gusmão (1)
Tretospeira Piroz. (1)
Tretovularia Deighton (1)
Tribolospora D.A. Reid (1)
Tricellula Beverw. (8)
Trichobolbus Bat. (1)
Trichobotrys Penz. & Sacc. (4)
Trichoconis Clem. (21)
Trichodiscula Vouaux (1)
Trichodochium Syd. (3)
Trichomatoclava G.F. Sepúlveda, Pereira-Carv. & Dianese (1)
Trichomatomyces Dorn.-Silva & Dianese (1)
Trichomatosphaera Pereira-Carv., G.F. Sepúlveda & Dianese (1)
Trichopeltulum Speg. (1)
Trichoseptoria Cavara (2)
Trichosporiella Kamyschko (4)
Trichosporodochium Dorn.-Silva & Dianese (1)
Trichotheca P. Karst. (1)
Tricladiella K. Ando & Tubaki (1)
Tricladiopsis Descals (2)
Tricladiospora Nawawi & Kuthub. (3)
Tricornispora Bonar (1)
Trifurcospora K. Ando & Tubaki (2)
Trigonosporium Tassi (2)
Tripoconidium Subram. (1)
Triposporina Höhn. (2)
Triramulispora Matsush. (3)
Triscelophorus Ingold (8)

1267
Triscelosporium Nawawi & Kuthub. (1)
Trisulcosporium H.J. Huds. & B. Sutton (1)
Tromeropsis Sherwood (1)
Troposporium Harkn. (1)
Troposporopsis Whitton, McKenzie & K.D. Hyde (2)
Tryblidiopycnis Höhn. (1)
Tryssglobulus B. Sutton & Pascoe (1)
Tuberculispora Deighton & Piroz. (1)
Tunicago B. Sutton & Pollack (2)
Turturconchata J.L. Chen, T.L. Huang & Tzean (2)
Tympanosporium W. Gams (1)
Uberispora Piroz. & Hodges (4)
Ubrizsya Negru (1)
Ulocoryphus Michaelides, L. Hunter & W.B. Kendr. (1)
Umbellidion B. Sutton & Hodges (1)
Uniseta Ciccar. (1)
Urohendersonia Speg. (5)
Urohendersoniella Petr. (1)
Uvarispora Goos & Piroz. (1)
Vagnia D. Hawksw. & Miądl. (1)
Vanakripa Bhat et al. (9)
Vanbeverwijkia Agnihothr. (1)
Vanderystiella Henn. (1)
Vanterpoolia A. Funk (1)
Vasudevella Chona et al. (1)
Velloziomyces Armando, Z.M. Chaves & Dianese (1)
Velutipila D. Hawksw. (1)
Ventrographium H.P. Upadhyay, Cavalc. & A.A. Silva (1)
Venustocephala Matsush. (2)
Venustosynnema R.F. Castañeda & W.B. Kendr. (3)
Veracruzomyces Mercado, Guarro, Heredia & J. Mena (1)
Veramycella G. Delgado (1)
Veramyces Matsush. (1)
Verdipulvinus A.W. Ramaley (1)
Veronaella Subram. & K.R.C. Reddy (1)
Veronidia Negru (1)
Verrucariella S. Ahmad (1)
Verrucaster Tobler (1)
Verrucophragmia Crous, M.J. Wingf. & W.B. Kendr. (1)
Verticicladus Matsush. (3)
Vesicladiella Crous & M.J. Wingf. (1)
Vesiculohyphomyces Armando, Pereira-Carv. & Dianese (1)
Vestigium Piroz. & Shoemaker (2)
Virgariella S. Hughes (11)
Viridiannula Etayo (1)
Vittalia Gaws & Bhat (1)
Vizellopsidites M.A. Khan, M. Bera & Bera (1)
Vouauxiella Petr. & Syd. (3)
Waihonghopes Yanna & K.D. Hyde (1)
Wardinella Bat. & Peres (1)
Websteromyces W.A. Baker & Partr. (2)
Weufia Bhat & B. Sutton (1)

1268
Wolkia Ramsb. (1)
Xenidiocercus Nag Raj (1)
Xenochora Petr. (1)
Xenodomus Petr. (1)
Xenoheteroconium Bhat, W.B. Kendr. & Nag Raj (1)
Xenokylindria DiCosmo, S.M. Berch & W.B. Kendr. (2)
Xenomyxa Syd. (1)
Xenopeltis Syd. & P. Syd. (1)
Xenoplaca Petr. (1)
Xenostroma Höhn. (1)
Xeroconium D. Hawksw. (1)
Xiphomyces Syd. & P. Syd. (2)
Xiuguozhangia K. Zhang, R.F. Castañeda, Jian Ma & L.G. Ma (5)
Xylochia B. Sutton (2)
Xyloglyphis Clem. (1)
Xylohypha (Fr.) E.W. Mason (6)
Xylohyphopsis W.A. Baker & Partr. (3)
Yalomyces Nag Raj (6)
Yinmingella Goh, C.K.M. Tsui & K.D. Hyde (1)
Ypsilomyces D.A.C. Almeida & Gusmão (1)
Yuccamyces Gour, Dyko & B. Sutton (6)
Zakatoshia B. Sutton (2)
Zebrospora McKenzie (1)
Zelandiocoela Nag Raj (1)
Zelodactylaria A.C. Cruz, Gusmão & R.F. Castañeda (1)
Zelopelta B. Sutton & R.D. Gaur (1)
Zelosatchmopsis Nag Raj (1)
Zetesimomyces Nag Raj (1)
Zevadia J.C. David & D. Hawksw. (1)
Zilingia Petr. (1)
Zinzipegasa Nag Raj (1)
Zopheromyces B. Sutton & Hodges (1)
Zunura Nag Raj (1)
Zythia Fr. (1)
Zyxiphora B. Sutton (1)

BASIDIOBOLOMYCOTA Doweld
Basidiobolomycetes Doweld*
Basidiobolales Jacz. & P.A. Jacz.*
Basidiobolaceae Engl. & E. Gilg
Basidiobolus Eidam (10)
Schizangiella J. Dwyer, B. Burwell, Humber, C. Mcleod, M. Fleetwood & T. Johnson bis
(1)

BASIDIOMYCOTA R.T. Moore


Basidiomycota R.T. Moore
Agaricomycotina Doweld
Agaricomycetes Doweld
Agaricales Underw.
Agaricaceae Chevall.
Abstoma G. Cunn. (8)
Acutocapillitium P. Ponce de León (3)

1269
Agaricus L. (ca. 500)
Arachnion Schwein. (13)
Barcheria T. Lebel (1)
Battarrea Pers. (3)
Battarreoides T. Herrera (1)
Calvatiopsis Hollós (1)
Chamaemyces Battarra ex Earle (2)
Chlamydopus Speg. (1)
Chlorolepiota Sathe & S.D. Deshp. (3)
Chlorophyllum Massee (19)
Clarkeinda Kuntze (5)
Clavogaster Henn. (2)
Coniolepiota Vellinga (1)
Coprinus Pers. (ca. 17)
Crucispora E. Horak (2)
Cystolepiota Singer (ca. 12)
Dictyocephalos L.M. Underwood ex V.S. White (1)
Disciseda Czern. (15)
Echinoderma (Locq. ex Bon) Bon (ca. 15)
Endolepiotula Singer (1)
Eriocybe Vellinga (1)
Gasterellopsis Routien (1)
Glyptoderma R. Heim & Perr.-Bertr. (1)
Heinemannomyces Watling (2)
Hiatulopsis Singer & Grinling (2)
Holocotylon Lloyd (3)
Hymenagaricus Heinem. (20)
Janauaria Singer (1)
Japonogaster Kobayasi (1)
Lepiota (Pers.) Gray (ca. 450)
Leucoagaricus Locq. ex Singer (ca. 135)
Leucocoprinus Pat. (ca. 50)
Lycoperdopsis Henn. (1)
Macrolepiota Singer (ca. 40)
Melanophyllum Velen. (3)
Metrodia Raithelh. (2)
Micropsalliota Höhn. (ca. 70)
Montagnea Fr. (5)
Mycenastrum Desv. (18)
Neosecotium Singer & A.H. Sm. (2)
Panaeolopsis Singer (4)
Phellorinia Berk. (1)
Phyllogaster Pegler (1)
Podaxis Desv. (10)
Pseudoauricularia Kobayasi (1)
Pseudolepiota Z.W. Ge (1)
Queletia Fr. (2)
Rugosospora Heinem. (2)
Schinzinia Fayod (1)
Schizostoma Ehrenb. ex Lév. (1)
Singerina Sathe & S.D. Deshp. (1)
Smithiogaster J.E. Wright (1)

1270
Smithiomyces Singer (3)
Termiticola E. Horak (1)
Tulostoma Pers. (ca. 83)
Xanthagaricus (Heinem.) Little Flower, Hosag. & T.K. Abraham (12)
Xerocoprinus Maire (1)

Amanitaceae E.-J. Gilbert


Amanita Pers. (ca. 570)
Catatrama Franco-Mol. (2)
Limacella Earle (ca. 15)
Limacellopsis Zhu L. Yang, Q. Cai & Y.Y. Cui (2)
Zhuliangomyces Redhead (5)

Biannulariaceae Jülich
Anupama K.N.A. Raj, K.P.D. Latha & Manim. (1)
Callistosporium Singer (14)
Catathelasma Lovejoy (4)
Guyanagarika Sánchez-García, T.W. Henkel & Aime (3)
Macrocybe Pegler & Lodge (7)
Pleurocollybia Singer (6)
Pseudolaccaria Vizzini, Contu& Z.W. Ge (1)

Bolbitiaceae Singer
Agrogaster D.A. Reid (1)
Bolbitius Fr. (ca. 70)
Conocybe Fayod (ca. 221)
Cyttarophyllopsis R. Heim (1)
Descolea Singer (ca. 15)
Galerella Earle (8)
Galeropsis Velen. (9)
Gymnoglossum Massee (1)
Pholiotina Fayod (56)
Ptychella Roze & Boud. (1)
Rhodoarrhenia Singer (8)
Tubariella E. Horak & Hauskn. (1)
Tubariopsis R. Heim (1)
Tympanella E. Horak (1)
Wielandomyces Raithelh. (1)

Broomeiaceae Zeller
Broomeia Berk. (2)

Chromocyphellaceae Knudsen
Chromocyphella De Toni & Levi (5)

Clavariaceae Chevall.
Camarophyllopsis Herink (26)
Clavaria Vaill. ex L. (32)
Clavicorona Doty (10)
Clavulinopsis Overeem (34)
Hirticlavula J.H. Petersen & Læssøe (1)
Hodophilus R. Heim (13)

1271
Hyphodontiella Å. Strid (2)
Lamelloclavaria Birkebak & Adamčík (1)
Ramariopsis (Donk) Corner (48)
Setigeroclavula R.H. Petersen (1)

Cortinariaceae R. Heim ex Pouzar


Cortinarius (Pers.) Gray (ca. 2250)
Protoglossum Massee (8)
Pyrrhoglossum Singer (12)
Quadrispora Bougher & Castellano (3)
Stephanopus M.M. Moser & E. Horak (5)

Crassisporiaceae Vizzini, Consiglio & M. Marchetti


Crassisporium Matheny, P.-A. Moreau & Vizzini (3)
Romagnesiella Contu, Matheny, P.-A. Moreau, Vizzini & A. de Haan (2)

Crepidotaceae (S. Imai) Singer


Crepidotus (Fr.) Staude (ca. 200)
Episphaeria Donk (1)
Nanstelocephala Oberw. & R.H. Petersen (1)
Pellidiscus Donk (3)
Pleuroflammula Singer (10)
Simocybe P. Karst. (26)

Cyphellaceae Lotsy
Asterocyphella W.B. Cooke (3)
Campanophyllum Cifuentes & R.H. Petersen (1)
Catilla Pat. (1)
Cheimonophyllum Singer (4)
Chondrostereum Pouzar (4)
Cunninghammyces Stalpers (2)
Cyphella Fr. (2)
Gloeocorticium Hjortstam & Ryvarden (1)
Gloeostereum S. Ito & S. Imai (1)
Granulobasidium Jülich (1)
Hyphoradulum Pouzar (1)
Incrustocalyptella Agerer (3)
Phaeoporotheleum (W.B. Cooke) W.B. Cooke (2)
Seticyphella Agerer (3)
Sphaerobasidioscypha Agerer (2)
Thujacorticium Ginns (1)

Cystostereaceae Jülich
Cericium Hjortstam (1)
Crustomyces Jülich (3)
Cystidiodontia Hjortstam (2)
Cystostereum Pouzar (7)
Parvobasidium Jülich (3)
Parvodontia Hjortstam & Ryvarden (2)
Rigidotubus J. Song, Y.C. Dai & B.K. Cui (1)

Entolomataceae Kotl. & Pouzar

1272
Clitocella Kluting, T.J. Baroni & Bergemann (6)
Clitopilopsis Maire (2)
Clitopilus (Fr. ex Rabenh.) P. Kumm. (ca. 140)
Entocybe T.J. Baroni, V. Hofst. & Largent (10)
Entoloma P. Kumm. (ca. 1800)
Rhodocybe Maire (ca. 50)
Rhodophana Kühner (7)

Hemigasteraceae Gäum. & C.W. Dodge


Hemigaster Juel (1)

Hydnangiaceae Gäum. & C.W. Dodge


Hydnangium Wallr. (ca. 20)
Laccaria Berk. & Broome (ca. 85)
Maccagnia Mattir. (1)
Podohydnangium G.W. Beaton, Pegler & T.W.K. Young (1)

Hygrophoraceae Lotsy
Acantholichen P.M. Jørg. (6)
Aeruginospora Höhn. (2)
Ampulloclitocybe Redhead, Lutzoni, Moncalvo & Vilgalys (3)
Aphroditeola Redhead & Manfr. Binder (1)
Arrhenia Fr. (ca. 36)
Cantharocybe H.E. Bigelow & A.H. Sm. (3)
Chromosera Redhead, Ammirati & Norvell (5)
Chrysomphalina Clémençon (4)
Cora Fr. (189)
Corella Vain. (2)
Cuphophyllus (Donk) Bon (ca. 25)
Cyphellostereum D.A. Reid (9)
Dictyonema C. Agardh ex Kunth (28)
Eonema Redhead, Lücking & Lawrey (1)
Gliophorus Herink (ca. 17)
Haasiella Kotl. & Pouzar (2)
Humidicutis (Singer) Singer (12)
Hygroaster Singer (3)
Hygrocybe (Fr.) P. Kumm. (ca. 120)
Hygrophorus Fr. (ca. 200)
Lichenomphalia Redhead, Lutzoni, Moncalvo & Vilgalys (14)
Neohygrocybe Herink (5)
Porpolomopsis Bresinsky (5)
Pseudoarmillariella Singer (3)
Semiomphalina Redhead (1)
SinohygrocybeC.Q. Wang, Ming Zhang & T.H. Li (1)

Hymenogastraceae Vittad.
Anamika K.A. Thomas, Peintner, M.M. Moser & Manim. (3)
Flammula (Fr.) P. Kumm. (ca. 10)
Galerina Earle (ca. 250)
Gymnopilus P. Karst. (ca. 200)
Hebeloma (Fr.) P. Kumm. (ca. 190)
Hymenogaster Vittad. (c.170)

1273
Naucoria (Fr.) P. Kumm. (30)
Phaeocollybia R. Heim (ca. 80)
Psathyloma Soop, J.A. Cooper & Dima (2)
Psilocybe (Fr.) P. Kumm. (ca. 326)

Inocybaceae Jülich
Auritella Matheny & Bougher (8)
Inocybe (Fr.) Fr. (ca. 1000)
Tubariomyces Esteve-Rav. & Matheny (3)

Limnoperdaceae G.A. Escobar


Limnoperdon G.A. Escobar (1)

Lycoperdaceae Chevall.
Apioperdon (Kreisel & D. Krüger) Vizzini (1)
Bovista Pers. Bryoperdon Vizzini (ca. 58)
Calbovista Morse ex M.T. Seidl (1)
Calvatia Fr. (ca. 43)
Gastropila Homrich & J.E. Wright (4)
Lycoperdon Pers. (ca. 55)
Morganella Zeller (7)

Lyophyllaceae Jülich
Asterophora Ditmar (3)
Blastosporella T.J. Baroni & Franco-Mol. (1)
Calocybe Kühner ex Donk (46)
Calocybella Vizzini, Consiglio & Setti (4)
Clitolyophyllum Sesli, Vizzini & Contu (1)
Gerhardtia Bon (ca. 7)
Hypsizygus Singer (3)
Lyophyllopsis Sathe & J.T. Daniel (1)
Lyophyllum P. Karst. (ca. 60)
Myochromella V. Hofst., Clémençon, Moncalvo & Redhead (2)
Ossicaulis Redhead & Ginns (2)
Rugosomyces Raithelh. (ca. 12)
Sagaranella V. Hofst., Clémençon, Moncalvo & Redhead (4)
Sphagnurus Redhead & V. Hofst. (1)
Tephrocybe Donk (ca. 47)
Tephrocybella Picillo, Vizzini & Contu (1)
Termitomyces R. Heim (ca. 34)
Tricholomella Zerova ex Kalamees (1)

Macrocystidiaceae Kühner
Macrocystidia Joss. (5)

Marasmiaceae Roze ex Kühner


Amyloflagellula Singer (4)
Brunneocorticium Sheng H. Wu (1)
Campanella Henn. (ca. 39)
Chaetocalathus Singer (ca. 20)
Crinipellis Pat. (ca. 65)
Hymenogloea Pat. (1)

1274
Marasmius Fr. (ca. 600)
Moniliophthora H.C. Evans, Stalpers, Samson & Benny (7)
Neocampanella Nakasone, Hibbett & Goranova (1)
Tetrapyrgos E. Horak (18)

Mycenaceae Overeem
Atheniella Redhead, Moncalvo, Vilgalys, Desjardin & B.A. Perry (7)
Cruentomycena R.H. Petersen, Kovalenko & O.V. Morozova (3)
Decapitatus Redhead & Seifert (1)
Favolaschia (Pat.) Pat. (ca. 54)
Flabellimycena Redhead (1)
Heimiomyces Singer (ca. 7)
Hemimycena Singer (ca. 60)
Hydropus Kühner ex Singer (ca. 100)
Mycena (Pers.) Roussel (ca. 600)
Mycopan Redhead, Moncalvo & Vilgalys (1)
Panellus P. Karst. (ca. 55)
Resinomycena Redhead & Singer (ca. 10)
Roridomyces Rexer (9)
Sarcomyxa P. Karst. (2)
Tectella Earle (3)
Xeromphalina Kühner & Maire (ca. 32)

Mythicomycetaceae Vizzini, Consiglio & M. Marchetti


Mythicomyces Redhead & A.H. Sm. (1)
Stagnicola Redhead & A.H. Sm. (1)

Niaceae Jülich
Digitatispora Doguet (2)
Flagelloscypha Donk (ca. 25)
Halocyphina Kohlm. & E. Kohlm. (1)
Lachnella Fr. (6)
Maireina W.B. Cooke (ca. 18)
Merismodes Earle (20)
Nia R.T. Moore & Meyers (3)
Peyronelina P.J. Fisher, J. Webster & D.F. Kane (1)
Woldmaria W.B. Cooke (1)

Omphalotaceae Bresinsky
Anthracophyllum Ces. (12)
Caripia Kuntze (1)
Connopus R.H. Petersen (1)
Gymnopanella Sand.-Leiva, J.V. McDonald & Thorn (1)
Gymnopus (Pers.) Gray (ca. 325)
Hymenoporus Tkalčec, Mešić & Chun Y. Deng (1)
Lentinula Earle (8)
Marasmiellus Murrill (ca. 260)
Mycetinis Earle (15)
Neonothopanus R.H. Petersen & Krisai (3)
Omphalotus Fayod (6)
Rhodocollybia Singer (ca. 35)
Paragymnopus J.S. Oliveira (6)

1275
Pusillomyces J.S. Oliveira (3)

Physalacriaceae Corner
Anastrophella E. Horak & Desjardin (3)
Armillaria (Fr.) Staude (39)
Cibaomyces Zhu L. Yang, Y.J. Hao & J. Qin (1)
Cribbea A.H. Sm. & D.A. Reid (5)
Cryptomarasmius T.S. Jenkinson & Desjardin (15)
Cylindrobasidium Jülich (7)
Cyptotrama Singer (16)
Dactylosporina (Clémençon) Dörfelt (5)
Desarmillaria (Herink) R. A. Koch & Aime (2)
Epicnaphus Singer (2)
Flammulina P. Karst. (14)
Gloiocephala Massee (ca. 40)
Guyanagaster T.W. Henkel, M.E. Sm. & Aime (2)
Hymenopellis R.H. Petersen (ca. 50)
Laccariopsis Vizzini (1)
Manuripia Singer (1)
Mucidula Pat. (2)
Mycaureola Maire & Chemin (1)
Naiadolina Redhead, Labbé & Ginns (1)
Oudemansiella Speg. (ca. 20)
Paraxerula R.H. Petersen (4)
Physalacria Peck (33)
Ponticulomyces R.H. Petersen (2)
Protoxerula R.H. Petersen (1)
Rhizomarasmius R.H. Petersen (5)
Rhodotus Maire (2)
Strobilurus Singer (10)
Xerula Maire (ca. 17)

Pleurotaceae Kühner
Agaricochaete Eichelb. (4)
Hohenbuehelia Schulzer (ca. 50)
Lignomyces R.H. Petersen & Zmitr. (1)
Pleurotus (Fr.) P. Kumm. (25)
Resupinatus Nees ex Gray (33)

Pluteaceae Kotl. & Pouzar


Pluteus Fr. (ca. 500)
Volvariella Speg. (ca. 50)
Volvopluteus Vizzini, Contu & Justo (4)

Porotheleaceae Murrill
Phloeomana Redhead (6)
Porotheleum Fr. (ca. 16)

Psathyrellaceae Vilgalys, Moncalvo & Redhead


Coprinellus P. Karst. (70)
Coprinopsis P. Karst. (ca. 150)
Cystoagaricus Singer (7)

1276
Gasteroagaricoides D.A. Reid (1)
Homophron (Britzelm.) Örstadius & E. Larss. (3)
Hormographiella Guarro & Gené (3)
Kauffmania Örstadius & E. Larss. (1)
Lacrymaria Pat. (14)
Macrometrula Donk & Singer (1)
Parasola Redhead, Vilgalys & Hopple (ca. 27)
Psathyrella (Fr.) Quél. (ca. 420)
Rhacophyllus Berk. & Broome (1)
Typhrasa Örstadius & E. Larss. (2)

Pseudoclitocybaceae Vizzini, Consiglio, P.-A. Moreau & P. Alvarado


Bonomyces Vizzini (3)
Cleistocybe Ammirati, A.D. Parker & Matheny (5)
Clitopaxillus G. Moreno, Vizzini, Consiglio & P. Alvarado (2)
Harmajaea Dima, P. Alvarado & Kekki (3)
Musumecia Vizzini & Contu (4)
Pogonoloma (Singer) Sánchez-García (3)
Pseudoclitocybe (Singer) Singer (16)

Pterulaceae Corner
Actiniceps Berk. & Broome (6)
Allantula Corner (1)
Aphanobasidium Jülich (17)
Chaetotyphula Corner (7)
Coronicium J. Erikss. & Ryvarden (5)
Deflexula Corner (ca. 11)
Lepidomyces Jülich (2)
Merulicium J. Erikss. & Ryvarden (1)
Parapterulicium Corner (3)
Pterula Fr. (ca. 50)
Pterulicium Corner (1)
Radulomyces M.P. Christ. (10)
Radulotubus Y.C. Dai, S.H. He & C.L. Zhao (1)

Schizophyllaceae Quél.
Auriculariopsis Maire (3)
Porodisculus Murrill (2)
Schizophyllum Fr. (6)

Stephanosporaceae Oberw. & E. Horak


Athelidium Oberw. (3)
Cristinia Parmasto (10)
Lindtneria Pilát (10)
Mayamontana Castellano, Trappe & Lodge (1)
Stephanospora Pat. (6)

Strophariaceae Singer & A.H. Sm.


Agrocybe Fayod (ca. 100)
Bogbodia Redhead (1)
Brauniella Rick ex Singer (1)
Deconica (W.G. Sm.) P. Karst. (44)

1277
Hypholoma (Fr.) P. Kumm. (ca. 45)
Leratiomyces Bresinsky & Manfr. Binder ex Bridge, Spooner, Beever & D.C. Park (13)
Melanotus Pat. (ca. 33)
Pholiota (Fr.) P. Kumm. (ca. 157)
Protostropharia Redhead, Moncalvo & Vilgalys (14)
Pseudogymnopilus Raithelh. (1)
Stropharia (Fr.) Quél. (ca. 20)

Tricholomataceae R. Heim ex Pouzar


Albomagister Sánchez-García, Birkebak & Matheny (2)
Corneriella Sánchez-García (3)
Dennisiomyces Singer (5)
Dermoloma J.E. Lange ex Herink (ca. 25)
Leucopaxillus Boursier (ca. 16)
Porpoloma Singer (ca. 13)
Pseudobaeospora Singer (ca. 26)
PseudoporpolomaVizzini & Consiglio (1)
Pseudotricholoma (Singer) Sánchez-García & Matheny (2)
Tricholoma (Fr.) Staude (ca. 210)

Tubariaceae Vizzini
Cyclocybe Velen. (6)
Flammulaster Earle (10)
Hemistropharia Jacobsson & E. Larss. (1)
Pachylepyrium Singer (1)
Phaeomarasmius Scherff. (ca. 20)
Pleuromyces Dima, P.-A. Moreau & V. Papp (1)*
Tubaria (W.G. Sm.) Gillet (ca. 21)

Typhulaceae Jülich
Lutypha Khurana, K.S. Thind & Berthier (1)
Macrotyphula R.H. Petersen (6)
Tygervalleyomyces Crous (1)
Typhula (Pers.) Fr. (ca. 100)

Agaricales genera incertae sedis


Acanthocorticium Baltazar, Gorjón & Rajchenb. (1)
Acinophora Raf. (1)
Aleurocystis Lloyd ex G. Cunn. (3)
Amparoina Singer (2)
Amylolepiota Harmaja (1)
Aphyllotus Singer (1)
Arthromyces T.J. Baroni & Lodge (2)
Arthrosporella Singer (1)
Asproinocybe R. Heim (5)
Aspropaxillus Kühner & Maire (3)
Atractosporocybe P. Alvarado, G. Moreno & Vizzini (2)
Austroclitocybe Raithelh. (2)
Austroomphaliaster Garrido (1)
Baeospora Singer (13)
Callistodermatium Singer (1)
Calyptella Quél. (20)

1278
Caulorhiza Lennox (3)
Cellypha Donk (10)
Cephaloscypha Agerer (1)
Cercopemyces T.J. Baroni, Kropp & V.S. Evenson (3)
Clavomphalia E. Horak (1)
Clitocybe (Fr.) Staude (ca. 300)
Clitocybula (Singer) Singer ex Métrod (25)
Coccobotrys Boud. & Pat. (2)
Collybia (Fr.) Staude (3)
Conchomyces Overeem (2)
Crucibulum Tul. & C. Tul. (7)
Cyathus Haller (ca. 59)
Cymatella Pat. (4)
Cymatellopsis Parmasto (1)
Cynema Maas Geest. & E. Horak (1)
Cyphellocalathus Agerer (1)
Cystoderma Fayod (ca. 36)
Cystodermella Harmaja (16)
Deigloria Agerer (5)
Delicatula Fayod (ca. 3)
Dendrocollybia R.H. Petersen & Redhead (1)
Dendrothele Höhn. & Litsch. (58)
Disporotrichum Stalpers (1)
Fayodia Kühner (ca. 10)
Fibulochlamys A.I. Romero & Cabral (2)
Fissolimbus E. Horak (1)
Fistulina Bull. (9)
Floccularia Pouzar (6)
Gamundia Raithelh. (ca. 7)
Gerronema Singer (58)
Giacomia Vizzini & Contu (1)
Glabrocyphella W.B. Cooke (12)
Gloioxanthomyces Lodge, Vizzini, Ercole & Boertm. (2)
Gramincola Velen. (1)
Hemipholiota (Singer) Bon (2)*
Henningsomyces Kuntze (ca. 21)
Hispidocalyptella E. Horak & Desjardin (1)
Hygrophorocybe Vizzini & Contu (1)
Infundibulicybe Harmaja (22)
Lactocollybia Singer (20)
Lecanocybe Desjardin & E. Horak (1)
Lepista (Fr.) W.G. Sm. (ca. 50)
Lepistella T.J. Baroni & Ovrebo (ca. 50)
Leucocalocybe X.D. Yu & Y.J. Yao (1)
Leucocortinarius (J.E. Lange) Singer (1)
Leucocybe Vizzini, P. Alvarado, G. Moreno & Consiglio (3)
Leucoinocybe Singer ex Antonín, Borovička, Holec & Kolařík (3)
Leucopholiota (Romagn.) O.K. Mill., T.J. Volk & Bessette (2)
Lignomphalia Antonín, Borovička, Holec & Kolařík (1)
Lulesia Singer (3)
Lycogalopsis E. Fisch. (1)
Megacollybia Kotl. & Pouzar (9)

1279
Melanoleuca Pat. (ca. 60)
Melanomphalia M.P. Christ. (1)
Meottomyces Vizzini (1)
Mesophelliopsis Bat. & A.F. Vital (1)
Metraria (Cooke) Cooke & Massee (2)
Metulocyphella Agerer (2)
Mucronella Fr. (8)
Mycenella (J.E. Lange) Singer (10)
Mycoalvimia Singer (1)
Mycocalia J.T. Palmer (7)
Mycospongia Velen. (1)
Myxomphalia Hora (ca. 2)
Neoclitocybe Singer (11)
Neopaxillus Singer (6)
Nidula V.S. White (6)
Nidularia Fr. (3)
Nochascypha Agerer (3)
Notholepista Vizzini & Contu (1)
Omphaliaster Lamoure (7)
Omphalina Quél. (ca. 50)
Palaeocephala Singer (1)
Panaeolina Maire (2)
Panaeolus (Fr.) Quél. (15)
Paralepistopsis Vizzini (2)
Peglerochaete Sarwal & Locq. (1)
Pegleromyces Singer (1)
Phaeodepas D.A. Reid (2)
Phaeolepiota Maire ex Konrad & Maubl. (1)
Phaeomycena R. Heim ex Singer & Digilio (5)
Phaeopholiota Locq. & Sarwal (1)
Phlebonema R. Heim (1)
Phlebophyllum R. Heim (1)
Phyllotopsis E.-J. Gilbert & Donk ex Singer (5)
Physocystidium Singer (1)
Pleurella E. Horak (1)
Pleurocybella Singer (5)
Plicatura Peck (1)
Polygaster Fr. (1)
Pseudoclitopilus Vizzini & Contu (2)
Pseudofistulina O. Fidalgo & M. Fidalgo (3)
Pseudohiatula (Singer) Singer (ca. 5)
Pseudohygrophorus Velen. (1)
Pseudolasiobolus Agerer (1)
Pseudoomphalina (Singer) Singer (ca. 6)
Pseudotyphula Corner (1)
Radulomycetopsis Dhingra, Priyanka & J. Kaur (1)
Rectipilus Agerer (11)
Rhizocybe Vizzini, G. Moreno, P. Alvarado & Consiglio (4)
Rimbachia Pat. (11)
Ripartitella Singer (1)
Ripartites P. Karst. (5)
Secotium Kunze (ca. 10)

1280
Singerocybe Harmaja (7)
Skepperiella Pilát (4)
Squamanita Imbach (10)
Stanglomyces Raithelh. (1)
Stemastrum Raf. (1)
Stromatocyphella W.B. Cooke (3)
Tephroderma Contu & Musumeci (1)
Trichocybe Vizzini (1)
Tricholomopsis Singer (ca. 33)
Tricholosporum Guzmán (7)
Trogia Fr. (ca. 94)
Ugola Adans. (3)
Vanromburghia Holterm. (1)
Verrucospora E. Horak (2)

Amylocorticiales K.H. Larss., Manfr. Binder & Hibbett


Amylocorticiaceae Jülich
Amyloathelia Hjortstam & Ryvarden (3)
Amylocorticiellum Spirin & Zmitr. (4)
Amylocorticium Pouzar (11)
Amyloxenasma (Oberw.) Hjortstam & Ryvarden (6)
Anomoloma Niemelä & K.H. Larss. (6)
Anomoporia Pouzar (8)
Ceraceomyces Jülich (16)
Irpicodon Pouzar (1)
Plicaturopsis D.A. Reid (2)
Podoserpula D.A. Reid (2)
Serpulomyces (Zmitr.) Zmitr. (1)

Atheliales Jülich
Atheliaceae Jülich
Amphinema P. Karst. (4)
Athelia Pers. (32)
Athelicium K.H. Larss. & Hjortstam (2)
Athelocystis Hjortstam & Ryvarden (1)
Athelopsis Oberw. ex Parmasto (14)
Butlerelfia Weresub & Illman (1)
Byssocorticium Bondartsev & Singer (11)
Elaphocephala Pouzar (1)
Hypochnella J. Schröt. (2)
Hypochniciellum Hjortstam & Ryvarden (1)
Leptosporomyces Jülich (15)
Lobulicium K.H. Larss. & Hjortstam (1)
Lyoathelia Hjortstam & Ryvarden (1)
Melzericium Hauerslev (3)
Mycostigma Jülich (1)
Piloderma Jülich (6)
Pteridomyces Jülich (4)
Taeniospora Marvanová (2)
Tretomyces K.H. Larss., Kotir. & Saaren. (2)
Tylospora Donk (2)

1281
Auriculariales J. Schröt.
Auriculariaceae Fr.
Amphistereum Spirin & Malysheva (2)
Auricularia Bull. (ca. 21)
Eichleriella Bres. (ca. 14)
Elmerina Bres. (7)
Exidia Fr. (ca. 26)
Exidiopsis (Bref.) Möller (ca. 30)
Fibulosebacea K. Wells & Raitv. (1)
Heterochaete Pat. (ca. 40)
Heteroradulum Lloyd ex Spirin & Malysheva (7)
Protodaedalea Imazeki (2)
Pseudostypella McNabb (1)
Sclerotrema Spirin & Malysheva (1)

Hyaloriaceae Lindau
Helicomyxa R. Kirschner & Chee J. Chen (1)
Hyaloria Möller (3)
Myxarium Wallr. (14)

Auriculariales genera incertae sedis


Basidiodendron Rick (ca. 15)
Bourdotia (Bres.) Bres. & Torrend (1)
Ceratosebacina P. Roberts (3)
Dendrogloeon Spirin & Miettinen (1)
Ductifera Lloyd (ca. 11)
Endoperplexa P. Roberts (6)
Gelacantha V. Malysheva & Spirin (1)
Grammatus H.S. Yuan & C. Decock (2)
Guepinia Fr. (1)
Hauerslevia P. Roberts (1)
Heterorepetobasidium Chee J. Chen & Oberw. (2)
Heteroscypha Oberw. & Agerer (1)
Hyalodon V. Malysheva & Spirin (2)
Hydrophana V. Malysheva & Spirin (1)
Metabourdotia L.S. Olive (1)
Microsebacina P. Roberts (2)
Mycostilla Spirin & V. Malysheva (1)
Myxariellum Spirin & V. Malysheva (2)
Ofella Spirin & V. Malysheva (1)
Porpopycnis R. Kirschner (1)
Protoacia Spirin & V. Malysheva (1)
Protodontia Höhn. (3)
Protograndinia Rick (1)
Protohydnum Möller (3)
Protomerulius Möller (7)
Protoradulum Rick (1)
Pseudohydnum P. Karst. (1)
Renatobasidium Hauerslev (1)
Stypella Möller (4)
Stypellopsis Spirin & V. Malysheva (2)
Tremellacantha Jülich (1)

1282
Boletales E.-J. Gilbert
Boletaceae Chevall.
Afroboletus Pegler & T.W.K. Young (8)
Afrocastellanoa M.E. Sm. & Orihara (1)
Alessioporus Gelardi, Vizzini & Simonini (2)
Aureoboletus Pouzar (33)
Australopilus Halling & N.A. Fechner (1)
Austroboletus (Corner) Wolfe (ca. 36)
Baorangia G. Wu & Zhu L. Yang (4)
Binderoboletus T.W. Henkel & M.E. Sm. (1)
Boletellus Murrill (ca. 50)
Boletochaete Singer (5)
Boletus L. (ca. 350)
Borofutus Hosen & Zhu L. Yang (1)
Bothia Halling, T.J. Baroni & Manfr. Binder (2)
Buchwaldoboletus Pilát (11)
Butyriboletus Arora & J.L. Frank (ca. 25)
Caloboletus Vizzini (14)
Carolinigaster M.E. Sm. & S. Cruz (1)
Castellanea T.W. Henkel & M.E. Sm. (1)
Chalciporus Bataille (ca. 30)
Chamonixia Rolland (8)
Chiua Y.C. Li & Zhu L. Yang (4)
Corneroboletus N.K. Zeng & Zhu L. Yang (1)
Costatisporus T.W. Henkel & M.E. Sm. (1)
Crocinoboletus N.K. Zeng, Zhu L. Yang & G. Wu (2)
Cupreoboletus Simonini, Gelardi & Vizzini (1)
Cyanoboletus Gelardi, Vizzini & Simonini (7)
Durianella Desjardin, A.W. Wilson & Manfr. Binder (1)
Erythrophylloporus Ming Zhang & T.H. Li (1)
Fistulinella Henn. (ca. 25)
Gastroboletus Lohwag (14)
Gastroleccinum Thiers (1)
Guyanaporus T.W. Henkel & M.E. Sm. (1)
Gymnogaster J.W. Cribb (1)
Harrya Halling, Nuhn & Osmundson (6)
Heimioporus E. Horak (14)
Heliogaster Orihara & K. Iwase (1)
Hemileccinum Šutara (5)
Hortiboletus Simonini, Vizzini & Gelardi (7)
Hourangia Xue T. Zhu & Zhu L. Yang (4)
Hymenoboletus Y.C. Li & Zhu L. Yang (1)
Imleria Vizzini (5)
Imperator G. Koller, Assyov, Bellanger, Bertéa, Loizides, G. Marques, P.-A. Moreau, J.A.
Muñoz, Oppicelli, Puddu & F. Richard (3)
Indoporus A. Parihar, K. Das, Hembrom & Vizzini (1)
Ionosporus O. Khmelnitsky (2)
Jimtrappea T.W. Henkel, M.E. Sm. & Aime (2)
Kombocles Castellano, T.W. Henkel & Dentinger (1)
Lanmaoa G. Wu & Zhu L. Yang (7)
Leccinellum Bresinsky & Manfr. Binder (17)
Leccinum Gray (ca. 130)

1283
Mackintoshia Pacioni & Sharp (1)
Mucilopilus Wolfe (1)
Mycoamaranthus Castellano, Trappe & Malajczuk (3)
Neoboletus Gelardi, Simonini & Vizzini (11)
Nigroboletus Gelardi, Vizzini, E. Horak, T.H. Li & Ming Zhang (1)
Octaviania Vittad. (ca. 40)
Parvixerocomus G. Wu & Zhu L. Yang (2)
Paxillogaster E. Horak (1)
Phylloboletellus Singer (1)
Phyllobolites Singer (1)
Phylloporus Quél. (ca. 90)
Porphyrellus E.-J. Gilbert (ca. 20)
Pseudoaustroboletus Y.C. Li & Zhu L. Yang (3)
Pseudoboletus Šutara (2)
Pulchroboletus Gelardi, Vizzini & Simonini (1)
Pulveroboletus Murrill (38)
Retiboletus Manfr. Binder & Bresinsky (12)
Rheubarbariboletus Vizzini, Simonini & Gelardi (2)
Rhodactina Pegler & T.W.K. Young (3)
Rossbeevera T. Lebel, Orihara & N. Maek. (10)
Royoungia Castellano, Trappe & Malajczuk (6)
Rubroboletus Kuan Zhao & Zhu L. Yang (14)
Rugiboletus G. Wu & Zhu L. Yang (2)
Setogyroporus Heinem. & Rammeloo (1)
Singerocomus T.W. Henkel & M.E. Sm. (2)
Singeromyces M.M. Moser (1)
Solioccasus Trappe, Osmundson, Manfr. Binder, Castellano & Halling (1)
Spongiforma Desjardin, Manfr. Binder, Roekring & Flegel (2)
Spongispora G. Wu, S.M.L. Lee, E. Horak & Zhu L. Yang (1)
Strobilomyces Berk. (ca. 27)
Suillellus Murrill (23)
Sutorius Halling, Nuhn & N.A. Fechner (2)
Tengioboletus G. Wu & Zhu L. Yang (2)
Tubosaeta E. Horak (6)
Turmalinea Orihara & N. Maek. (4)
Tylocinum Y.C. Li & Zhu L. Yang (1)
Tylopilus P. Karst. (ca. 100)
Veloporphyrellus L.D. Gómez & Singer (7)
Wakefieldia Corner & Hawker (2)
Xanthoconium Singer (10)
Xerocomellus Šutara (17)
Xerocomus Quél. (ca. 120)
Zangia Y.C. Li & Zhu L. Yang (6)

Boletinellaceae P.M. Kirk, P.F. Cannon & J.C. David


Boletinellus Murrill (1)
Phlebopus (R. Heim) Singer (14)

Calostomataceae E. Fisch.
Calostoma Desv. (16)

Coniophoraceae Ulbr.

1284
Chrysoconia McCabe & G.A. Escobar (1)
Coniophora DC. (20)
Coniophoropsis Hjortstam & Ryvarden (2)
Gyrodontium Pat. (3)
Sedecula Zeller (1)

Diplocystidiaceae Kreisel
Astraeus Morgan (11)
Diplocystis Berk. & M.A. Curtis (2)
Endogonopsis R. Heim (1)
Tremellogaster E. Fisch. (1)

Gasterellaceae Zeller
Gasterella Zeller & L.B. Walker (1)

Gomphidiaceae Maire ex Jülich


Chroogomphus (Singer) O.K. Mill. (25)*
Cystogomphus Singer (1)
Gomphidius Fr. (10)
Gomphogaster O.K. Mill. (1)

Gyroporaceae (Singer) Manfr. Binder & Bresinsky


Gyroporus Quél. (24)

Hygrophoropsidaceae Kühner
Hygrophoropsis (J. Schröt.) Maire ex Martin-Sans (16)
Leucogyrophana Pouzar (13)

Paxillaceae Lotsy
Alpova C.W. Dodge (16)
Austrogaster Singer (4)
Gyrodon Opat. (10)
Hoehnelogaster Lohwag (1)
Hydnomerulius Jarosch & Besl (1)
Meiorganum R. Heim (3)
Melanogaster Corda (ca. 26)
Neoalpova Vizzini (1)
Paragyrodon (Singer) Singer (1)
Paxillus Fr. (19)

Protogastraceae Zeller
Protogaster Thaxt. (1)

Rhizopogonaceae Gäum. & C.W. Dodge


Fevansia Trappe & Castellano (1)
Rhizopogon Fr. (ca. 157)
Rhopalogaster J.R. Johnst. (1)

Sclerodermataceae Corda
Chlorogaster Læssøe & Jalink (1)
Favillea Fr. (2)
Horakiella Castellano & Trappe (2)

1285
Pisolithus Alb. & Schwein. (17)
Scleroderma Pers. (ca. 46)

Serpulaceae Jarosch & Bresinsky


Austropaxillus Bresinsky & Jarosch (9)
Gymnopaxillus E. Horak (4)
Serpula (Pers.) Gray (ca. 11)

Suillaceae Besl & Bresinsky


Psiloboletinus Singer (1)
Suillus Gray (ca. 60)

Tapinellaceae C. Hahn
Bondarcevomyces Parmasto (1)
Pseudomerulius Jülich (4)
Tapinella E.-J. Gilbert (2)

Boletales genera incertae sedis


Corditubera Henn. (5)
Corneromyces Ginns (2)
Marthanella States & Fogel (1)
Phaeoradulum Pat. (1)

Cantharellales Gäum.
Aphelariaceae Corner
Aphelaria Corner (20)
Phaeoaphelaria Corner (1)
Tumidapexus D.A. Crawford (1)

Botryobasidiaceae Jülich
Acladium Link (20)
Allescheriella Henn. (5)
Alysidium Kunze (4)
Botryobasidium Donk (ca. 58)
Suillosporium Pouzar (4)

Ceratobasidiaceae G.W. Martin


Ceratobasidium D.P. Rogers (ca. 19)
Ceratoporia Ryvarden & de Meijer (1)
Ceratorhiza R.T. Moore (7)
Rhizoctonia DC. (ca. 50)
Scotomyces Jülich (1)
Thanatephorus Donk (12)

Hydnaceae Chevall.
Burgoa Goid. (9)
Burgella Diederich & Lawrey (2)
Burgellopsis Diederich & Lawrey (1)
Cantharellus Adans.ex Fr. (ca. 300)
Clavulina J. Schröt. (ca. 75)
Corallofungus Kobayasi (2)
Craterellus Pers. (ca. 80)

1286
Gloeomucro R.H. Petersen (10)
Hydnum L. (49)
Ingoldiella D.E. Shaw (3)
Membranomyces Jülich (1)
Multiclavula R.H. Petersen (13)
Neoburgoa Diederich, E. Zimm. & Lawrey (1)
Parastereopsis Corner (1)
Osteomorpha G. Arnaud ex Watling & W.B. Kendr. (1)
Paullicorticium J. Erikss. (5)
Repetobasidiellum J. Erikss. & Hjortstam (1)
Repetobasidium J. Erikss. (12)
Rogersiomyces J.L. Crane & Schokn. (2)
Sistotrema Fr. (ca. 55)
Sistotremella Hjortstam (3)

Oliveoniaceae P. Roberts
Oliveonia Donk (5)

Tulasnellaceae Juel
Pseudotulasnella Lowy (1)
Tulasnella J. Schröt. (ca. 70)

Cantharellales genera incertae sedis


Boidinella Nakasone (2)
Bulbilla Diederich, Flakus & Etayo (1)
Clavulicium Boidin (3)
Minimedusa Weresub & P.M. LeClair (3)
Odontiochaete Rick (1)
Radulochaete Rick (2)
Schildia Franchi & M. Marchetti (1)
Stilbotulasnella Oberw. & Bandoni (1)

Corticiales K.H. Larss.


Corticiaceae Herter
Capillosclerotium Prameela & Deeba (1)
Corticirama Pilát (2)
Corticium Pers. (25)
Erythricium J. Erikss. & Hjortstam (6)
Galzinia Bourdot (9)
Giulia Tassi (1)
Laetisaria (Burds. (7)
Lawreymyces Lücking & Moncada (7)
Marchandiomyces Diederich & D. Hawksw. (3)
Necator Massee (1)
Tretopileus B.O. Dodge (3)
Waitea Warcup & P.H.B. Talbot (1)

Dendrominiaceae Ghobad-Nejhad
Dendrominia Ghobad-Nejhad & Duhem (4)

Punctulariaceae Donk
Dendrocorticium M.J. Larsen & Gilb. (9)

1287
Punctularia Pat. (2)
Punctulariopsis Ghobad-Nejhad (4)

Vuilleminiaceae Maire ex Lotsy


Australovuilleminia Ghobad-Nejhad & Hallenb. (1)
Cytidia Quél. (5)
Vuilleminia Maire (8)

Corticiales genera incertae sedis


Ambivina Katz (1)
Amylobasidium Ginns (1)
Leptocorticium Hjortstam & Ryvarden (8)
Melzerodontia Hjortstam & Ryvarden (3)
Nothocorticium Gresl. & Rajchenb. (1)
Papyrodiscus D. A. Reid (1)
Ripexicium Hjortstam (1)

Geastrales K. Hosaka & Castellano


Geastraceae Corda
Geasteroides Long (1)
Geastrum Pers. (130)
Myriostoma Desv. (4)
Nidulariopsis Greis (2)
Phialastrum Sunhede (1)
Schenella T. Macbr. (4)
Sphaerobolus Tode (3)

Sclerogastraceae Locq. ex P. M. Kirk


Sclerogaster R. Hesse (11)

Geastrales genus incertae sedis


Boninogaster Kobayasi (1)

Gloeophyllales Thorn
Gloeophyllaceae Jülich
Boreostereum Parmasto (4)
Campylomyces Nakasone (2)
Chaetodermella Rauschert (1)
Gloeophyllum P. Karst. (13)
Griseoporia Ginns (2)
Heliocybe Redhead & Ginns (1)
Hispidaedalea Y. C. Dai & S.H. He (1)
Mycothele Jülich (1)
Neolentinus Redhead & Ginns (14)
Osmoporus Singer (2)
Stiptophyllum Ryvarden (1)
Veluticeps Cooke (12)

Gloeophyllales genus incertae sedis


Pileodon P. Roberts & Hjortstam (2)

Gomphales Jülich

1288
Clavariadelphaceae Corner
Beenakia D. A. Reid (7)
Clavariadelphus Donk (20)

Gomphaceae Donk
Araeocoryne Corner (1)
Ceratellopsis Konrad & Maubl. (9)
Delentaria Corner (1)
Destuntzia Fogel & Trappe (5)
Gautieria Vittad. (37)
Gloeocantharellus Singer (12)
Gomphus Pers. (7)
Phaeoclavulina Brinkmann (41)
Protogautieria A. H. Sm. (2)
Pseudogomphus R. Heim (1)
Ramaria Fr. ex Bonord. (230)
Ramaricium J. Erikss. (5)
Terenodon Maas Geest. (1)
Turbinellus Earle (5)

Lentariaceae Jülich
Hydnocristella R.H. Petersen (2)
Kavinia Pilát (4)
Lentaria Corner (19)

Hymenochaetales Oberw.
Hymenochaetaceae Donk
Arambarria Rajchenb. & Pildain (1)
Asterodon Pat. (1)
Aurificaria D.A. Reid (2)
Botryodontia (Hjortstam & Ryvarden) Hjortstam (6)
Clavariachaete Corner (2)
Coltricia Gray (40)
Coltriciella Murrill (13)
Coniferiporia L.W. Zhou & Y. C. Dai (3)
Cylindrosporus L.W. Zhou (1)
Deviodontia (Parmasto) Hjortstam & Ryvarden (1)
Dichochaete Parmasto (2)
Erythromyces Hjortstam & Ryvarden (1)
Fomitiporella Murrill (13)
Fomitiporia Murrill (46)
Fulvifomes Murrill (33)
Fuscoporia Murrill (62)
Hastodontia (Parmasto) Hjortstam & Ryvarden (2)
Hydnochaete Bres. (1)
Hymenochaete Lév. (149)
Hymenochaetopsis S. H. He & Jiao Yang (16)
Inocutis Fiasson & Niemelä (9)
Inonotopsis Parmasto (1)
Inonotus P. Karst. (120)
Mensularia Lázaro Ibiza (6)
Neomensularia F. Wu, L. W. Zhou & Y.C. Dai (4)

1289
Nothophellinus Rajchenb. (1)
Onnia P. Karst. (8)
Phellinidium (Kotl.) Fiasson & Niemelä (5)
Phellinopsis Y. C. Dai (10)
Phellinotus Drechsler-Santos, Robledo & Rajchenb. (2)
Phellinus Quél. (202)
Phellopilus Niemelä, T. Wagner & M. Fisch. (1)
Phylloporia Murrill (38)
Porodaedalea Murrill (14)
Pseudoinonotus T. Wagner & M. Fisch. (8)
Pyrrhoderma Imazeki (2)
Sanghuangporus Sheng H. Wu, L.W. Zhou & Y. C. Dai (13)
Tropicoporus L.W. Zhou, Y. C. Dai & Sheng H. Wu (12)
Tubulicrinis Donk (34)
Xanthoporia Murrill (3)

Neoantrodiellaceae Y.C. Dai, B. K. Cui, Jia J. Chen & H. S. Yuan


Neoantrodiella Y. C. Dai, B. K. Cui, Jia J. Chen & H.S. Yuan (2)

Nigrofomitaceae Jülich
Nigrofomes Murrill (3)

Oxyporaceae Zmitr. & V. Malysheva


Oxyporus (Bourdot & Galzin) Donk (18)

Rickenellaceae Vizzini
Alloclavaria Dentinger & D. J. McLaughlin (1)
Atheloderma Parmasto (2)
Contumyces Redhead, Moncalvo, Vilgalys & Lutzoni (3)
Cotylidia P. Karst. (10)
Globulicium Hjortstam (1)
Peniophorella P. Karst. (25)
Resinicium Parmasto (8)
Rickenella Raithelh. (10)

Schizoporaceae Jülich
Alutaceodontia (Parmasto) Hjortstam & Ryvarden (1)
Basidioradulum Nobles (1)
Echinoporia Ryvarden (3)
Fibrodontia Parmasto (6)
Hyphodontia J. Erikss. (86)
Lagarobasidium Jülich (5)
Leucophellinus Bondartsev & Singer (1)
Paratrichaptum Corner (1)
Poriodontia Parmasto (1)
Rogersella Liberta & A.J. Navas (1)
Odontiopsis Hjortstam & Ryvarden (2)
Schizopora Velen. (7)
Xylodon (Pers.) Gray (60)

Hymenochaetales genera incertae sedis


Caeruleomyces Stalpers (1)

1290
Cantharellopsis Kuyper (1)
Cyanotrama Ghobad-Nejhad & Y.C. Dai (1)
Fibricium J. Erikss. (5)
Ginnsia Sheng H. Wu & Hallenb. (1)
Gyroflexus Raithelh. (1)
Kurtia Karasiński (3)
Lawrynomyces Karasiński (1)
Muscinupta Redhead, Lücking & Lawrey (1)
Physodontia Ryvarden & H. Solheim (1)
Sidera Miettinen & K. H. Larss. (6)
Skvortzovia Bononi & Hjortstam (1)
Subulicium Hjortstam & Ryvarden (3)
Trichaptum Murrill (27)
Tsugacorticium Nakasone & Burds. (1)

Hysterangiales K. Hosaka & Castellano


Gallaceaceae Locq. ex P. M. Kirk
Austrogautieria E. L. Stewart & Trappe (7)
Gallacea Lloyd (6)
Hallingea Castellano (3)

Hysterangiaceae E. Fisch.
Aroramyces Castellano & Verbeken (5)
Circulocolumella S. Ito & S. Imai (1)
Clathrogaster Petri (2)
Hysterangium Vittad. (54)

Mesophelliaceae Jülich
Andebbia Trappe, Castellano & Amar. (1)
Castoreum Cooke & Massee (3)
Chondrogaster Maire (2)
Gummiglobus Trappe, Castellano & Amar. (2)
Gummivena Trappe & Bougher (1)
Malajczukia Trappe & Castellano (8)
Mesophellia Berk. (15)
Nothocastoreum G. W. Beaton (1)

Phallogastraceae Locq.
Phallogaster Morgan (1)
Protubera Möller (13)

Trappeaceae P.M. Kirk


Phallobata G. Cunn. (1)
Restingomyces Sulzbacher, Grebenc & Baseia (1)
Trappea Castellano (1)

Jaapiales Manfr. Binder, K. H. Larss. & Hibbett


Jaapiaceae Manfr. Binder, K. H. Larss. & Hibbett
Jaapia Bres. (2)

Lepidostromatales B.P. Hodk. & Lücking


Lepidostromataceae Ertz, Eb. Fisch., Killmann, Sérus. & Lawrey

1291
Ertzia B.P. Hodk. & Lücking (1)
Lepidostroma Mägd. & S. Winkl. (1)
Sulzbacheromyces B. P. Hodk. & Lücking (6)

Phallales E. Fisch.
Claustulaceae G. Cunn.
Claustula K.M. Curtis (1)
Gelopellis Zeller (6)
Kjeldsenia W. Colgan, Castellano & Bougher (1)
Phlebogaster Fogel (2)
Pseudogelopellis K. Tao & B. Liu (1)

Gastrosporiaceae Pilát
Gastrosporium Mattir. (2)

Phallaceae Corda
Abrachium Baseia & T. S. Cabral (1)
Aporophallus Möller (1)
Aseroë Labill. (2)
Blumenavia Möller (3)
Calvarula Zeller (1)
Clathrus P. Micheli ex L. (20)
Colus Cavalier & Séchier (4)
Echinophallus Henn. (1)
Endoclathrus B. Liu, Yin H. Liu & Z.J. Gu (1)
Endophallus M. Zang & R. H. Petersen (1)
Ileodictyon Tul. & C. Tul. (2)
Itajahya Möller (4)
Kobayasia S. Imai & A. Kawam. (1)
Laternea Turpin (2)
Ligiella J.A. Sáenz (1)
Lysurus Fr. (30)
Mutinus Fr. (21)
Neolysurus O. K. Mill., Ovrebo & Burk (1)
PhallusJunius ex L. (34)
Protuberella S. Imai & A. Kawam. (1)
Pseudoclathrus B. Liu & Y.S. Bau (5)
Pseudocolus Lloyd (2)
Staheliomyces E. Fisch. (1)
Staurophallus Mont. (1)
Stephanophallus MacOwan (1)
Xylophallus (Schltdl.) E. Fisch. (2)

Phallales genera incertae sedis


Saprogaster Fogel & States (1)
Vandasia Velen. (1)

Polyporales Gäum.
Cerrenaceae Miettinen, Justo & Hibbett
Cerrena Gray (7)
Irpiciporus Murrill (1)
Pseudolagarobasidium J. C. Jang & T. Chen (7)

1292
Radulodon Ryvarden (11)

Dacryobolaceae Jülich
Amylocystis Bondartsev & Singer ex Singer (1)
Dacryobolus Fr. (7)
Jahnoporus Nuss (4)
Oligoporus Bref. (15)
Osteina Donk (1)
Postia Fr. (40)
Spongiporus Murrill (7)

Fomitopsidaceae Jülich*
Adustoporia Audet (1)
Anthoporia Karasiński & Niemelä (1)
Antrodia P. Karst. (80)
Antrodiopsis Audet (1)
Brunneoporus Audet (5)
Buglossoporus Kotl. & Pouzar (9)
Daedalea Pers. (12)
Dentiporus Audet (1)
Flavidoporia Audet (3)
Fomitopsis P. Karst. (40)
Fragifomes B. K. Cui, M.L. Han & Y. C. Dai (1)
Laricifomes Kotl. & Pouzar (1)
Lentoporia Audet (1)
Neoantrodia Audet (13)
Neolentiporus Rajchenb. (2)
Niveoporofomes B. K. Cui, M. L. Han & Y. C. Dai (1)
Ranadivia Zmitr. (5)*
Resinoporia Audet (11)
Rhizoporia Audet (1)
Rhodofomes Kotl. & Pouzar (5)
Rhodofomitopsis B.K. Cui, M. L. Han & Y. C. Dai (4)
Rubellofomes B. K. Cui, M.L. Han & Y. C. Dai (2)
Subantrodia Audet (2)
Ungulidaedalea B. K. Cui, M.L. Han & Y. C. Dai (1)
Wolfiporia Ryvarden & Gilb. (6)

Fragiliporiaceae Y. C. Dai, B.K. Cui & C. L. Zhao


Fragiliporia Y. C. Dai, B.K. Cui & C. L. Zhao (1)

Gelatoporiaceae Miettinen, Justo & Hibbett


Cinereomyces Jülich (2)
Gelatoporia Niemelä (2)
Obba Miettinen & Rajchenb. (2)
Sebipora Miettinen (1)

Grifolaceae Jülich
Aegis Gómez-Montoya, Rajchenb. & Robledo (1)
Grifola Gray (5)

Hyphodermataceae Jülich

1293
Hyphoderma Fr. (20)

Incrustoporiaceae Jülich
Gloeoporellus Zmitr. (1)*
Incrustoporia Domanski (5)
Piloporia Niemelä (2)
Skeletocutis Kotl. & Pouzar (40)
Tyromyces P. Karst. (41)

Irpicaceae Spirin & Zmitr.


Byssomerulius Parmasto (8)
Ceriporia Donk (ca. 50)
Cytidiella Pouzar (2)
Efibula Sheng H. Wu (18)
Emmia Zmitr., Spirin & Malysheva (2)
FlavodonRyvarden (3)
Gloeoporus Mont. (13)
Hydnopolyporus D. A. Reid (2)
Irpex Fr. (10)
Leptoporus Quél. (1)
Meruliopsis Bondartsev (4)
Raduliporus Spirin & Zmitr. (1)
Resiniporus Zmitr. (2)*
Trametopsis Tomšovský (4)

Ischnodermataceae Jülich
Ischnoderma P. Karst. (10)

Laetiporaceae Jülich
Kusaghiporia J. Hussein, S. Tibell & Tubuhwa (1)*
Laetiporus Murrill (15)
Phaeolus (Pat.) Pat. (3)

Meripilaceae Jülich
Meripilus P. Karst. (5)
Pseudonadsoniella T. O. Kondr. & S. Y. Kondr. (1)
Rigidoporus Murrill (30)

Meruliaceae Rea
Aurantiopileus Ginns, D. L. Lindner & T. J. Baroni (3)
Aurantiporus Murrill (6)
Ceriporiopsis Domański (40)
Climacodon P. Karst. (7)
Crustodontia Hjortstam & Ryvarden (1)
Geesterania Westphalen, Tomšovský & Rajchenb. (2)*
Hermanssonia Zmitr. (1)*
Hydnophanerochaete Sheng H. Wu & C.C. Chen (1)*
Hydnophlebia Parmasto (5)
Lilaceophlebia (Parmasto) Spirin & Zmitr. (2)
Luteoporia F. Wu, Jia J. Chen & S. H. He (1)
Merulius Fr. (150)
Mycoacia Donk (16)

1294
Mycoaciella J. Erikss. & Ryvarden (5)
Odoria V. Papp & Dima (1)
Pappia Zmitr. (1)*
Phlebia Fr. (60)
Phlebiporia Jia J. Chen, B. K. Cui & Y. C. Dai (1)
Physisporinus P. Karst. (15)
Sarcodontia Schulzer (1)
Scopuloides (Massee) Höhn. & Litsch. (5)
Stereophlebia Zmitr. (1)*

Panaceae Miettinen, Justo & Hibbett


Cymatoderma Jungh. (11)
Panus Fr. (20)

Phanerochaetaceae Jülich
Bjerkandera P. Karst. (5)
Crepatura C.L. Zhao (1)*
Donkia Pilát (1)
Efibulella Zmitr. (1)
Geliporus Yuan Yuan, Jia J. Chen & S. H. He (1)
Hapalopilus P. Karst. (11)
Hyphodermella J. Erikss. & Ryvarden (7)
Odontoefibula C. C. Chen & Sheng H. Wu (1)*
Oxychaete Miettinen (1)
Phaeophlebiopsis D. Floudas & Hibbett (3)
Phanerina Miettinen (1)
Phanerochaete P. Karst. (80)
Phlebiopsis Jülich (22)
Pirex Hjortstam & Ryvarden (1)
Porostereum Pilát (15)
Rhizochaete Gresl., Nakasone & Rajchenb. (13)
Riopa D.A. Reid (3)
Terana Adans. (1)

Podoscyphaceae D.A. Reid


Abortiporus Murrill (4)
Podoscypha Pat. (36)
Pouzaroporia Vampola (1)

Polyporaceae Fr. ex Corda


Abundisporus Ryvarden (8)
Amauroderma Murrill (40)
Atroporus Ryvarden (3)
Australoporus P.K. Buchanan & Ryvarden (1)
Bresadolia Speg. (4)*
Cerarioporia F. Wu, L.W. Zhou & J. Si (1)
Cerioporus Quél. (15)
Cinereomycetella Zmitr. (1)*
Colospora Miettinen & Spirin (2)
Coriolopsis Murrill (19)
Cryptoporus (Peck) Shear (2)
Daedaleopsis J. Schröt. (7)

1295
Datronia Donk (9)
Datroniella B.K. Cui, Hai J. Li & Y.C. Dai (6)
Dentocorticium (Parmasto) M.J. Larsen & Gilb. (3)*
Dextrinoporus H.S. Yuan (1)*
Dichomitus D.A. Reid (13)
Donkioporia Kotl. & Pouzar (2)
Donkioporiella L.W. Zhou (1)
Earliella Murrill (1)
Echinochaete D.A. Reid (5)
Endopandanicola Tibpromma & K.D. Hyde (1)
Epithele (Pat.) Pat. (17)
Epithelopsis Jülich (2)
Favolus Fr. (20)
Flammeopellis Y.C. Dai, B.K. Cui & C.L. Zhao (1)
Fomes (Fr.) Fr. (3)
Fomitella Murrill (2)
Globifomes Murrill (1)
Foraminispora Robledo, Costa-Rezende & Drechsler-Santos (1)
Funalia Pat. (10)
Furtadoa Costa-Rezende, Robledo & Drechsler-Santos (3)
Ganoderma P. Karst. (180)
Grammothele Berk. & M.A. Curtis (20)
Grammothelopsis Jülich (7)
Haddowia Steyaert (3)
Haploporus Bondartsev & Singer (13)
Hexagonia Fr. (17)
Hornodermoporus Teixeira (2)
Humphreya Steyaert (4)
Laccocephalum Mc Alpine & Tepper (5)
Leifiporia Y.C. Dai, F. Wu & C.L. Zhao (2)
Lentinus Fr. (55)
Lignosus Lloyd ex Torrend (8)
Lopharia Kalchbr. & MacOwan (7)
Megasporia B.K. Cui, Y.C. Dai & Hai J. Li (10)
Megasporoporia Ryvarden & J.E. Wright (3)
Megasporoporiella B.K. Cui, Y.C. Dai & Hai J. Li (5)
Melanoderma B.K. Cui & Y.C. Dai (2)
Microporellus Murrill (20)
Microporus P. Beauv. (13)
Mollicarpus Ginns (1)
Murinicarpus B.K. Cui & Y.C. Dai (2)
Myriothele Nakasone (1)
Navisporus Ryvarden (8)
Neodatronia B.K. Cui, Hai J. Li & Y.C. Dai (2)
Neodictyopus Palacio, Robledo, Reck & Drechsler-Santos (3)
Neofavolus Sotome & T. Hatt. (4)
Neofomitella Y.C. Dai, Hai J. Li & Vlasák (3)
Pachykytospora Kotl. & Pouzar (4)
Perenniporia Murrill (100)
Perenniporiella Decock & Ryvarden (5)
Perenniporiopsis C.L. Zhao (1)
Phaeotrametes Lloyd ex J. E. Wright (1)

1296
Picipes Zmitr. & Kovalenko (16)
Pilatotrama Zmitr. (1)*
Podofomes Pouzar (3)
Polyporopsis Audet (1)
Polyporus [P. Micheli ex Adans.] Fr. (35)
Porogramme (Pat.) Pat. (4)
Pseudofavolus Pat. (4)
Pseudomegasporoporia X.H. Ji & F. Wu (1)
Pseudopiptoporus Ryvarden (2)
Pyrofomes Kotl. & Pouzar (8)
Rubroporus Log.-Leite, Ryvarden & Groposo (1)
Sparsitubus L.W. Hsu & J.D. Zhao (1)
Szczepkamyces Zmitr. (1)*
Theleporus Fr. (9)
Thermophymatospora Udagawa, Awao & Abdullah (1)
Tinctoporellus Ryvarden (4)
Tomophagus Murrill (2)
Trametes Fr. (70)
Truncospora Pilát (23)
Vanderbylia D.A. Reid (7)
Yuchengia B.K. Cui & K.T. Steffen (1)

Sparassidaceae Jülich
Crustoderma Parmasto (16)
Pycnoporellus Murrill (2)
Sparassis Fr. (7)

Steccherinaceae Parmasto
Antella Miettinen (3)
Antrodiella Ryvarden & I. Johans. (50)
Atraporiella Ryvarden (2)
Austeria Miettinen (1)
Butyrea Miettinen (2)
Cabalodontia Piątek (5)
Caudicicola Miettinen, M. Kulju & Kotir. (1)
Citripora Miettinen (2)
Elaphroporia Z.Q. Wu & C.L. Zhao (1)
Flabellophora G. Cunn. (18)
Flaviporus Murrill (14)
Frantisekia Spirin & Zmitr. (4)
Junghuhnia Corda (35)
Lamelloporus Ryvarden (1)
Loweomyces (Kotl. & Pouzar) Jülich (6)
Metuloidea G. Cunn. (5)
Mycorrhaphium Maas Geest. (6)
Niemelaea Zmitr., Ezhov & Khimich (5)
Nigroporus Murrill (3)
Steccherinum Gray (40)
Trullella Zmitr. (6)*
Xanthoporus Audet (2)

1297
Polyporales genera incertae sedis
Aegeritopsis Höhn. (1)
Amaropostia B.K. Cui, L.L. Shen & Y.C. Dai (2)
Amaurohydnum Jülich (1)
Amauromyces Jülich (1)
Amethicium Hjortstam (1)
Amyloporia Singer (5)
Aquascypha D.A. Reid (1)
Auriporia Ryvarden (4)
Australicium Hjortstam & Ryvarden (2)
Australohydnum Jülich (2)
Austrolentinus Ryvarden (1)
Bourdotiella Duhem & Schultheis (1)
Bulbillomyces Jülich (1)
Calcipostia B.K. Cui, L.L. Shen & Y.C. Dai (1)
Candelabrochaete Boidin (12)
Climacocystis Kotl. & Pouzar (2)
Columnodontia Jülich (1)
Conohypha Jülich (2)
Coralloderma D.A. Reid (2)
Cordochaete Sanyal, Samita, Dhingra & Avn. P. Singh (1)
Cryptomphalina R. Heim (1)
Cyanodontia Hjortstam (1)
Cyanosporus McGinty (1)
Cystidiopostia B.K. Cui, L.L. Shen & Y.C. Dai (3)
Dendrophlebia Dhingra & Priyanka (1)
Diacanthodes Singer (3)
Diplomitoporus Domański (25)
Erastia Niemelä & Kinnunen (1)
Faerberia Pouzar (1)
Fibroporia Parmasto (10)
Fuscopostia B.K. Cui, L.L. Shen & Y.C. Dai (4)
Gilbertsonia Parmasto (1)
Globosomyces Jülich (1)
Globuliciopsis Hjortstam & Ryvarden (2)
Gyrophanopsis Jülich (2)
Henningsia Möller (5)
Hymenogramme Mont. & Berk. (1)
Hyphodontiastra Hjortstam (1)
Hypochnicium J. Erikss. (30)
Inflatostereum D.A. Reid (2)
Irpicochaete Rick (1)
Laetifomes T. Hatt. (1)
Macrohyporia I. Johans. & Ryvarden (2)
Meruliophana Duhem & Buyck (1)
Mycoleptodonoides Nikol. (4)
Mycorrhaphoides Hembrom, K. Das & Hallenb. (1)
Nigrohydnum Ryvarden (1)
Phaneroites Hjortstam & Ryvarden (1)
Phanerodontia Hjortstam & Ryvarden (4)
Phlebiella P.Karst. (20)
Piptoporellus B.K. Cui, M.L. Han & Y.C. Dai (3)

1298
Pseudofibroporia Yuan Y. Chen, B.K. Cui & Y.C. Dai (1)
Repetobasidiopsis Dhingra & Avn. P. Singh (1)
Rhodonia Niemelä (1)
Rickiopora Westphalen, Tomšovský & Rajchenb. (1)
Roseofavolus T. Hatt. (1)
Roseograndinia Hjortstam & Ryvarden (1)
Ryvardenia Rajchenb. (2)
Sarcoporia P. Karst. (9)
Skeletohydnum Jülich (1)
Sparassiella Schwarzman (1)
Spathulina Pat. (1)
Spongioides Lázaro Ibiza (1)
Spongipellis Pat. (8)
Stegiacantha Maas Geest. (1)
Taiwanofungus Sheng H. Wu, Z.H. Yu, Y.C. Dai & C.H. Su (2)
Uncobasidium Hjortstam & Ryvarden (2)

Russulales Kreisel ex P. M. Kirk, P. F. Cannon & J. C. David


Albatrellaceae Nuss
Albatrellopsis Teixeira (8)
Albatrellus Gray (22)
Byssoporia M.J. Larsen & Zak (1)
Leucogaster R. Hesse (20)
Leucophleps Harkn. (3)
Mycolevis A.H. Sm. (1)
Polyporoletus Snell (4)
Scutiger Paulet (10)

Auriscalpiaceae Maas Geest.


Amylonotus Ryvarden (6)
Artomyces Jülich (17)
Auriscalpium Gray (8)
Dentipratulum Domański (3)
Lentinellus P. Karst. (30)
Stalpersia Parmasto (1)

Bondarzewiaceae Kotl. & Pouzar


Amylaria Corner (1)
Amylosporus Ryvarden (12)
Bondarzewia Singer (14)
Gloiodon P. Karst. (3)
Heterobasidion Bref. (15)
Laurilia Pouzar (2)
Lauriliella Nakasone & S.H. He (2)
Stecchericium D.A. Reid (7)
Wrightoporia Pouzar (32)

Echinodontiaceae Donk
Echinodontiellum S.H. He & Nakasone (1)
Echinodontium Ellis & Everh. (4)
Larssoniporia Y.C. Dai, Jia J. Chen & B.K. Cui (2)

1299
Hericiaceae Donk
Dentipellicula Y.C. Dai & L.W. Zhou (3)
Dentipellis Donk (7)
Hericium Pers. (23)
Laxitextum Lentz (3)
Pseudowrightoporia Y.C. Dai, Jia J. Chen & B.K. Cui (10)
Wrightoporiopsis Y.C. Dai, Jia J. Chen & B.K. Cui (5)

Hybogasteraceae Jülich
Hybogaster Singer (1)

Peniophoraceae Lotsy
Amylofungus Sheng H. Wu (2)
Asterostroma Massee (19)
Baltazaria Leal-Dutra, Dentinger & G.W. Griff. (4)
Dendrophora (Parmasto) Chamuris (3)
Dichostereum Pilát (11)
Duportella Pat. (13)
Entomocorticium H.S. Whitney, Bandoni & Oberw. (1)
Gloiothele Bres. (12)
Lachnocladium Lév. (40)
Licrostroma P.A. Lemke (1)
Metulodontia Parmasto (1)
Peniophora Cooke (60)
Sceptrulum K.H. Larss. (1)
Scytinostroma Donk (35)
Vararia P. Karst. (50)
Vesiculomyces E. Hagstr. (1)

Russulaceae Lotsy
Boidinia Stalpers & Hjortstam (11)
Gloeopeniophorella Rick (6)
Lactarius Pers. (450)
Lactifluus (Pers.) Roussel (207)
Multifurca Buyck & V. Hofst. (10)
Pseudoxenasma K.H. Larss. & Hjortstam (1)
Russula Pers. (>3000)

Stereaceae Pilát
Acanthobasidium Oberw. (6)
Acanthofungus Sheng H. Wu, Boidin & C.Y. Chien (6)
Acanthophysellum Parmasto (14)
Acanthophysium (Pilát) G. Cunn. (20)
Aleurobotrys Boidin (10)
Aleurodiscus Rabenh. ex J. Schröt. (27)
Aleuromyces Boidin & Gilles (1)
Amylohyphus Ryvarden (1)
Amylosporomyces S. S. Rattan (2)
Confertextum Priyanka & Dhingra (2)
Conferticium Hallenb. (4)
Dextrinocystidium Sheng H. Wu (2)
Gloeocystidiellum Donk (8)

1300
Gloeocystidiopsis Jülich (1)
Gloeomyces Sheng H. Wu (3)
Gloeosoma Bres. (1)
Matula Massee (2)
Megalocystidium Jülich (7)
Neoaleurodiscus Sheng H. Wu (2)
Scotoderma Jülich (1)
Stereum Hill ex Pers. (40)
Xylobolus P. Karst. (10)

Xenasmataceae Oberw.
Xenasma Donk (16)
Xenasmatella Oberw. (14)
Xenosperma Oberw. (4)

Russulales genera incertae sedis


Aleurocystidiellum P.A. Lemke (3)
Dentipellopsis Y.C. Dai & L.W. Zhou (1)
Dichantharellus Corner (2)
Dichopleuropus D. A. Reid (1)
Gloeoasterostroma Rick (1)
Gloeodontia Boidin (8)
Gloeohypochnicium (Parmasto) Hjortstam (2)
Haloaleurodiscus N. Maek., Suhara & K. Kinjo (1)
Laeticutis Audet (1)
Neoalbatrellus Audet (4)
Perplexostereum Ryvarden & S. Tutka (1)
Polypus Audet (1)
Scopulodontia Hjortstam (3)
Scytinostromella Parmasto (6)
Xeroceps Audet (2)

Sebacinales M. Weiss, Selosse, Rexer, A. Urb. & Oberw.


Sebacinaceae K. Wells & Oberw.
Chaetospermum Sacc. (4)
Ditangium P. Karst. (3)
Efibulobasidium K. Wells (1)
Globulisebacina Oberw., Garnica & K. Riess (2)
Helvellosebacina Oberw., Garnica & K. Riess (2)
Paulisebacina Oberw., Garnica & K. Riess (1)
Sebacina Tul. & C. Tul. (17)
Tremelloscypha D.A. Reid (4)

Serendipitaceae M. Weiss, Waller, A. Zuccaro & Selosse


Serendipita P. Roberts (11)

Stereopsidales Sjökvist, E. Larss., B.E. Pfeil & K.H. Larss.


Stereopsidaceae Sjökvist, E. Larss., B.E. Pfeil & K.H. Larss.
Stereopsis D.A. Reid (15)

Thelephorales Corner ex Oberw.


Bankeraceae Donk

1301
Bankera Coker & Beers ex Pouzar (8)
Boletopsis Fayod (10)
Corneroporus T. Hatt. (1)
Hydnellum P. Karst. (39)
Sarcodon Quél. ex P. Karst. (49)

Thelephoraceae Chevall.
Amaurodon J. Schröt (10)
Lenzitopsis Malençon & Bertault (2)
Phellodon P. Karst. (18)
Polyozellus Murrill (1)
Pseudotomentella Svrček (17)
Skepperia Berk. (5)
Thelephora Ehrh. ex Willd. (50)
Tomentella Pers. ex Pat. (100)
Tomentellopsis Hjortstam (8)

Thelephorales genus incertae sedis


Thelephorella P. Karst. (1)

Trechisporales K.H. Larss.


Hydnodontaceae Jülich
Brevicellicium K. H. Larss. & Hjortstam (13)
Dextrinocystis Gilb. & M. Blackw. (2)
Dextrinodontia Hjortstam & Ryvarden (1)
Hydnodon Banker (1)
Litschauerella Oberw. (3)
Luellia K.H. Larss. & Hjortstam (3)
Porpomyces Jülich (1)
Scytinopogon Singer (5)
Sistotremastrum J. Erikss. (6)
Sphaerobasidium Oberw. (3)
Subulicystidium Parmasto (20)
Trechispora P. Karst. (48)
Tubulicium Oberw. (7)

Tremellodendropsidales Vizzini
Tremellodendropsidaceae Jülich
Tremellodendropsis (Corner) D.A. Crawford (8)

Agaricomycetes genera incertae sedis


Akenomyces G. Arnaud ex D. Hornby (1)
Aldridgea Massee (1)
Arthrodochium R.F. Castañeda & W.B. Kendr. (1)
Arualis Katz (1)
Blasiphalia Redhead (1)
Bridgeoporus T.J. Volk, Burds. & Ammirati (2)
Cenangiomyces Dyko & B. Sutton (1)
Ceraceopsis Hjortstam & Ryvarden (1)
Cilicia Fr. (2)
Corticomyces A.I. Romero & S. E. López (1)
Cruciger R. Kirschner & Oberw. (1)

1302
Dendrosporomyces Nawawi, J. Webster & R.A. Davey (1)
Ellula Nag Raj (1)
Fibulocoela Nag Raj (1)
Fibulotaeniella Marvanová & Bärl. (1)
Geotrichopsis Tzean & Estey (1)
Gloeosynnema Seifert & G. Okada (2)
Glomerulomyces A.I. Romero & S.E. López (1)
Glutinoagger Sivan. & Watling (1)
Hallenbergia Dhingra & Priyanka (1)
Heteroacanthella Oberw. (3)
Intextomyces J. Erikss. & Ryvarden (4)
Korupella Hjortstam & P. Roberts (1)
Loreleia Redhead, Moncalvo, Vilgalys & Lutzoni (3)
Minostroscyta Hjortstam & Ryvarden (1)
Mylittopsis Pat. (1)
Myriococcum Fr. (1)
Odonticium Parmasto (7)
Pagidospora Drechsler (1)
Phlyctibasidium Jülich (1)
Purpureocorticium S.H. Wu (1)
Pycnovellomyces R.F. Castañeda (1)
Riessia Fresen. (5)
Riessiella Jülich (2)
Taiwanoporia T.T. Chang & W.N. Chou (1)
Titaeella G. Arnaud ex K. Ando & Tubaki (1)
Trechinothus E.C. Martini & Trichiès (1)
Trimitiella Dhingra (1)
Tubulicrinopsis Hjortstam & Kotir. (4)
Xerotus Fr. (4)

Bartheletiomycetes Thines
Bartheletiales Thines
Bartheletiaceae R. Bauer, Scheuer, M. Lutz & Grube
Bartheletia G. Arnaud ex Scheuer, R. Bauer, M. Lutz, Stabenth., Melnik & Grube (1)

Dacrymycetes Doweld
Dacrymycetales Henn.
Cerinomycetaceae Jülich
Cerinomyces G. W. Martin (13)

Dacrymycetaceae J. Schröt.
Calocera (Fr.) Fr. (18)
Cerinosterus R.T. Moore (1)
Dacrymyces Nees (50)
Dacryonaema Nannf. (1)
Dacryopinax G.W. Martin (24)
Dacryoscyphus R. Kirschner & Zhu L. Yang (1)
Ditiola Fr. (10)
Femsjonia Fr. (7)
Guepiniopsis Pat. (8)
Heterotextus Lloyd (6)

1303
Unilacrymales Shirouzu, Tokum. & Oberw.
Unilacrymaceae Shirouzu, Tokum. & Oberw.
Unilacryma Shirouzu, Tokum. & Oberw. (1)

Tremellomycetes Doweld
Cystofilobasidiales Fell, Roeijmans & Boekhout
Cystofilobasidiaceae K. Wells & Bandoni
Cystofilobasidium Oberw. & Bandoni (8)

Mrakiaceae X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout


Itersonilia Derx (3)
Krasilnikovozyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (3)
Mrakia Y. Yamada & Komag. (12)
Phaffia M.W. Mill., Yoney. & Soneda (1)
Tausonia Babeva (3)
Udeniomyces Nakase & Takem. (4)
Vustinia Kachalkin, Turchetti & Yurkov (1)

Filobasidiales Jülich
Filobasidiaceae L.S. Olive
Filobasidium L.S. Olive (9)
Goffeauzyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (6)
Heterocephalacria Berthier (8)
Naganishia S. Goto (8)
Syzygospora G.W. Martin (2)

Piskurozymaceae X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout


Piskurozyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (12)
Solicoccozyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (7)

Holtermanniales Libkind, Wuczk., Turchetti & Boekhout


Holtermanniaceae Redhead
Holtermannia Sacc. & Traverso (8)
Holtermanniella Libkind, Wuczk., Turchetti & Boekhout (5)

Tremellales Fr.
Bulleraceae X. Zh. Liu, F.Y. Bai, M. Groenew. & Boekhout
Bullera Derx (4)
Fonsecazyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (3)
Genolevuria X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (4)
Pseudotremella X.Z. Liu, F.Y. Bai, A.M. Yurkov, M. Groenew. & Boekhout (4)

Bulleribasidiaceae X. Z. Liu, F.Y. Bai, M. Groenew. & Boekhout


Bulleribasidium J.P. Samp., M. Weiss & R. Bauer (11)
Derxomyces F.Y. Bai & Q.M. Wang (24)
Dioszegia Zsolt (18)
Hannaella F.Y. Bai & Q.M. Wang (11)
Nielozyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (2)
Vishniacozyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (11)

Carcinomycetaceae Oberw. & Bandoni


Carcinomyces Oberw. & Bandoni (3)

1304
Cryptococcaceae Kütz. ex Castell. & Chalm.
Cryptococcus Vuill. (12)
Kwoniella Statzell & Fell (14)

Cuniculitremaceae J.P. Samp., R. Kirschner & M. Weiss


Fellomyces Y. Yamada & I. Banno (4)
Kockovaella Nakase, I. Banno & Y. Yamada (19)
Sterigmatosporidium G. Kraep. & U. Schulze (1)

Naemateliaceae X. Z. Liu, F. Y. Bai, M. Groenew. & Boekhout


Dimennazyma X. Z. Liu, F. Y. Bai, M. Groenew. & Boekhout (1)
Naematelia Fr. (4)

Phaeotremellaceae A.M. Yurkov & Boekhout


Gelidatrema A.M. Yurkov, X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (1)
Phaeotremella Rea (11)

Phragmoxenidiaceae Oberw. & R. Bauer


Phragmoxenidium Oberw. (1)

Rhynchogastremaceae Oberw. & B. Metzler


Papiliotrema J.P. Samp., M. Weiss & R. Bauer (30)
Rhynchogastrema B. Metzler & Oberw. (9)
Tetragoniomyces Oberw. & Bandoni (1)

Sirobasidiaceae Lindau
Fibulobasidium Bandoni (3)

Tremellaceae Fr.
Hormomyces Bonord. (6)
Mycocryptococcus Pollacci & Nann. (1)
Tremella Pers. (>500)

Trimorphomycetaceae X. Z. Liu, F.Y. Bai, M. Groenew. & Boekhout


Carlosrosaea A.M. Yurkov, X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (3)
Saitozyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (4)
Sugitazyma A.M. Yurkov, X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (1)
Trimorphomyces Bandoni & Oberw. (2)

Tremellales genera incertae sedis


Biatoropsis Räsänen (4)
Dictyotremella Kobayasi (1)
Neotremella Lowy (1)
Sigmogloea Bandoni & J.C. Krug (1)
Sirobasidium Lagerh. & Pat. (8)
Sirotrema Bandoni (3)
Tremellina Bandoni (1)
Xenolachne D.P. Rogers (2)

Trichosporonales Boekhout & Fell


Tetragoniomycetaceae Oberw. & Bandoni
Bandonia A.M. Yurkov, X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (1)

1305
Cryptotrichosporon Okoli & Boekhout (5)
Takashimella Q.M. Wang (4)

Trichosporonaceae Nann.
Apiotrichum Stautz (21)
Cutaneotrichosporon X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (15)
Effuseotrichosporon A.M. Yurkov, X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (1)
Haglerozyma X.Z. Liu, F.Y. Bai, M. Groenew. & Boekhout (1)
Pascua Takashima, Manabe, Nishimura, Sriswasdi, Ohkuma, Iwasaki & Sugita (1)
Prillingera Takashima, Manabe, Nishimura, Sriswasdi, Ohkuma, Iwasaki & Sugita (1)
Trichosporon Behrend (12)
Vanrija R. T. Moore (9)

Tremellomycetes genera incertae sedis


Heteromycophaga P. Roberts (2)
Phyllopta (Fr.) Fr. (1)
Trichosporonoides Haskins & J.F.T. Spencer (6)

Pucciniomycotina R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Agaricostilbomycetes R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.
Agaricostilbales Oberw. & R. Bauer
Agaricostilbaceae Oberw. & R. Bauer
Agaricostilbum J.E. Wright (4)
Pseudobensingtonia F.Y. Bai, Q.M. Wang, M. Groenewald & Boekhout (2)
Sterigmatomyces Fell (5)

Chionosphaeraceae Oberw. & Bandoni


Ballistosporomyces Nakase, G. Okada & Sugiy. (4)
Chionosphaera D.E. Cox (6)
Cystobasidiopsis R. Bauer, B. Metzler, Begerow & Oberw. (3)
Kurtzmanomyces Y. Yamada, Itoh, H. Kawas., I. Banno & Nakase (4)
Stilbum Tode (10)

Kondoaceae R. Bauer, Begerow, J. P. Samp., M. Weiss & Oberw.


Bensingtonia Ingold (5)
Kondoa Y. Yamada, Nakagawa & I. Banno (10)

Ruineniaceae Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout


Ruinenia Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (5)

Agaricostilbales genera incertae sedis


Jianyunia Q. M. Wang, F. Y. Bai, M. Groenew. & Boekhout (1)
Mycogloea L. S. Olive (7)

Atractiellomycetes R. Bauer, Begerow, J. P. Samp., M. Weiss & Oberw.


Atractiellales Oberw. & Bandoni
Atractogloeaceae Oberw. & R. Bauer
Atractogloea Oberw. & Bandoni (1)

Hoehnelomycetaceae Jülich
Basidiopycnis Oberw., R. Kirschner, R. Bauer, Begerow & Arenal (1)

1306
Proceropycnis M. Villarreal, Arenal, V. Rubio, Begerow, R. Bauer, R. Kirschner & Oberw.
(2)

Phleogenaceae Gäum.
Atractidochium Oono, Urbina & Aime (1)
Atractiella Sacc. (7)
Bourdotigloea Aime (9)
Helicogloea Pat. (25)
Hobsonia Berk. ex Massee (2)
Phleogena Link (1)
Saccosoma Spirin (9)

Classiculomycetes R. Bauer, Begerow, J. P. Samp., M. Weiss & Oberw.


Classiculales R. Bauer, Begerow, Oberw. & Marvanová
Classiculaceae R. Bauer, Begerow, Oberw. & Marvanová
Classicula R. Bauer, Begerow, Oberw. & Marvanová (2)
Jaculispora H. J. Huds. & Ingold (1)

Cryptomycocolacomycetes R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Cryptomycocolacales Oberw. & R. Bauer
Cryptomycocolacaceae Oberw. & R. Bauer
Colacosiphon R. Kirschner, R. Bauer & Oberw. (1)
Cryptomycocolax Oberw. & R. Bauer (1)

Cystobasidiomycetes R. Bauer, Begerow, J. P. Samp., M. Weiss & Oberw.


Buckleyzymales R.L. Zhao & K.D. Hyde
Buckleyzymaceae Q. M. Wang, F.Y. Bai, M. Groenew. & Boekhout
Buckleyzyma Q. M. Wang, F.Y. Bai, M. Groenew. & Boekhout (5)

Cystobasidiales R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Cystobasidiaceae Gäum.
Cystobasidium (Lagerh.) Neuhoff (20)
Halobasidium Z. Guo, Y.R. Wang, Q.C. Hou, W.C. Li, H. J. Zhao, Z. H. Sun & Z.D. Zhang
(1)
Occultifur Oberw. (?9)

Erythrobasidiales R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Erythrobasidiaceae Denchev
Bannoa Hamam. (4)
Erythrobasidium Hamam, Sugiy. & Komag. (3)

Erythrobasidiales genera incertae sedis


Cyphobasidium Millanes, Diederich & Wedin (2)
Cyrenella Goch. (1)
Hasegawazyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (1)

Naohideales R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Naohideaceae Denchev
Naohidea Oberw. (1)

Sakaguchiales R.L. Zhao & K. D. Hyde


Sakaguchiaceae Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout

1307
Sakaguchia Y. Yamada, K. Maeda & Mikata (5)

Cystobasidiomycetes families incertae sedis


Microsporomycetaceae Q.M. Wang, F. Y. Bai, M. Groenew. & Boekhout
Microsporomyces Q.M. Wang, F. Y. Bai, M. Groenew. & Boekhout (4)

Symmetrosporaceae Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout


Symmetrospora Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (10)

Cystobasidiomycetes genus incertae sedis


Queiroziella C.R. Félix, J.D.P. Bezerra, R.P. Neves & Landell (1)

Microbotryomycetes R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Heterogastridiales Oberw. & R. Bauer
Heterogastridiaceae Oberw. & R. Bauer
Hyalopycnis Höhn. (1)
Krieglsteinera Pouzar (1)
Pycnopulvinus Toome & Aime (1)

Kriegeriales Toome & Aime


Camptobasidiaceae R.T. Moore
Camptobasidium Marvanová & Suberkr. (1)
Glaciozyma Turchetti, Connell, Thomas-Hall & Boekhout (4)

Kriegeriaceae Toome & Aime


Kriegeria Bres. (1)
Meredithblackwellia Toome & Aime (1)
Phenoliferia Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (4)
Yamadamyces Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (1)

Leucosporidiales Sampaio, M. Weiss & Bauer


Leucosporidiaceae Sampaio, M. Weiss & Bauer
Leucosporidium Fell, Statzell, I.L. Hunter & Phaff (11)

Microbotryales R. Bauer & Oberw.


Microbotryaceae R.T. Moore
Bauerago Vánky (9)
Microbotryum Lév. (100)
Sphacelotheca de Bary (50)
Zundeliomyces Vánky (1)

Ustilentylomataceae R. Bauer & Oberw.


Aurantiosporium M. Piepenbr., Vánky & Oberw. (4)
Fulvisporium Vánky (1)
Microbotryozyma S.O. Suh, D.A. Maslov, Molestina & J.J. Zhou (1)
Ustilentyloma Savile (4)

Sporidiobolales Doweld
Sporidiobolaceae R.T. Moore
Rhodosporidiobolus Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (11)
Rhodotorula F.C. Harrison (15)
Sporobolomyces Kluyver & C.B. Niel (10)

1308
Microbotryomycetes families incertae sedis
Chrysozymaceae Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout
Bannozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (2)
Chrysozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (2)
Fellozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (1)
Hamamotoa Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (4)

Colacogloeaceae Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout


Colacogloea Oberw. & Bandoni (13)

Microbotryomycetes genera incertae sedis


Atractocolax R. Kirschner, R. Bauer & Oberw. (1)
Curvibasidium Samp. & Golubev (3)
Heitmania X.Z. Liu, F.Y. Bai, M. Groenew. & T. Boekhout (3)
Libkindia Mašínová, A. Pontes, J.P. Samp. & Baldrian (1)
Oberwinklerozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (3)
Pseudohyphozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (3)
Pseudoleucosporidium V. de Garcia, M.A. Coelho, T. Maia, L.H. Rosa, A.B.M. Vaz, C.A.
Rosa, J.P. Samp., P. Gonç., M.R. Van Broock & Libkind (1)
Sampaiozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (2)
Slooffia Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (4)
Spencerozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (1)
Trigonosporomyces Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (1)
Udeniozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (1)
Vonarxula Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (1)
Yunzhangia Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (2)
Yurkovia Mašínová, A. Pontes, J.P. Samp. & Baldrian (1)

Mixiomycetes R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Mixiales R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.
Mixiaceae C.L. Kramer
Mixia C.L. Kramer (1)

Pucciniomycetes R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Helicobasidiales R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.
Helicobasidiaceae P.M. Kirk
Helicobasidium Pat. (6)
Tuberculina Tode ex Sacc. (26)

Pachnocybales R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.


Pachnocybaceae Oberw. & R. Bauer
Pachnocybe Berk. (1)

Platygloeales R.T. Moore


Eocronartiaceae Jülich
Eocronartium G.F. Atk. (1)
Herpobasidium Lind (6)
Jola Möller (1)
Platycarpa Couch (2)
Ptechetelium Oberw. & Bandoni (1)

1309
Platygloeaceae Racib.
Glomerogloea Doweld (1)
Glomopsis D.M. Hend. (2)
Insolibasidium Oberw. & Bandoni (1)
Platygloea J. Schröt. (16)

Pucciniales Clem. & Shear


Chaconiaceae Cummins & Y. Hirats.
Achrotelium Syd. (5)
Aplopsora Mains (6)
Botryorhiza Whetzel & Olive (1)
Ceraceopsora Kakish., T. Sato & S. Sato (1)
Chaconia Juel (12)
Goplana Racib. (13)
Maravalia Arthur (41)
Olivea Arthur (8)
Telomapea G.F. Laundon (1)

Coleosporiaceae Dietel
Ceropsora B.K. Bakshi & Suj. Singh (1)
Chrysomyxa Unger (38)
Coleosporium Lév. (125)
Diaphanopellis P.E. Crane (2)
Gallowaya Arthur (3)

Cronartiaceae Dietel
Cronartium Fr. (34)
Endocronartium Y. Hirats. (2)
Peridermium (Link) J.C. Schmidt & Kunze (50)

Melampsoraceae Dietel
Melampsora Castagne (100)

Mikronegeriaceae Cummins & Y. Hirats.


Blastospora Dietel (5)
Chrysocelis Lagerh. & Dietel (5)
Mikronegeria Dietel (3)

Phakopsoraceae Cummins & Hirats. f.


Aeciure Buriticá & J.F. Hennen (1)
Arthuria H.S. Jacks. (6)
Cerotelium Arthur (27)
Crossopsora Syd. & P. Syd. (16)
Dasturella Mundk. & Khesw. (3)
Kweilingia Teng (4)
Macabuna Buriticá & J.F. Hennen (7)
Monosporidium Barclay (3)
Newinia Thaung (3)
Nothoravenelia Dietel (3)
Phakopsora Dietel (116)
Phragmidiella Henn. (8)
Pucciniostele Tranzschel & K.L. Kom. (4)

1310
Scalarispora Buriticá & J.F. Hennen (1)
Uredopeltis Henn. (7)

Phragmidiaceae Corda
Arthuriomyces Cummins & Y. Hirats. (3)
Campanulospora Salazar-Yepes, Pardo-Card. & Buriticá (1)
Gerwasia Racib. (19)
Gymnoconia Lagerh. (4)
Hamaspora Körn. (15)
Joerstadia Gjaerum & Cummins (4)
Kuehneola Magnus (22)
Morispora Salazar-Yepes, Pardo-Card. & Buriticá (1)
Phragmidium Link (100)
Physonema Lév. (1)
Scutelliformis Salazar-Yepes, Pardo-Card. & Buriticá (1)
Trachyspora Fuckel (5)
Xenodochus Schltdl. (2)

Pileolariaceae Cummins & Y. Hirats.


Atelocauda Arthur & Cummins (3)
Pileolaria Castagne (16)
Skierka Racib. (13)
Uromycladium McAlpine (11)

Pucciniaceae Chevall.
Allodus Arthur (1)
Chrysella Syd. (1)
Chrysocyclus Syd. (3)
Chrysopsora Lagerh. (1)
Cleptomyces Arthur (1)
Coleopucciniella Hara ex Hirats. (2)
Corbulopsora Cummins (3)
Cumminsiella Arthur (8)
Cystopsora E.J. Butler (2)
Endophyllum Lév. (43)
Gymnosporangium R. Hedw. ex DC.(64)
Kernella Thirum. (1)
Miyagia Miyabe ex Syd. & P. Syd. (3)
Polioma Arthur (5)
Puccinia Pers. (3300)
Ramakrishnania Ramachar & Bhagyan. (1)
Roestelia Rebent. (15)
Stereostratum Magnus (1)
Uromyces (Link) Unger (1500)
Xenostele Syd. & P. Syd. (4)
Zaghouania Pat. (2)

Pucciniastraceae Gäum. ex Leppik


Hyalopsora Magnus (21)
Melampsorella J. Schröt. (2)
Melampsoridium Kleb. (11)
Milesia F.B. White 1878 (20)

1311
Milesina Magnus (65)
Naohidemyces S. Sato, Katsuya & Y. Hirats. (2)
Peridiopsora Kamat & Sathe (2)
Pucciniastrum G.H. Otth (50)
Thekopsora Magnus (7)
Uredinopsis Magnus (30)

Pucciniosiraceae Cummins & Y. Hirats.


Alveolaria Lagerh. (2)
Baeodromus Arthur (6)
Ceratocoma Buriticá & J.F. Hennen (1)
Chardoniella F. Kern (4)
Cionothrix Arthur (5)
Didymopsora Dietel (6)
Dietelia Henn. (13)
Gambleola Massee (1)
Pucciniosira Lagerh. (17)
Trichopsora Lagerh. (1)

Raveneliaceae Leppik
Allotelium Syd. (1)
Anthomyces Dietel (1)
Anthomycetella Syd. & P. Syd. (1)
Apra J.F. Hennen & F.O. Freire (1)
Bibulocystis J. Walker, Beilharz, Pascoe & Priest (3)
Cumminsina Petr. (1)
Cystomyces Syd. (1)
Diabole Arthur (1)
Diabolidium Berndt (1)
Dicheirinia Arthur (14)
Diorchidiella J.C. Lindq. (2)
Diorchidium Kalchbr. (20)
Endoraecium Hodges & D.E. Gardner (22)
Esalque J.F. Hennen, Figueiredo & A.A. Carvalho (1)
Hapalophragmium Syd. & P. Syd. (18)
Kernkampella Rajendren (8)
Lipocystis Cummins (1)
Nyssopsora Arthur (11)
Ravenelia Berk. (250)
Sphenospora Dietel (6)
Spumula Mains (7)
Triphragmiopsis Naumov (3)
Triphragmium Link (7)
Ypsilospora Cummins (3)

Sphaerophragmiaceae Cummins & Y. Hirats.


Austropuccinia Beenken (1)
Sphaerophragmium Magnus (24)

Uncolaceae Buriticá
Calidion Syd. & P. Syd. (4)
Uncol Buriticá & P.A. Rodr. (1)

1312
Uropyxidaceae (P. Syd. & Syd.) Cummins & Y. Hirats.
Canasta A.A. Carvalho & J.F. Hennen (3)
Dasyspora Berk. & M.A. Curtis (13)
Didymopsorella Thirum. (2)
Dipyxis Cummins & J.W. Baxter (2)
Kimuromyces Dianese, L.T.P. Santos, R.B. Medeiros & Furlan. (1)
Leucotelium Tranzschel (3)
Macruropyxis Azbukina (2)
Mimema H.S. Jacks. (1)
Ochropsora Dietel (3)
Phragmopyxis Dietel (4)
Poliomopsis A.W. Ramaley (1)
Porotenus Viégas (7)
Prospodium Arthur (84)
Sorataea Syd. (8)
Tranzschelia Arthur (19)
Uropyxis J. Schröt. (15)

Pucciniales genera incertae sedis


Aecidiconium Vuill. (1)
Aecidiolum Unger (12)
Aecidium Pers. (ca. 800)
Caeoma Link (ca. 50)
Caetea Salazar-Yepes & A.A. Carvalho (1)
Cerradoa J.F. Hennen & Y. Ono (1)
Coleopuccinia Pat. (1)
Desmella Syd. & P. Syd. (4)
Desmellopsis J.M. Yen (1)
Desmosorus Ritschel, Oberw. & Berndt (1)
Edythea H.S. Jacks. (5)
Elateraecium Thirum., F. Kern & B.V. Patil (3)
Flaminia Sacc. & P. Syd. (1)
Hemileia Berk. & Broome (55)
Hennenia Buriticá (1)
Intrapes J.F. Hennen & Figueiredo (1)
Masseeëlla Dietel (6)
Mehtamyces Mundk. & Thirum. (1)
Phragmotelium Syd. (10)
Puccorchidium Beenken (2)
Schroeteriaster Magnus (4)
Sphenorchidium Beenken (2)
Uraecium Arthur (12)
Uredo Pers. (600)

Septobasidiales Couch ex Donk


Septobasidiaceae Racib.
Aphelariopsis Jülich (2)
Auriculoscypha D.A. Reid & Manim. (1)
Coccidiodictyon Oberw. (1)
Johncouchia S. Hughes & Cavalc. (1)
Septobasidium Pat. (200)
Uredinella Couch (2)

1313
Spiculogloeomycetes Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout
Spiculogloeales R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.
Spiculogloeaceae Denchev
Phyllozyma Q.M. Wang, F.Y. Bai, M. Groenew. & Boekhout (7)
Spiculogloea P. Roberts (5)

Tritirachiomycetes Aime & Schell


Tritirachiales Aime & Schell
Tritirachiaceae Aime & Schell
Tritirachium Limber (4)
Paratritirachium Beguin, Pyck & Detandt (2)

Pucciniomycotina genera incertae sedis


Kryptastrina Oberw (1)
Paraphelaria Corner (2)
Zygogloea P. Roberts (1)

Ustilaginomycotina Doweld
Exobasidiomycetes Begerow, M. Stoll & R. Bauer
Ceraceosorales Begerow, M. Stoll & R. Bauer
Ceraceosoraceae Denchev & R.T. Moore
Ceraceosorus B.K. Bakshi (3)

Doassansiales R. Bauer & Oberw.


Doassansiaceae R.T. Moore ex P.M. Kirk, P.F. Cannon & J.C. David
Burrillia Setch. (4)
Doassansia Cornu (12)
Doassinga Vánky, R. Bauer & Begerow (1)
Entylomaster Vánky & R.G. Shivas (2)
Heterodoassansia Vánky (8)
Nannfeldtiomyces Vánky (2)
Narasimhania Thirum. & Pavgi (1)
Pseudodermatosorus Vánky (2)
Pseudodoassansia (Setch.) Vánky (2)
Pseudotracya Vánky (1)
Tracya Syd. & P. Syd. (2)

Melaniellaceae R. Bauer, Vánky, Begerow & Oberw.


Melaniella R. Bauer, Vánky, Begerow & Oberw. (2)

Rhamphosporaceae R. Bauer & Oberw.


Rhamphospora D.D. Cunn. (2)

Entylomatales R. Bauer & Oberw.


Entylomataceae R. Bauer & Oberw.
Entyloma de Bary (163)
Tilletiopsis Derx (3)

Exobasidiales Henn.
Brachybasidiaceae Gäum.
Brachybasidium Gäum. (1)
Dicellomyces L.S. Olive (4)

1314
Kordyana Racib. (8)
Meira Boekhout, Scorzetti, Gerson & Sztejnb. (4)
Proliferobasidium J.L. Cunn. (1)

Cryptobasidiaceae Malençon ex Donk


Acaromyces Boekhout, Scorzetti, Gerson & Sztejnb. (1)
Botryoconis Syd. & P.Syd. (2)
Clinoconidium Pat. (6)
Coniodictyum Har. & Pat. (1)
Drepanoconis J. Schröt. & Henn. (3)
Phacellula Syd. (1)

Exobasidiaceae J. Schröt.
Arcticomyces Savile (1)
Austrobasidium Palfner (1)
Exobasidium Woronin (51)
Muribasidiospora Kamat & Rajendren (3)

Graphiolaceae Clem. & Shear


Graphiola Poit. (12)
Stylina Syd. & P. Syd. (1)

Laurobasidiaceae Pinruan, Sommai, Suetrong, Somrith. & E.B.G. Jones


Laurobasidium Jülich (2)

Georgefischeriales R. Bauer, Begerow & Oberw.


Eballistraceae R. Bauer, Begerow, A. Nagler & Oberw.
Eballistra R. Bauer, Begerow, A. Nagler & Oberw. (4)

Georgefischeriaceae R. Bauer, Begerow & Oberw.


Georgefischeria Thirum. & Naras. (4)
Jamesdicksonia Thirum., Pavgi & Payak (20)

Gjaerumiaceae R. Bauer, M. Lutz & Oberw.


Gjaerumia R. Bauer, M. Lutz & Oberw. (3)

Tilletiariaceae R.T. Moore


Phragmotaenium R. Bauer, Begerow, A. Nagler & Oberw. (5)
Tilletiaria Bandoni & B.N. Johri (1)
Tolyposporella G.F. Atk. (6)

Golubeviales Q.M. Wang, Begerow, F.Y. Bai & Boekhout


Golubeviaceae Q.M. Wang, F.Y. Bai, Begerow & Boekhout
Golubevia Q.M. Wang, F.Y. Bai, Begerow & Boekhout (1)

Microstromatales R. Bauer & Oberw.


Microstromataceae Jülich
Microstroma Niessl (16)

Quambalariaceae Z.W. de Beer, Begerow & R. Bauer


Quambalaria J.A. Simpson (7)

1315
Volvocisporiaceae Begerow, R. Bauer & Oberw.
Volvocisporium Begerow, R. Bauer & Oberw. (2)

Microstromatales genera incertae sedis


Jaminaea Sipiczki & Kajdacsi ex T. Kij. & Aime (4)
Parajaminaea T. Kij. & Aime (2)
Pseudomicrostroma T. Kij. & Aime (3)
Sympodiomycopsis Sugiy., Tokuoka & Komag. (3)

Robbauerales Boekhout, Begerow, Q.M. Wang & F.Y. Bai


Robbaueraceae Boekhout, Begerow, Q.M. Wang & F.Y. Bai
Robbauera Boekhout, Begerow, Q.M. Wang & F.Y. Bai (1)

Tilletiales Kreisel ex R. Bauer & Oberw.


Erratomycetaceae Denchev & T. Denchev
Erratomyces M. Piepenbr. & R. Bauer (5)

Tilletiaceae J. Schröt.
Conidiosporomyces Vánky (3)
Ingoldiomyces Vánky (1)
Neovossia Körn. (1)
Oberwinkleria Vánky & R. Bauer (1)
Salmacisia D.R. Huff & A. Chandra (1)
Tilletia Tul. & C. Tul. (179)

Malasseziomycetes Q.M. Wang & F.Y. Bai


Malasseziales R.T. Moore
Malasseziaceae Denchev & R.T. Moore
Malassezia Baillon (18)

Moniliellomycetes Q.M. Wang, F.Y. Bai & Boekhout


Moniliellales Q.M. Wang, F.Y. Bai & Boekhout
Moniliellaceae Q.M. Wang, F.Y. Bai & Boekhout
Moniliella Stolk & Dakin (15)

Ustilaginomycetes R. Bauer, Oberw. & Vánky


Uleiellales Garnica, K. Riess, M. Schön, H. Butin, M. Lutz, Oberw. & R. Bauer
Uleiellaceae Vánky
Uleiella J. Schröt. (2)

Urocystidales R. Bauer & Oberw.


Doassansiopsidaceae Begerow, R. Bauer & Oberw.
Doassansiopsis (Setch.) Dietel (14)

Fereydouniaceae S. Nasr, Soudi, H.D.T. Nguyen, M. Lutz & Piątek


Fereydounia S. Nasr, M.R. Soudi, H.D.T. Nguyen, M. Lutz & Piątek (1)

Floromycetaceae S. Nasr, Soudi, H.D.T. Nguyen, M. Lutz & Piątek


Antherospora R. Bauer, M. Lutz, Begerow, Piątek & Vánky (12)
Floromyces Vánky, M. Lutz & R. Bauer (1)

Glomosporiaceae Cif.

1316
Thecaphora Fingerh. (61)

Mycosyringaceae R. Bauer & Oberw.


Mycosyrinx Beck (4)

Urocystidaceae Begerow, R. Bauer & Oberw.


Flamingomyces R. Bauer, M. Lutz, Piątek, Vánky & Oberw. (1)
Melanoxa M. Lutz, Vánky & R. Bauer (2)
Melanustilospora Denchev (2)
Mundkurella Thirum. (5)
Urocystis Rabenh. ex Fuckel (166)
Ustacystis Zundel (2)
Vankya Ershad (3)

Ustilaginales G. Winter
Anthracoideaceae Denchev
Anthracoidea Bref. (112)
Cintractia Cornu (13)
Dermatosorus Sawada ex L. Ling (6)
Farysia Racib. (23)
Farysporium Vánky (1)
Heterotolyposporium Vánky (2)
Kuntzeomyces Henn. Ex Sacc. & P. Syd. (2)
Leucocintractia M. Piepenbr., Begerow & Oberw. (4)
Moreaua Liou & H.C. Cheng (39)
Orphanomyces Savile (3)
Pilocintractia Vánky (2)
Planetella Savile (1)
Portalia V. González, Vánky & Platas (1)
Schizonella J. Schröt. (5)
Stegocintractia M. Piepenbr., Begerow & Oberw. (6)
Testicularia Klotzsch (3)
Tolyposporium Woronin ex J. Schröt. (5)
Trichocintractia M. Piepenbr. (1)
Ustanciosporium Vánky (22)

Cintractiellaceae Vánky
Cintractiella Boedijn (2)

Clintamraceae Vánky
Clintamra Cordas & Durán (1)

Geminaginaceae Vánky
Geminago Vánky & R. Bauer (1)

Melanotaeniaceae Begerow, R. Bauer & Oberw.


Exoteliospora R. Bauer, Oberw. & Vánky (1)
Melanotaenium de Bary (9)
Yelsemia J. Walker (4)

Pericladiaceae Vánky
Pericladium Pass. (3)

1317
Ustilaginaceae Tul. & C. Tul.
Anthracocystis Bref. (134)
Macalpinomyces Langdon & Full. (41)
Moesziomyces Vánky (8)
Sporisorium Ehrenb. ex Link (195)
Tranzscheliella Lavrov (17)
Ustilago (Pers.) Roussel (170)

Websdaneaceae Vánky
Restiosporium Vánky (21)
Websdanea Vánky (1)

Ustilaginales genera incertae sedis


Ahmadiago Vánky (1)
Aizoago Vánky (2)
Anomalomyces Vánky, M. Lutz & R.G. Shivas (2)
Bambusiomyces Vánky (1)
Centrolepidosporium R.G. Shivas & Vánky (1)
Dirkmeia F.Y. Bai, Q.M. Wang, Begerow & Boekhout (1)
Eriocaulago Vánky (2)
Eriocortex Vánky & R.G. Shivas (1)
Eriosporium Vánky (2)
Franzpetrakia Thirum. & Pavgi (3)
Kalmanozyma Q.M. Wang, F.Y. Bai, Begerow & Boekhout (3)
Langdonia McTaggart & R.G. Shivas (8)
Melanopsichium Beck (2)
Mycosarcoma Bref (5)
Parvulago R. Bauer, M. Lutz, Piątek, Vánky & Oberw. (1)
Pattersoniomyces Piątek, M. Lutz & C.A. Rosa (1)
Shivasia Vánky, M. Lutz & Piątek (1)
Stollia McTaggart & R.G. Shivas (5)
Triodiomyces McTaggart & R.G. Shivas (6)
Yunchangia L. Guo & B. Xu (1)

Violaceomycetales Albu, Toome & Aime


Violaceomycetaceae Albu, Toome & Aime
Violaceomyces Albu, Toome & Aime (1)

Ustilaginomycetes genus incertae sedis


Capitulocladosporium L.Y. Sun, X. Sun & L.D. Guo (1)

Wallemiomycotina Doweld
Wallemiomycetes Zalar, de Hoog & Schroers
Geminibasidiales H.D.T. Nguyen, N.L. Nick. & Seifert
Geminibasidiaceae H.D.T. Nguyen, N.L. Nick. & Seifert
Basidioascus Matsush. (3)
Geminibasidium H.D.T. Nguyen, N.L. Nick. & Seifert (2)

Wallemiales Zalar, de Hoog & Schroers


Wallemiaceae R.T. Moore
Wallemia Johan-Olsen (8)

1318
Wallemiomycetes genus incertae sedis
Chernovia A.M. Yurkov & Begerow (1)

Basidiomycota genera incertae sedis


Anastomyces W.P. Wu, B. Sutton & Gange (1)
Anguillomyces Marvanová & Bärl. (1)
Arcispora Marvanová & Bärl. (1)
Arrasia Bernicchia, Gorjón & Nakasone (1)
Brevicellopsis Hjortstam & Ryvarden (1)
Celatogloea P. Roberts (1)
Cystogloea P. Roberts (1)
Microstella K. Ando & Tubaki (1)
Neotyphula Wakef. (1)
Radulodontia Hjortstam & Ryvarden (1)
Restilago Vánky (1)

Blastocladiomycota T.Y. James


Blastocladiomycetes Doweld
Blastocladiales H.E. Petersen
Blastocladiaceae H.E. Petersen
Allomyces E.J. Butler (13)
Blastocladia Reinsch (31)
Blastocladiopsis Sparrow (2)

Catenariaceae Couch
Catenophlyctis Karling (2)
Nematoceromyces Doweld (3)

Paraphysodermataceae Doweld
Paraphysoderma Boussiba, Zarka & T.Y. James (1)

Sorochytriaceae Dewel
Sorochytrium Dewel (1)

Blastocladiales genus incertae sedis


Endoblastidium Codreanu (1)

Callimastigales Doweld
Callimastigaceae Fonseca
Callimastix Weissenb. (2)

Catenomycetales Doweld
Catenomycetaceae Doweld
Catenomyces A.M. Hanson (2)

Coelomomycetaceae Couch
Coelomomyces Keilin (66)
Coelomycidium Debais. (2)

Blastocladiomycetes genus incertae sedis


Microallomyces R. Emers. & J.A. Robertson (1)

1319
Physodermatomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.
May, M. Ryberg & Abarenkov
Physodermatales Caval.-Sm.
Physodermataceae Sparrow
Physoderma Wallr. (99)

Calcarisporiellomycota Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Calcarisporiellomycotina Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel,
T.W. May, M. Ryberg & Abarenkov
Calcarisporiellomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.
May, M. Ryberg & Abarenkov
Calcarisporiellales Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.
May, M. Ryberg & Abarenkov
Calcarisporiellaceae Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.
May, M. Ryberg & Abarenkov
Calcarisporiella de Hoog (1)
Echinochlamydosporium X.Z. Jiang, H.Y. Yu, M.C. Xiang, X.Y. Liu & Xing Z. Liu (1)

Caulochytriomycota Doweld
Caulochytriomycetes Doweld
Caulochytriales Doweld
Caulochytriaceae Subram.
Caulochytrium Voos & L.S. Olive (2)

Chytridiomycota Doweld
Chytridiomycetes Caval.-Sm.
Chytridiales Cohn
Asterophlyctaceae Doweld
Asterophlyctis H.E. Petersen (2)
Wheelerophlyctis P.M Letcher, M.J. Powell, W.J. Davis (2)*

Chytridiaceae Nowak.
Chytridium A. Braun (143)
Dendrochytridium Letcher, Longcore & M.J. Powell (1)
Dinochytrium Lesham, Letcher & M.J. Powell (1)
Irineochytrium Letcher, Longcore & M.J. Powell (1)
Polyphlyctis Karling (3)
Zopfochytrium M.J. Powell, Longcore, Letcher (1)*

Chytriomycetaceae Letcher
Avachytrium Vélez & Letcher (1)
Chytriomyces Karling (33)
Entophlyctis A. Fisch. (29)
Fayochytriomyces W.J. Davis, Letcher, Longcore & M.J. Powell (1)
Obelidium Nowak. (3)
Odontochytrium Vélez & Letcher (1)
Pendulichytrium K. Seto & Degawa (1)
Physocladia Sparrow (1)
Podochytrium Pfitzer (7)
Rhizoclosmatium H.E. Petersen (4)
Siphonaria H.E. Petersen (3)

1320
Phlyctochytriaceae Doweld
Phlyctochytrium J. Schröt. (73)

Phlyctorhizaceae Doweld
Phlyctorhiza A.M. Hanson (3)

Pseudorhizidiaceae Doweld
Pseudorhizidium M.J. Powell, Letcher & Longcore (1)

Scherffeliomycetaceae Doweld
Scherffeliomyces Sparrow (4)

Zygorhizidiaceae Doweld
Zygorhizidium Löwenthal (12)

Chytridiales genus incertae sedis


Delfinachytrium Vélez & Letcher (1)

Nephridiophagales Doweld
Nephridiophagaceae R. Radek, Letcher, Wijayaw., P.M. Kirk & K.D. Hyde
Coleospora Gibbs (1)
Nephridiophaga Ivanić (12)
Oryctospora Purrini & Weiser (1)
Peltomyces Léger (1)

Polyphagales Doweld
Polyphagaceae F. Maekawa
Polyphagus Nowak. (15)

Saccopodiales Doweld
Saccopodiaceae Jacz. & P.A. Jacz.
Saccopodium Sorokīn (1)

Chytridiomycetes families incertae sedis


Amoebochytriaceae Doweld
Amoebochytrium Zopf (1)

Sparrowiaceae Doweld
Sparrowia Willoughby (2)

Sphaeromonadaceae Doweld
Sphaeromonas E. Liebet. (6)

Tetrachytriaceae Doweld
Tetrachytrium Sorokīn (1)

Thalassochytriaceae Doweld
Thalassochytrium Nyvall, M. Pedersén & Longcore (1)
Chytridiomycetes genera incertae sedis
Aphanistis Sorokīn (2)
Bertramia Mesnil & Caullery (3)

1321
Blyttiomyces A.F. Bartsch (11)
Canteria Karling (1)
Dangeardia Schröd. (11)
Dangeardiana Valkanov ex A. Batko (4)
Dictyomorpha Mullins (2)
Gamolpidium Vlădescu (2)
Ichthyochytrium Plehn (1)
Loborhiza A.M. Hanson (1)
Macrochytrium Minden (1)
Megachytrium Sparrow (1)
Mitochytridium P.A. Dang. (2)
Mucophilus Plehn (1)
Nowakowskia Borzí (1)
Olpidiaster Pascher (4)
Perolpidium Doweld (2)
Physorhizophidium Scherff. (1)
Plasmophagus De Wild. (3)
Pseudopileum Canter (1)
Rhizidiocystis Sideris (1)
Rhizosiphon Scherff. (4)
Rhopalophlyctis Karling (1)
Riethophlyctis Doweld (1)
Saccomyces Serbinow (2)
Sagittospora Lubinsky (1)
Scherffeliomycopsis Geitler (1)
Schizolpidium Doweld (1)
Septolpidium Sparrow (1)
Septosperma Whiffen ex R.L. Seym. (5)
Solutoparies Whiffen ex W.H. Blackw. & M.J. Powell (1)
Sorokinocystis Doweld (1)
Sporophlyctidium Sparrow (2)
Sporophlyctis Serbinow (2)
Trematophlyctis Pat. (1)
Truittella Karling (1)
Volvorax Doweld (1)
Zygochytrium Sorokīn (1)
Zygophlyctis Doweld (1)

Cladochytriomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Cladochytriales Mozl.-Standr.
Catenochytridiaceae Doweld
Catenochytridium Berdan (6)

Cladochytriaceae J. Schröt.
Cladochytrium Nowak. (51)

Endochytriaceae Sparrow ex D.J.S. Barr


Diplophlyctis J. Schröt. (12)
Endochytrium Sparrow (7)

Nowakowskiellaceae Sparrow ex Mozl.-Standr.

1322
Nowakowskiella J. Schröt. (18)

Septochytriaceae Mozl.-Standr.
Septochytrium Berdan (5)

Cladochytriales genera incertae sedis


Allochytridium D.J.S. Barr & Désauln. (2)
Cylindrochytridium Karling (2)
Nephrochytrium Karling (8)

Lobulomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May, M.


Ryberg & Abarenkov
Lobulomycetales D.R. Simmons
Alogomycetaceae Doweld
Alogomyces D.R. Simmons & Letcher (1)

Lobulomycetaceae D.R. Simmons


Clydaea D.R. Simmons (1)
Cyclopsomyces K. Seto & Degawa (1)
Lobulomyces D.R. Simmons (2)
Maunachytrium D.R. Simmons (1)

Lobulomycetales genus incertae sedis


Algochytrops Doweld (1)

Mesochytriomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Gromochytriales Karpov & Aleoshin
Gromochytriaceae Karpov & Aleoshin
Gromochytrium Karpov & Aleoshin (1)

Mesochytriales Doweld
Mesochytriaceae Doweld
Mesochytrium B.V. Gromov, Mamkaeva & Pljusch (1)

Polychytriomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Polychytriales Longcore & D.R. Simmons
Arkayaceae Doweld
Arkaya Longcore & D.R. Simmons (2)

Polychytriaceae Doweld
Karlingiomyces Sparrow (8)
Lacustromyces Longcore (1)
Neokarlingia Longcore & D.R. Simmons (1)
Polychytrium Ajello (1)

Rhizophydiomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Rhizophydiales Letcher
Alphamycetaceae Letcher
Alphamyces Letcher (1)

1323
Betamyces Letcher (1)
Gammamyces Letcher (1)

Angulomycetaceae Letcher
Angulomyces Letcher (1)

Aquamycetaceae Letcher
Aquamyces Letcher (1)

Batrachochytriaceae Doweld
Batrachochytrium Longcore, Pessier & D.K. Nichols (2)

Collimycetaceae K. Seto & Degawa


Collimyces K. Seto & Degawa (1)

Coralloidiomycetaceae Doweld
Coralloidiomyces Letcher (1)

Dinomycetaceae Karpov & Guillou


Dinomyces Karpov & Guillou (1)

Globomycetaceae Letcher
Globomyces Letcher (1)
Urceomyces Letcher (1)

Gorgonomycetaceae Letcher
Gorgonomyces Letcher (1)

Halomycetaceae Letcher & M.J. Powell


Halomyces Letcher & M.J. Powell (1)
Paludomyces Letcher & M.J. Powell (1)
Paranamyces Letcher & M.J. Powell (1)
Ulkenomyces Letcher & M.J. Powell (1)

Kappamycetaceae Letcher
Kappamyces Letcher & M.J. Powell (1)

Operculomycetaceae Doweld
Operculomyces M.J. Powell, Letcher & Longcore (1)

Pateramycetaceae Letcher
Pateramyces Letcher (1)

Protrudomycetaceae Letcher
Protrudomyces Letcher (1)

Rhizophydiaceae Letcher
Rhizophydium Schenk ex Rabenh. (218)

Staurastromycetaceae S. Van den Wyngaert, K. Seto & K. Rojas


Staurastromyces Van den Wyngaert, K. Seto & K. Rojas (1)

1324
Terramycetaceae Letcher
Boothiomyces Letcher (1)
Terramyces Letcher (1)

Uebelmesseromycetaceae M.J. Powell & Letcher


Uebelmesseromyces M.J. Powell & Letcher (1)

Rhizophydiales genus incertae sedis


Homolaphlyctis Longcore, Letcher & T.Y. James (1)

Rhizophlyctidomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Rhizophlyctidales Letcher
Arizonaphlyctidaceae Letcher
Arizonaphlyctis Letcher (1)

Borealophlyctidaceae Letcher
Borealophlyctis Letcher (2)

Rhizophlyctidaceae H.E. Petersen


Rhizophlyctis A. Fisch. (31)

Sonoraphlyctidaceae Letcher
Sonoraphlyctis Letcher (1)

Spizellomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,


M. Ryberg & Abarenkov
Spizellomycetales D.J.S. Barr
Powellomycetaceae D.R. Simmons
Fimicolochytrium D.R. Simmons & Longcore (2)
Geranomyces D.R. Simmons (4)
Powellomyces Longcore (2)
Thoreauomyces D.R. Simmons & Longcore (1)

Spizellomycetaceae D.J.S. Barr


Barromyces M.J. Powell & Letcher (1)
Brevicalcar Letcher & M.J. Powell (1)
Bulbosomyces Letcher & Longcore (1)
Gaertneriomyces D.J.S. Barr (4)
Gallinipes Letcher & M.J. Powell (3)
Kochiomyces D.J.S. Barr (1)
Spizellomyces D.J.S. Barr (8)
Triparticalcar D.J.S. Barr (2)

Synchytriomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,


M. Ryberg & Abarenkov
Synchytriales Doweld
Synchytriaceae J. Schröt.
Carpenterophlyctis Doweld (2)
Endodesmidium Canter (1)
Johnkarlingia Pavgi & S.L. Singh (1)
Synchytrium de Bary & Woronin (255)

1325
Synchytriales genus incertae sedis
Micromyces P.A. Dang. (19)

Chytridiomycota genera incertae sedis


Achlyella Lagerh. (1)
Coenomyces K.N. Deckenb. (1)
Achlyogeton Schenk (4)

Entomophthoromycota Humber
Entomophthoromycotina Humber
Entomophthoromycetes Humber
Entomophthorales G. Winter
Ancylistaceae J. Schröt.
Ancylistes Pfitzer (6)
Conidiobolus Bref. (54)
Macrobiotophthora Reukauf (2)

Completoriaceae Humber
Completoria Lohde (1)

Entomophthoraceae Nowak.
Batkoa Humber (10)
Entomophaga A. Batko (22)
Entomophthora Fresen. (63)
Erynia (Nowak. ex A. Batko) Remaud. & Hennebert (27)
Eryniopsis Humber (5)
Furia (A. Batko) Humber (16)
Massospora Peck (15)
Orthomyces Steinkr., Humber & J.B. Oliv. (1)
Strongwellsea A. Batko & J. Weiser (3)
Tarichium Cohn sensu stricto (26)
Zoophthora A. Batko (38)

Meristacraceae Humber
Meristacrum Drechsler (= Tabanomyces Couch, R.J. Andrejeva, Laird & Nolan) (2)

Neozygitomycetes Humber
Neozygitales Humber
Neozygitaceae Ben Ze'ev, R.G. Kenneth & Uziel
Apterivorax S. Keller (2)
Neozygites Witlaczil (22)
Tarichium Cohn pro parte (27)
Thaxterosporium Ben Ze’ev & R.G. Kenneth (1)

Entorrhizomycota R. Bauer, Garnica, Oberw., Riess, Weiß & Begerow


Entorrhizomycetes Begerow, M. Stoll & R. Bauer
Entorrhizales R. Bauer & Oberw.
Entorrhizaceae R. Bauer & Oberw.
Entorrhiza C.A. Weber (c.15)

Talbotiomycetales K. Riess, R. Bauer, R. Kellner, Kemler, Piątek, Vánky & Begerow

1326
Talbotiomycetaceae K. Riess, R. Bauer, R. Kellner, Kemler, Piątek, Vánky & Begerow
Talbotiomyces Vánky, R. Bauer & Begerow (1)

Glomeromycota C. Walker & A. Schüssler


Archaeosporomycetes Sieverd., G.A. Silva, B.T. Goto & Oehl
Archaeosporales C. Walker & A. Schüssler
Ambisporaceae C. Walker, Vestberg & A. Schüssler (= Appendicisporaceae C. Walker, Vestberg
& A. Schüssler)
Ambispora C. Walker, Vestberg & A. Schüssler (basionym Appendicispora Spain, Oehl &
Sieverding) (11)

Archaeosporaceae J.B. Morton & D. Redecker


Archaeospora J.B. Morton & D. Redecker (6)
Intraspora Oehl & Sieverd. (1)
Palaeospora Oehl, Palenz., Sánchez-Castro & G.A. Silva (1)

Geosiphonaceae Engl. & E. Gilg


Geosiphon F. Wettst. (1)

Glomeromycetes Caval.-Sm. emend. Oehl, G.A. Silva, B.T. Goto & Sieverd.
Diversisporales C. Walker & A. Schüssler emend. Oehl, G.A. Silva & Sieverd.
Acaulosporaceae J.B. Morton & Benny
Acaulospora Gerd. & Trappe (= Kuklospora Oehl & Sieverd.) (57)

Diversisporaceae C. Walker & A. Schüssler


Corymbiglomus Błaszk. & Chwat (3)
Desertispora Błaszk., Kozłowska, Ryszka, Al-Yahya’ei & Symanczik (1)
Diversispora C. Walker & A. Schüssler (21)
Otospora Oehl, Palenz. & N. Ferrol (1)
Redeckera C. Walker & A. Schüssler (6)
Sieverdingia Błaszk., Niezgoda & B.T. Goto (1)
Tricispora Oehl, Sieverd., G.A. Silva & Palenz. (1)

Pacisporaceae C. Walker, Błaszk., A. Schüssler & Schwarzott


Pacispora Sieverd. & Oehl (7)

Sacculosporaceae Oehl, Sieverd., G.A. Silva, B.T. Goto, Sánchez-Castro & Palenz.
Sacculospora Oehl, Sieverd., G.A. Silva, B.T. Goto, I.C. Sánchez & Palenz. (2)

Gigasporales S.P. Gautam & U.S. Patel (= Gigasporales Sieverd., G.A. Silva, B.T. Goto & Oehl)
Dentiscutataceae F.A. Souza, Oehl & Sieverd.
Dentiscutata Sieverd., F.A. Souza & Oehl (9)
Fuscutata Oehl, F.A. Souza & Sieverd. (5)
Quatunica F.A. Souza, Sieverd. & Oehl (1)

Gigasporaceae J.B. Morton & Benny


Gigaspora Gerd. & Trappe (7)

Intraornatosporaceae B.T. Goto & Oehl


Intraornatospora B.T. Goto, Oehl & G.A. Silva (1)
Paradentiscutata B.T. Goto, Oehl & G.A. Silva (2)

1327
Racocetraceae Oehl, Sieverd. & F.A. Souza
Cetraspora Oehl, F. A. Souza & Sieverd. (8)
Racocetra Oehl, F.A. Souza & Sieverd. (13)

Scutellosporaceae Sieverd., F.A. Souza & Oehl


Bulbospora Oehl& G.A. Silva (1)
Orbispora Oehl, G.A. Silva & D.K. Silva (2)
Scutellospora C. Walker & F.E. Sanders (10)

Glomerales J.B. Morton & Benny emend. Oehl, G.A. Silva, B.T. Goto & Sieverd.
Entrophosporaceae Oehl & Sieverd.
Albahypha Oehl, G.A. Silva, B.T. Goto & Sieverd. (2)
Claroideoglomus C. Walker & A. Schüssler (6)
Entrophospora R.N. Ames & R.W. Schneid. (2)*

Glomeraceae Piroz. &Dalpé emend. Oehl, G.A. Silva & Sieverd.


Dominikia Błaszk., Chwat &Kovács (11)
Funneliglomus Corazon-Guivin, G.A. Silva & Oehl (1)
Funneliformis C. Walker & A. Schüssler emend. Oehl, G.A. Silva & Sieverd. (11)
Glomus Tul. & C. Tul. emend. Oehl, G.A. Silva & Sieverd. (49)
Halonatospora Błaszk., Niezgoda, B.T. Goto & Kozłowska (1)
Kamienskia Błaszk., Chwat & Kovács (1)
Microdominikia Oehl, Corazon-Guivin & G.A. Silva (1)
Microkamienskia Corazon-Guivin, G.A. Silva & Oehl (3)*
Nanoglomus Corazon-Guivin, G.A. Silva & Oehl (1)
Oehlia Błaszk., Kozłowska, Niezgoda, B.T. Goto & Dalpé (1)
Orientoglomus G.A. Silva, Oehl & Corazon-Guivin (1)
Rhizoglomus Sieverd., G.A. Silva & Oehl (22)*
Sclerocystis Berk. & Broome (8)
Sclerocarpum B.T. Goto, Błaszk., Niezgoda, Kozłowska & Jobim (1)
Septoglomus Sieverd., G.A. Silva & Oehl (13)
Simiglomus Sieverd., G.A. Silva & Oehl (1)
Viscospora Sieverd. Oehl & G.A. Silva (1)

Paraglomeromycetes Oehl, G.A. Silva, B.T. Goto & Sieverd.


Paraglomerales C. Walker & A. Schüssler
Paraglomeraceae J.B. Morton & D. Redecker
Paraglomus J.B. Morton & D. Redecker (8)
Innospora Błaszk., Kovács, Chwat & Kozłowska (1)

Pervetustaceae Błaszk., Chwat, Kozłowska, Symanczik & Al-Yahya’ei


Pervetustus Błaszk., Chwat, Kozłowska, Symanczik & Al-Yahya'ei (1)

Kickxellomycota Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,


M. Ryberg & Abarenkov
Asellariomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,
M. Ryberg & Abarenkov
Asellariales Manier ex Manier & Lichtw.
Asellariaceae Manier ex Manier & Lichtw.
Asellaria R.A. Poiss. (9)

Asellariales genus incertae sedis

1328
Baltomyces Cafaro (1)

Barbatosporomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Barbatosporales Doweld
Barbatosporaceae Doweld
Barbatospora M.M. White, Siri & Lichtw. (1)

Dimargaritomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Dimargaritales R.K. Benj.
Dimargaritaceae R.K. Benj.
Dimargaris Tiegh. (7)
Dispira Tiegh. (4)
Tieghemiomyces R.K. Benj. (2)

Dimargaritales genus incertae sedis


Spinalia Vuill. (1)

Harpellomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,


M. Ryberg & Abarenkov
Harpellales Lichtw. & Manier
Harpellaceae L. Léger & Duboscq ex P.M. Kirk & P.F. Cannon
Carouxella Manier, Rioux & Whisler (2)
Harpella L. Léger & Duboscq (7)
Harpellomyces Lichtw. & S.T. Moss (4)
Klastostachys Lichtw., M.C. Williams & M.M. White (1)
Stachylina L. Léger & M. Gauthier (40)
Stachylinoides Lichtw. & López-Lastra (1)

Legeriomycetaceae Pouzar
Austrosmittium Lichtw. & M.C. Williams (5)
Bactromyces William & Strongman (1)
Baetimyces L.G. Valle & Santam. (1)
Bojamyces Longcore (3)
Capniomyces S.W. Peterson & Lichtw. (3)
Caudomyces Lichtw., Kobayasi & Indoh (3)
Coleopteromyces Ferrington, Lichtw. & López-Lastra (1)
Dacryodiomyces Lichtw. (1)
Ejectosporus S.W. Peterson, Lichtw. & M.C. Williams (1)
Ephemerellomyces M.M. White & Lichtw. (1)
Furculomyces Lichtw. & M.C. Williams (3)
Gauthieromyces Lichtw. (3)
Genistelloides S.W. Peterson, Lichtw. & B.W. Horn (5)
Genistellospora Lichtw. (6)
Glotzia M. Gauthier ex Manier & Lichtw. (7)
Graminella L. Léger & M. Gauthier ex Manier (3)
Laculus William & Strongman (1)
Lancisporomyces Santam. (5)
Legerioides M.M. White (1)
Legeriomyces Pouzar (11)
Legeriosimilis M.C. Williams, Lichtw., M.M. White & J.K. Misra (8)

1329
Orphella L. Léger & M. Gauthier (12)
Pennella Manier (8)
Plecopteromyces Lichtw., Ferrington & López-Lastra (3)
Pseudoharpella Ferrington, M.M. White & Lichtw. (1)
Pteromaktron Whisler (2)
Simuliomyces Lichtw. (1)
Sinotrichium Juan Wang (1)
Smittium R.A. Poiss. (1)
Spartiella Tuzet & Manier ex Manier (3)
Stipella L. Léger & M. Gauthier (2)
Stypomyces Doweld (2)
Tectimyces L.G. Valle & Santam. (3)
Trichozygospora Lichtw. (1)
Trifoliellum Strongman & M.M. White (1)
Zancudomyces Yan Wang, Tretter, Lichtw. & M.M. White (1)
Zygopolaris S.T. Moss, Lichtw. & Manier (2)
Zygopolaropsis Hirok. Sato & Degawa (1)

Harpellales genus incertae sedis


Trissocladomyces Doweld (1)

Kickxellomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,


M. Ryberg & Abarenkov
Kickxellales Kreisel ex R.K. Benj.
Kickxellaceae Linder
Coemansia Tiegh. & G. Le Monn. (25)
Dipsacomyces R.K. Benj. (1)
Kickxella Coem. (1)
Linderina Raper & Fennell (2)
Martensella Coem. (1)
Martensiomyces J.A. Mey. (1)
Mycoemilia Kurihara, Degawa & Tokum. (1)
Myconymphaea Kurihara, Degawa & Tokum. (1)
Pinnaticoemansia Kurihara & Degawa (1)
Spirodactylon R.K. Benj. (1)
Spiromyces R.K. Benj. (2)

Ramicandelaberomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel,


T.W. May, M. Ryberg & Abarenkov
Ramicandelaberales Doweld
Ramicandelaberaceae Doweld
Ramicandelaber Y. Ogawa, S. Hayashi, Degawa & Yaguchi (4)

Kickxellomycotina genera incertae sedis


Aenigmatospora R.F. Castañeda, Saikawa, Guarro & M. Calduch (1)
Ballocephala Drechsler (1)
Zygnemomyces K. Miura (2)

Monoblepharomycota Doweld
Hyaloraphidiomycetes Doweld
Hyaloraphidiales Doweld

1330
Hyaloraphidiaceae Doweld
Hyaloraphidium Korshikov (1)

Monoblepharidomycetes J.H. Schaffn.


Monoblepharidales Sparrow
Gonapodyaceae H.E. Petersen ex P.M. Kirk, P.F. Cannon & J.C. David
Gonapodya A. Fisch. (5)
Monoblepharella Sparrow (5)

Harpochytriaceae Wille
Harpochytrium Lagerh. (12)

Monoblepharidaceae A. Fisch.
Monoblepharis Cornu (15)

Oedogoniomycetaceae D.J.S. Barr


Oedogoniomyces Kobayasi & M. Ôkubo (1)

Telasphaerulaceae Longcore & T.Y. James


Telasphaerula Longcore & T.Y. James (1)

Sanchytriomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Sanchytriales Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May, M.
Ryberg & Abarenkov
Sanchytriaceae Karpov & Aleoshin
Amoeboradix Karpov, López-García, Mamkaeva & Moreira (1)
Sanchytrium Karpov & Aleoshin (1)

Mortierellomycota Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W. May,


M. Ryberg & Abarenkov
Mortierellomycotina Kerst. Hoffm., K. Voigt & P.M. Kirk
Mortierellomycetes Doweld
Mortierellales Caval.-Sm.
Mortierellaceae A. Fisch.
Aquamortierella Embree & Indoh (1)
Dissophora Thaxt. (3)
Gamsiella (R.K. Benj.) Benny & M. Blackw. (1)
Lobosporangium M. Blackw. & Benny (1)
Modicella Kanouse (2)
Mortierella Coem. (112)*

Mucoromycota Doweld
Mucoromycotina Benny
Endogonomycetes Doweld
Endogonales Jacz. & P.A. Jacz.
Densosporaceae Desirò, M.E. Sm., Bidartondo, Trappe & Bonito
Densospora McGee (9)*

Endogonaceae Paol.
Endogone Link (26)
Jimgerdemannia Trappe (2)

1331
Peridiospora C.G. Wu & Suh J. Lin (2)
Sclerogone Warcup (1)
Sphaerocreas Sacc. & Ellis (4)

Mucoromycetes Doweld
Mucorales Fr.
Backusellaceae K. Voigt & P.M. Kirk
Backusella Hesselt. & J.J. Ellis (13)

Choanephoraceae J. Schröt.
Blakeslea Thaxt. (2)
Choanephora Curr. (2)
Gilbertella Hesselt. (2)
Poitrasia P.M. Kirk (1)

Cunninghamellaceae Naumov ex R.K. Benj.


Absidia Tiegh. (20)
Chlamydoabsidia Hesselt. & J.J. Ellis (1)
Cunninghamella Matr. (13)
Gongronella Ribaldi (6)
Halteromyces Shipton & Schipper (1)
Hesseltinella H.P. Upadhyay (1)

Lentamycetaceae K. Voigt & P.M. Kirk


Lentamyces Kerst. Hoffm. & K. Voigt (4)

Lichtheimiaceae Kerst. Hoffm., Walther & K. Voigt


Circinella Tiegh. & G. Le Monn. (11)
Dichotomocladium Benny & R.K. Benj. (5)
Fennellomyces Benny & R.K. Benj. (4)
Lichtheimia Vuill. (7)
Phascolomyces Boedijn ex Benny & R.K. Benj. (1)
Rhizomucor Lucet & Costantin (6)
Thamnostylum Arx & H.P. Upadhyay (4)
Thermomucor Subrahm., B.S. Mehrotra & Thirum. (1)
Zychaea Benny & R.K. Benj. (1)

Mucoraceae Dumort.
Actinomucor Schostak. (1)
Ambomucor R.Y. Zheng & X.Y. Liu (3)
Benjaminiella Arx (3)
Chaetocladium Fresen. (2)
Cokeromyces Shanor (1)
Dicranophora J. Schröt. (1)
Ellisomyces Benny & R.K. Benj. (1)
Helicostylum Corda (2)
Hyphomucor Schipper & Lunn (1)
Isomucor J.I. Souza, Pires-Zottar. & Harakava (2)
Kirkiana L.S. Loh, Kuthub. & Nawawi (1)
Kirkomyces Benny (1)
Mucor Fresen. (91)*
Nawawiella L.S. Loh & Kuthub. (1)

1332
Parasitella Bainier (1)
Pilaira Tiegh. (7 and 1 subspecies)
Pirella Bainier (2)
Rhizopodopsis Boedijn (1)
Thamnidium Link (1)
Tortumyces L.S. Loh (2)

Mycocladaceae Kerst. Hoffm.


Mycocladus Beauverie (1)

Mycotyphaceae Benny & R.K. Benj.


Mycotypha Fenner (4)

Phycomycetaceae Arx
Phycomyces Kunze (3)
Spinellus Tiegh. (5)

Pilobolaceae Corda
Pilobolus Tode (10 and 1 subspecies)
Utharomyces Boedijn ex P.M. Kirk & Benny (1 sp. and 1 subspecies)

Radiomycetaceae Hesselt. & J.J. Ellis


Radiomyces Embree (3)

Rhizopodaceae K.voigt & P.M. Kirk


Rhizopus Ehrenb. (13)
Sporodiniella Boedijn (1)
Syzygites Ehrenb. (1)

Saksenaeaceae Hesselt. & J.J. Ellis


Apophysomyces P.C. Misra (5)
Saksenaea S.B. Saksena (5)

Syncephalastraceae Naumov ex R.K. Benj.


Protomycocladus Schipper & Samson (1)
Syncephalastrum J. Schröt. (2)

Umbelopsidomycetes Tedersoo, Sanchez-Ramirez, Kõljalg, Bahram, M. Döring, Schigel, T.W.


May, M. Ryberg & Abarenkov
Umbelopsidales Spatafora & Stajich
Umbelopsidaceae W. Gams & W. Mey.
Umbelopsis Amos & H.L. Barnett (16)

Mucoromycotina genera incertae sedis


Bifiguratus Torr.-Cruz & Porras-Alfaro (1)
Mucorodium K.W. Zaleski (1)
Palaeoendogone Strullu-Derr., Kenrick, Pressel, Duckett, J.P. Rioult & Strullu (1)
Planticonsortium C. Walker & D. Redecker (1)

Mucoromycota genus incertae sedis


Nothadelphia Degawa & W. Gams

1333
Neocallimastigomycota M.J. Powell
Neocallimastigomycetes M.J. Powell
Neocallimastigales J.L. Li, I.B. Heath & L. Packer
Neocallimastigaceae I.B. Heath (= Piromonadaceae Doweld; = Anaeromycetaceae Doweld)
Anaeromyces Breton, Bernalier, Dusser, Fonty, B. Gaillard & J. Guillot (4)
Buwchfawromyces T.M. Callaghan & G.W. Griff. (1)
Caecomyces J.J. Gold (5)
Cyllamyces Ozkose, B.J. Thomas, D.R. Davies, G.W. Griff. & Theodorou (1)
Feramyces Radwa Hanafy, Mostafa Elshahed & Noha Youssef (1)
Liebetanzomyces Joshi, G.W. Griff. & Dagar (1)
Neocallimastix Vávra & Joyon ex I.B. Heath (7)
Oontomyces Dagar (1)
Orpinomyces D.J.S. Barr, H. Kudo, Jakober & K.J. Cheng (2)
Pecoramyces Hanafy, N.H. Youssef, G.W. Griff. & Elshahed (1)
Piromyces J.J. Gold, I.B. Heath & Bauchop (= Piromonas E. Liebet.) (6)

Olpidiomycota Doweld
Olpidiomycetes Doweld
Olpidiales Caval.-Sm.
Olpidiaceae J. Schröt.
Chytridhaema Moniez (1)
Cibdelia Juel (1)
Leiolpidium Doweld (5)
Olpidium (A. Braun) J. Schröt. (ca. 50)

Rozellomycota Doweld
Rudimicrosporea Sprague
Metchnikovellida Vivier
Amphiacanthidae Larsson
Amphiacantha Caullery & Mesnil (3)

Metchnikovellidae Caullery & Mesnil emend. Larsson


Amphiamblys Caullery & Mesnil (7)
Caulleryetta Dogiel (8)
Desportesia Issi & Voronin (1)
Metchnikovella Caullery & Mesnil (8)

Microsporidea Corliss & Levine


Amblyosporida Tokarev & Issi
Amblyosporidae Weiser emend. Tokarev & Issi
Aedispora Kilochitskii (1)
Amblyospora Hazard & Oldacre (90)
Andreanna Simakova, Vossbrinck & Andreadis (1)
Becnelia Tonka & Weiser (1)
Crepidulospora Simakova, Pankova & Issi (1)
Cristulospora Khodzhaeva & Issi (3)
Culicospora Weiser (2)
Culicosporella Weiser (1)
Dimeiospora Simakova, Pankova & Issi (1)
Edhazardia Becnel, V. Sprague & Fukuda (1)
Hyalinocysta Hazard & Oldacre (1)
Intrapredatorus Chen, Kuo & Wu (1)

1334
Novothelohania Andreadis, Simakova, Vossbrinck, Shepard & Yurchenko (1)
Parastempellia Khodzhaeva (2)
Parathelohania Codreanu (25)
Trichoctosporea Larsson (1)
Tricornia Pell & Canning (1)

Caudosporidae Weiser emend. Tokarev & Issi


Binucleospora Bronnvall & Larsson (1)
Caudospora J. Weiser (1)
Flabelliforma Canning, R. Killick-Kendrick & Killick-Kendrick (4)
Myrmecomorba R.M. Plowes, J.J. Becnel, E.G. LeBrun, D.H. Oi, S.M. Valles, N.T. Jones &
L.E. Gilbert (1)
Neoflabelliforma Morris & Freeman (2)
Octosporea Flu (18)
Polydispyrenia Canning & Hazard (2)
Ringueletium Garcia (1)
Scipionospora Bylén & Larsson (1)
Weiseria Doby & Saguez (3)

Gurleyidae Sprague emend. Tokarev & Issi


Agglomerata Larsson& Yan (5)
Binucleata Refardt, Decaestecker, Johnson & Vávra (1)
Conglomerata Vavra, Fiala, Krylova, Petrusek, Hylis (1)
Episeptum Larsson (6)
Gurleya Doflein (10)
Lanatospora Voronin (4)
Larssonia Vidtmann & Sokolova (2)
Marssoniella Lemmermann (1)
Norlevinea Vávra (1)
Paraepiseptum Hyliš, Oborník, Nebesářová & Vávra (4)
Pseudoberwaldia Vavra, Fiala, Krylova, Petrusek, Hylis (1)
Senoma Simakova, Pankova, Tokarev & Issi (1)
Zelenkaia Hyliš, Oborník, Nebesářová & Vávra (1)

Amblyosporida genera incertae sedis


Alfvenia Larsson (4)
Hazardia Weiser (2)
Multilamina Becnel, Scheffrahn, Vossbrinck & Bahder (1)
Takaokaspora Andreadis, Takaoka, Otsuka & Vossbrinck (1)
Trichotuzetia Vávra, Larsson & Baker (1)

Neopereziida Tokarev & Issi


Berwaldiidae Simakova, Tokarev & Issi
Berwaldia Larsson (4)
Fibrillanosema Slothouber Galbreath, Smith, Terry, Becnel & Dunn (1)

Neopereziidae Voronin emend. Issi, Tokarev, Seliverstova & Voronin


Bacillidium Janda (5)
Bryonosema Canning, Refardt, Vossbrinck, Okamura & Curry (2)
Neoperezia Issi & Voronin (2)
Pseudonosema Canning, Refardt, Vossbrinck, Okamura & Curry (1)
Schroedera Morris & Adams (2)

1335
Trichonosema Canning, Refardt, Vossbrinck, Okamura & Curry (2)

Tubulinosematidae Franzen, Fischer, Schröder, Schölmerich & Schneuwly emend. Tokarev &
Issi
Anncaliia Issi, Krylova & Nikolaeva (6)
Kneallhazia Sokolova & Fuxa (2)
Tubulinosema Franzen, Fischer, Schröder, Schölmerich & Schneuwly (5)

Neopereziida genera incertae sedis


Janacekia Larsson (6)
Systenostrema Hazard & Oldacre (5)

Ovavesiculida Tokarev & Issi


Ovavesiculidae Sprague, Becnel & Hazard emend. Tokarev & Issi
Antonospora Fries, Paxton, Tengo, Slemenda, da Silva, & Pieniazek (2)
Ovavesicula Andreadis & Hanula (1)
Paranosema Sokolova, Dolgikh, Morzhina, Nassonova, Issi, Terry, Ironside, Smith (4)

Ovavesiculida genus incertae sedis


Nematocida Troemel, Félix, Whiteman, Barrière & Ausubel (1)

Glugeida Gurley emend. Tokarev & Issi


Facilisporidae Jones, Prosperi-Porta & Kim
Facilispora Jones, Prosperi-Porta & Kim (1)

Glugeidae Gurley emend. Tokarev & Issi


Alloglugea Paperna & Lainson (1)
Amazonspora Azevedo & Matos (1)
Glugea Thélohan (40)
Ichthyosporidium Caullery & Mesnil (5)
Johenrea Lange, Becnel, Razafindratiana, Przybyszewski & Razafindrafara (1)
Loma Morrison & Sprague (12)
Parapleistophora Issi, Kadyrova, Pushkar, Khodzhaeva & Krylova (1)
Pseudoloma J.L. Matthews, A.M.V. Br., K. Larison, J.K. Bishop-Stewart, P. Rogers & M.L.
Kent (6)

Myosporidae Stentiford, Bateman, Small, Moss, Shields, Reece & Tuck


Myospora Stentiford, Bateman, Small, Moss, Shields, Reece & Tuck (1)

Pereziidae Loubes, Maurand, Comps & Campillo emend. Tokarev & Issi
Ameson Sprague (2)
Nadelspora Olson, Tiekotter & Reno (1)
Perezia Léger & Duboscq (12)
Pernicivesicula Bylén & Larsson (1)

Pleistophoridae Doflein emend. Tokarev & Issi


Dasyatispora Diamant, Goren, Yokeş, Galil, Klopman, Huchon, Szitenberg & Karhan (1)
Heterosporis Schubert (4)
Myosporidium Baquero, Rubio, Moura, Pieniazek & Jordana (1)
Ovipleistophora Pekkarinen, Lom & Nilsen (2)
Pleistophora Gurley (10)
Trachipleistophora Hollister, Canning, Weidner, Field, Kench & Marriott (3)

1336
Vavraia Weiser (10)

Spragueidae Weissenberg emend. Tokarev & Issi


Apotaspora Sokolova & Overstreet (1)
Inodosporus Overstreet & Weidner (2)
Microgemma Ralphs & Matthews (6)
Spraguea Weissenberg (2)
Potaspora Casal, Matos, Teles-Grilo & Azevedo (3)
Pseudokabatana Liu, Stentiford, Voronin, Sato, Li& Zhang (1)
Tetramicra Matthews & Matthews (1)

Thelohaniidae Hazard & Oldacre emend. Tokarev & Issi


Bohuslavia Larsson (1)
Chapmanium Hazard & Oldacre (4)
Coccospora Wallr. (1)
Cucumispora Ovcharenko, Bacela, Wilkinson, Ironside, Rigaud & Wattier (2)
Hyperspora Stentiford, Ramilo, Abollo, Kerr, Bateman, Feist, Bass & Villalba (1)
Napamichum Larsson (3)
Nudispora Larsson (1)
Octotetraspora Issi, Kadyrova, Pushkar, Khodzhaeva & Krylova (1)
Ormieresia Vivarès, Bouix & Manier (1)
Orthothelohania Codreanu & Codreanu-Balcescu (1)
Paradoxium Stentiford, Ross, Kerr, Bass & Bateman (1)
Pegmatheca Hazard & Oldacre (2)
Resiomeria Larsson (1)
Spherospora Garcia (1)
Thelohania Henneguy (50)

Unikaryonidae Sprague emend. Tokarev & Issi


Canningia Weiser, Wegensteiner & Žižka (2)
Dictyocoela Terry, Smith, Sharpe, Rigaud, Littlewood, Ironside, Rollinson, Bouchon,
MacNeil, Dick & Dunn (8)
Larssoniella Weiser & David (2)
Unikaryon Canning, Lai & Lie (18)

Glugeida genus incertae sedis


Triwangia Wang, Nai, Chih Wang, Solter, Hsu, Wang & Lo (1)

Nosematida Labbe emend. Tokarev & Issi


Encephalitozoonidae Voronin
Encephalitozoon Levaditi, Nicolau & Schoen (12)
Mockfordia Sokolova, Sokolov & C.E. Carlton (1)

Enterocytozoonidae Cali & Owen emend. Tokarev & Issi


Desmozoon Freeman & Sommerville (3)
Enterocytozoon Desportes, Le Charpentier, Galian, Bernard, Cochand-Priollet, Lavergne,
Ravisse & Modigliani (2)
Enterospora Stentiford, Bateman, Longshaw & Feist (2)
Hepatospora Stentiford, Bateman, Dubuffet, Chambers & Stone (1)
Nucleospora Hedrick, Groff & Baxa (3)
Obruspora Diamant, Rothman, Goren, Galil, Yokes, Szitenberg & Huchon (1)

1337
Heterovesiculidae Lange, Macvean, Henry & Streett
Heterovesicula Lange, Macvean, Henry & Streett (1)

Mrazekiidae Léger & Hesse emend. Tokarev & Issi


Agmasoma Hazard & Oldacre (3)
Anostracospora Rode, Landes, Lievens, Flaven, Segard, Jabbour-Zahab, Michalakis,
Agnew, Vivarés & Lenormand (1)
Euplotespora Fokin, Di Giuseppe, Erra & Dini (1)
Helmichia Larsson (5)
Hrabyeia Lom & Dyková (1)
Jirovecia Weiser (7)
Mrazekia Léger & Hesse (17)
Rectispora Larsson (1)

Nosematidae Tokarev, Huang, Solter, Malysh, Becnel & Vossbrinck


Nosema Nägeli (20)
Vairimorpha Pilley (15)

Ordosporidae Larsson, Ebert & Vávra


Ordospora Larsson, Ebert & Vávra (2)

Nosematida genera incertae sedis


Alternosema Lipa, Tokarev, Issi (1)
Anisofilariata Tokarev, Voronin, Seliverstova, Dolgikh, Pavlova, Ignatieva & Issi (1)
Crispospora Tokarev, Voronin, Seliverstova, Pavlova & Issi (1)
Cystosporogenes Canning, Barker, Nicholas & Page (4)
Endoreticulatus Brooks, Becnel & Kennedy (5)
Enterocytospora Rode, Landes, Lievens, Flaven, Segard, Jabbour-Zahab, Michalakis,
Agnew, Vivarés & Lenormand (1)
Enteropsectra Zhang, Sachse, Prevost, Luallen, Troemel & Felix(2)
Glugoides Larsson, Ebert, Vávra & Voronin (1)
Liebermannia Sokolova, Lange & Fuxa (3)
Orthosomella Canning, Wigley & Barker (2)
Pancytospora Zhang, Sachse, Prevost, Luallen, Troemel & Felix (2)
Parahepatospora Bojko, Clark, Bass, Dunn, Stewart-Clark, Stebbing & Stentiford (1)
Percutemincola Nishikori, Setiamarga, Tanji, Kuroda, Shiraishi & Okashi-Kobayashi (1)
Sporanauta Ardila-Garcia & Fast (1)
Vittaforma Silveira & Canning (1)

Microsporidia families incertae sedis


Abelsporidae Azevedo
Abelspora Azevedo (1)

Areosporiidae Stentiford, Bateman, Feist, Oyarzún, Uribe, Palacios & Stone


Areospora Stentiford, Bateman, Feist, Oyarzún, Uribe, Palacios & Stone (1)

Burenellidae Jouvenaz & Hazard


Burenella Jouvenaz & Hazard (1)
Pilosporella Hazard & Oldacre (2)
Tabanispora Bykova, Sokolova & Issi (2)

Cougourdellidae Poisson

1338
Cougourdella Hesse (7)

Cylindrosporidae Issi & Voronin


Cylindrospora Issi & Voronin (2)

Duboscqiidae R. Sprague
Duboscqia Pérez (11)
Mitoplistophora Codreanu (1)
Pulicispora Vedmed, Krylova & Issi (1)
Tardivesicula Larsson& Bylén (1)
Trichoduboscqia Léger (1)

Golbergiidae Issi
Golbergia Weiser (1)
Krishtalia Kilochitskii (1)
Simuliospora Khodzhaeva, Krylova & Issi (2)

Microfilidae Sprague, Becnel & Hazard


Microfilum Faye, Toguebaye & Bouix (1)

Neonosemoidiidae Faye, Toguebaye & Bouix


Neonosemoides Faye & Toguebaye (4)

Pleistosporidiidae Codreanu-Balcescu & Codreanu


Pleistosporidium Codreanu-Balcescu & Codreanu (1)

Pseudopleistophoridae Sprague
Pseudopleistophora Sprague (1)
Steinhausia Sprague, Ormières & Manier (4)

Striatosporidae Issi & Voronin


Striatospora Issi & Voronin (1)

Telomyxidae Léger & Hesse


Telomyxa Léger & Hesse (4)

Toxoglugeidae Larsson
Toxoglugea Léger & Hesse (15)
Toxospora Voronin (2)

Tuzetiidae Sprague, Tuzet & Maurand


Nelliemelba Larsson (1)
Pankovaia Simakova, Tokarev & Issi (1)
Paratuzetia Poddubnaya, Tokarev & Issi (1)
Tuzetia Maurand, Fize, Vernick & Michel (7)

Microsporidia genera incertae sedis


Auraspora Weiser & Purrini (1)
Baculea Loubès & Akbarieh (1)
Burkea Sprague (2)
Chytridioides Tregouboff (1)
Ciliatosporidium Foissner & Foissner (1)

1339
Cryptosporina Hazard & Oldacre (1)
Evlachovaia Voronin (1)
Geusia Rühl & Korn (1)
Gurleyides Voronin (1)
Hamiltosporidium Haag, Larsson, Refardt & Ebert (2)
Hirsutosporos Batson (1)
Holobispora Voronin (1)
Issia Weiser (3)
Kinorhynchospora Adrianov & Rybakov (1)
Mariona Stempell (1)
Merocinta Pell & Canning (1)
Microsporidium Balbiani (120)#
Myxocystis Mrazek (1)
Nematocenator Sapir, Dillman, Connon, Grupe, Ingels, Mundo-Ocampo, Levin, Bladwin,
Orphan & Sternberg (1)
Nosemoides Vinckier (5)
Pyrotheca Hesse (4)
Spiroglugea Léger & Hesse (1)
Stempellia Léger & Hesse (19)
Wittmannia Czaker (1)

Rozellomycota orders incertae sedis


Chytridiopsidea Weiser
Buxtehudidae Larsson
Jiroveciana Larsson (1)
Buxtehudea Larsson (1)

Chytridiopsidae Sprague, Ormières & Manier


Acarispora Radek and Alberti (1)
Chytridiopsis Schneider (11)
Intexta Larsson, Steiner & Bjørnson (1)
Nolleria Beard, Butler & Becnel (1)
Sheriffia Larsson (1)

Hesseidae Ormières & Sprague


Hessea Ormières & Sprague (1)

Rozellomycota genera incertae sedis


Mitosporidium Haag, James, Pombert, Larsson, Schaer, Refardt & Ebert (2)
Morellospora Corsaro, Walochnik, Venditti, Hauröder & Michel (1)
Nucleophaga Dangeard (2)
Paramicrosporidium Corsaro, Walochnik, Venditti, Steinmann, Müller & Michel (1)
Rozella Cornu (20)
#
Microsporidium is a collective genus which incorporate species with uncertain genus allocation

Zoopagomycota Gryganskyi, M.E. Sm., Spatafora & Stajich


Zoopagomycetes Doweld
Zoopagales Bessey ex R.K. Benj.
Cochlonemataceae Dudd.
Aenigmatomyces R. F. Castañeda & W.B. Kendr. (1)
Amoebophilus P.A. Dang. (4)

1340
Aplectosoma Drechsler (1)
Bdellospora Drechsler (1)
Cochlonema Drechsler (11)
Endocochlus Drechsler (4)
Euryancale Drechsler (4)

Helicocephalidaceae Boedijn
Brachymyces G.L. Barron (1)
Helicocephalum Thaxt. (6)
Rhopalomyces Corda (11)
Verrucocephalum Degawa (1)

Piptocephalidaceae J. Schröt.
Kuzuhaea R.K. Benj. (1)
Piptocephalis de Bary (ca. 25)
Syncephalis Tiegh. & G. Le Monn. (ca. 55)

Sigmoideomycetaceae Benny, R.K. Benj. & P.M. Kirk


Reticulocephalis Benny, R.K. Benj. & P.M. Kirk (2)
Sigmoideomyces Thaxt. (1)
Sphondylocephalum Stalpers (1)
Thamnocephalis Blakeslee (3)

Zoopagaceae Drechsler
Acaulopage Drechsler (27)
Cystopage Drechsler (9)
Lecophagus M.W. Dick (2)
Stylopage Drechsler (17)
Tentaculophagus Doweld (1)
Zoopage Drechsler (11)
Zoophagus Sommerst. (4)

Zoopagales genus incertae sedis


Massartia De Wild. (1)

Zoopagomycotina genus incertae sedis


Basidiolum Cienk. (1)

Outline of Fossil fungi


The legitimate fungal genera known so far are listed below (with number of species in each
genus in brackets). Here we list genera based on Saccardoan System (Table 4), fossil fungal
sporophores, mycelia and other fungal remains (Table 5) and modern fungal genera to which
fossil species have been assigned (Table 6).

Table 4 Fossil fungal spores (according to Saccardoan System).

Fungi Family Genera


Imperfecti
Amerosporae Asyregraamspora Locq. & Sal.-Cheb. (1)
Basidiosporites Elsik (4)*
Biporipsilonites Kalgutkar & Janson. (11)*

1341
Table 4 Continued.

Fungi Family Genera


Imperfecti
Biporisporites Ke & Shi (2)
Cervichlamydospora R. Kar, Mand. & R.K. Kar (1)
Diporisporites Hammen (ca. 34)*
Dremuspora Sal.-Cheb. & Locq. (1)
Exesisporites Elsik (4)*
Foliopollenites Sierotin (3)
Foveodiporites C.P. Varma & Rawat (11)*
Fusidiporosporonites Z.C. Song (1)
Geotrichites Stubblef., C.E. Mill., T.N. Taylor & G.T. Cole (1)
Graphiolites Fritel (1)
Haplographites Félix(2)
Hypoxylonites Elsik (ca. 60)*
Inapertisporites Hammen (ca. 67)*
Incertisporites Hammen (1)
Lacrimasporonites R.T. Clarke (9)
Magnosporites Rouse (1)
Microsporonites R.K. Jain (2)
Monoporisporites Hammen (ca. 58)
Nigrosporites Debi Mukh. (1)
Palaeoamphisphaerella Ramanujam & Srisailam (3)*
Palaeopericonia C.G. Ibañez & Zamuner (1)*
Portalites Hemer & Nygreen (1)
Psiamspora Locq. & Sal.-Cheb. in Sal.-Cheb. & Locq. (2)
Retidiporites C.P. Varma & Rawat (1)
Saccisporonites Kalgutkar & Janson. (1)
Spirotremesporites Dueñas (ca. 20)*
Sporotrichites Göpp. & Berendt (3)
Striadiporites C.P. Varma & Rawat (14)*
Trichosporites Félix (1)
Triporisporites Hammen (1)
Uncinulites Pampal. (3)*
Xylariasporites Debi Mukh. (1)
Xylohyphites Kalgutkar & Sigler (1)
Didymosporae Ampulliferinites Kalgutkar & Sigler (1)
Cladosporites Félix (3)*
Dicellaeporisporites Kalgutkar (3)
Dicellaesporites Elsik (ca. 58)
Didymoporisporonites Sheffy & Dilcher (ca. 25)
Didymosporonites Sal.-Cheb. & Locq. (1)
Dyadosporites Hammen ex R.T. Clarke (ca. 41)*
Felixites Elsik ex Janson. & Hills (2)
Fusiformisporites Rouse (21)*
Hilidicellites Kalgutkar & Janson. (21)*
Phragmosporae Alleppeysporonites Ramanujam & K.P. Rao (1)*
Anatolinites Elsik, V.S. Ediger & Bati (13)*
Axisporonites Kalgutkar & Janson. (1)*
Brachysporisporites R.T. Lange & P.H. Sm. (23)*
Ceratohirudispora R. Kar, Mand. & R.K. Kar (2)
Cercosporites E.S. Salmon (3)*
Chaetosphaerites Félix (4)*
Chordecystia C.B. Foster (1)
Circinoconites R. Kar, Mand. & R.K. Kar (1)
Cladosporiumsporinites Debi Mukh. (1)
Diporicellaesporites Elsik (ca. 61)
Diporipollis S.K. Dutta & S.C.D. Sah emend. Kalgutkar & Janson. (2)

1342
Table 4 Continued.

Fungi Family Genera


Imperfecti
Dwayabeejaesporonites Debi Mukh. (1)
Edmundmasonaesporites Debi Mukh. (1)
Foveoletisporonites Ramanujam & K.P. Rao (3)
Fractisporonites R.T. Clarke (9)
Heterocystinella Cookson & Eisenack (1)
Jansoniisporites Kalgutkar (1)
Kumarisporites Kalgutkar & Janson. (1)*
Mathurisporites Kalgutkar & Janson. (2)*
Monilites Pampal. (1)
Multicellaesporites Elsik emend. P. Kumar (ca. 14)
Multicellites Kalgutkar & Janson. (48)
Ornasporonites Ramanujam & K.P. Rao (1)*
Paragranatisporites Zhong Y. Zhang (5)
Phialophoronites Debi Mukh. (1)
Pluricellaesporites Hammen (ca. 72)*
Quilonia K.P. Jain & R.C. Gupta emend. Kalgutkar & Janson. (11)
Ramasricellites Kalgutkar & Janson. (2)*
Reduviasporonites L.R. Wilson (9)
Reticellites D.L.E. Glass, D.D. Br. & Elsik (1)
Scolecosporites R.T. Lange & P.H. Sm. (4)
Tripithonites Sat. K. Srivastava & Al-Tayyar (2)
Tympanicysta Malme (1)
Varmasporites Kalgutkar & Janson. (1)*
Dictyosporae Centonites Peppers (1)
Ctenosporites Elsik & Janson. (3)
Dictyosporites Félix emend. Kalgutkar & Janson. (ca. 20)
Dictyostromata R. Kar, Mand. & R.K. Kar (2)
Kutchiathyrites R.K. Kar emend. Kalgutkar & Janson. (7)*
Lirasporis R. Potonié & S.C.D. Sah (3)
Octosporites Sal.-Cheb. & Locq. (1)
Palambages Wetzel (3)
Papulosporonites Schmied. & G. Schwab (7)*
Polyadosporites Hammen (ca. 9)
Polycellaesporonites Anil Chandra, R.K. Saxena & Setty (7)*
Staphlosporonites Sheffy & Dilcher (c.21)*
Helicosporae Colligerites K.P. Jain & R.K. Kar (3)*
Elsikisporonites P. Kumar (1)
Helicominites Barlinge & Paradkar (1)
Helicoönites Kalgutkar & Sigler (1)
Helicosporiates Kalgutkar & Sigler (1)
Involutisporonites R.T. Clarke (ca. 8)
Palaeocirrenalia Ramanujam & Srisailam (3)
Paleoslimacomyces Kalgutkar & Sigler (3)*
Retihelicosporonites Ramanujam & K.P. Rao (1)*
Staurosporae Eoglobella W.H. Bradley (1)
Frasnacritetrus Taug. (7)*
Mossopisporites Kalgutkar & Janson. (1)*
Pesavis Elsik & Janson. (3)
Spegazzinites Félix (3)
Tribolites W.H. Bradley (2)*
Trihyphites Kalgutkar & Janson. (1)*
Triporicellaesporites Ke & Shi (4)*

1343
Table 5 Fossil fungal fructifications, mycelia and other fungal remains.

Phylum Order Genera


Ascomycota Botryosphaeriales Guignardiacarpites Debi Mukh. (1)
Capnodiales Mycosphaerellascoidetes Debi Mukh. (1)
Dothideales Cucurbitariaceites R.K. Kar, R.Y. Singh & S.C.D. Sah (2)*
Leptosphaerites Richon (2)
Palaeoleptosphaeria Barlinge & Paradkar (1)
Perisporiacites Félix (4)
Erysiphales Erisiphites Pampal. (1)
Meliolinites Selkirk (9)
Meliostroma R. Kar, Mand. & R.K. Kar (1)
Palaeosclerotium G.W. Rothwell (1)*
Perisporites Pampal. (2)
Eurotiales Coleocarpon Stubblef., T.N. Taylor, C.E. Miller & G.T. Cole (1)
Cryptocolax R.A. Scott (2)
Memnonillasporonites Debi Mukh. (1)
Mycocarpon S.A. Hutch. (7)*
Roannaisia T.N. Taylor, Galtier & Axsmith (1)
Sporocarpon Will. (13)*
Traquairia Carruth. ex Scott (4)
Hysteriales Hysterites Unger (16)
Microthyriales Appendicisporonites R.K. Saxena & S. Khare (1)
Asterinites Doub. & D. Pons ex Kalgutkar & Janson. (2)
Asterothyrites Cookson (16)
Brefeldiellites Dilcher (2)
Caldesites Puri (1)
Callimothallus Dilcher (11)
Cribrites R.T. Lange (1)
Dictyotopileos Dilcher (1)
Euthythyrites Cookson (4)
Haplopeltis Theiss. (5)
Kalviwadithyrites M.R. Rao (1)
Koshalia S. Sarkar & V. Prasad (1)
Mariusia D. Pons & Boureau (1)
Melanosporites Pampal. (1)
Microthyriacites Cookson (19)
Microthyrites Pampal. (1)
Molinaea Doub. & D. Pons (1)
Palmellathyrites Locq., D. Pons & Sal.-Cheb. (1)
Parmathyrites K.P. Jain & R.C. Gupta (5)
Pelicothallos Dilcher (1)
Phragmothyrites W.N. Edwards (24)*
Plochmopeltinites Cookson (3)
Polyhyphaethyrites R. Srivast. & R.K. Kar (1)
Ratnagiriathyrites R.K. Saxena & N.K. Misra (1)*
Spinosporonites R.K. Saxena & S. Khare (1)*
Stomiopeltites Alvin & M.D. Muir (3)
Trichopeltinites Cookson (5)
Trichothyrites Rosend. (13)*
Ussurithyrites Krassilov (1)
Patellariales Rhytidhysteriumites Debi Mukh. (1)
Pezizales Ascodesmisites Trivedi, Chaturv. & C.L. Verma (1)\
Paleomorchella Poinar (1)
Pezizites Göpp. & Berendt (4)
Phyllachorales Paleoserenomyces Currah, Stockey & B.A. LePage (1)
Pleosporales Cryptodidymosphaerites Currah, Stockey & B.A. LePage (1)*
Dictyosporiuminites Debi Mukh. (1)
Pleosporites Y. Suzuki (1)

1344
Table 5 Continued.

Phylum Order Genera


Sphaeriales Diploneurospora K.P. Jain & R.C. Gupta (1)*
Palaeosordaria Sahni & H.S. Rao (1)
Petrosphaeria Stopes & H. Fujii (1)
Valsarites Puri (1)
Uredinales Aeciosporonites Debi Mukh. (1)
Xylariales. Chaethomites Pampal. (1)
Sphaerites Unger (48)
Incertae sedis Cephalothecoidomyces G. Worobiec, Neumann & E. Worobiec (1)
Basidiomycota Agaricales. Archaeomarasmius Hibbett, D. Grimaldi & Donoghue (1)
Coprinites Poinar & Singer (1)*
Gondwanagaricites Heads, A.N. Mill & J.L. Crane (1)
Protomycena Hibbett, D .Grimaldi & Donoghue (1)
Polyporales. Eopolyporoides Rigby (1)
Phellinites Singer & S. Archang. (1)
Pseudopolyporus Hollick (1)
Trametites A. Straus (3)
Pucciniales. Shuklania J.N. Dwivedi (1)
Sphaeropsidales Archephoma Kyoto Watanabe, H. Nishida & Tak. Kobay. (1)
Ascochytites Barlinge & Paradkar (2)*
Deccanodia Singhai (1)
Diplodites D.N. Babajan & Tasl. ex Kalgutkar, Nambudiri & Tidwell
(5)*
Entopeltacites Selkirk (6)
Meniscoideisporites Kyoto Watanabe, H. Nishida & Tak. Kobay. (1)
Mohgaonidium Singhai (1)
Palaeocytosphaera R.B. Singh & G.V. Patil (1)
Palaeophoma Singhai (1)*
Phomites Fritel (2)
Rabenhorstinidium R.B. Singh & G.V. Patil (1)
Uredinales Aecidites Debey & Ettingsh. (4)
Aeciosporonites Debi Mukh. (1)
Hapalophragmites Ramanujam & Ramachar (1)
Milesites Ramanujam & Ramachar (1)
Pucciniasporonites Ramanujam & Ramachar (1)
Ustilaginales Chlamydosporites Paradkar (1)
Teliosporites R. Kar, Mand. & R.K. Kar (2)
Chytridiomycota Chytridiales Grilletia Renault & C.E. Bertrand (1)
Guizhounema X. Mu (1)
Krispiromyces T.N. Taylor, Hass & W. Remy (1)
Lyonomyces T.N. Taylor, Hass & W. (1)
Milleromyces T.N. Taylor, Hass & W. Remy (1)
Oochytrium Renault (1)
Mucoromycota Endogonales Chlamydospora R. Kar, Mand. & R.K. Kar (1)
Endochaetophora J.F. White & T.N. Taylor (1)
Gigasporites Carlie J. Phipps & T.N. Taylor (1)
Palaeogigaspora R. Kar, Mand. & R.K. Kar (1)
Palaeomycites Mesch. (21)*
Udaria A. Gupta (2)
Mucoromycota Lithomucorites R. Kar, Mand. & R.K. Kar (2)
genera incertae
sedis
Mycelia Sterilia Animikiea Bargh. (1)
Archaeorestis Bargh. (1)
Celyphus Batten (1)
Dendromyceliates K.P. Jain & R.K. Kar (2)
Entosphaeroides Bargh. (1)

1345
Table 5 Continued.

Phylum Order Genera


Eoastrion Bargh. (2)
Eomycetopsis J.W. Schopf (2)
Fungites Hallier (7)
Gunflintia Bargh. (2)
Laevitubulus N.D. Burgess & D. Edwards (5)
Ornatifilum N.D. Burgess & D. Edwards (2)
Palaeancistrus R.L. Dennis (1)
Palaeofibulus J.M. Osborn, T.N. Taylor & J.F. White (1)
Sclerotites A. Massal. (16)
Tormentella H.D. Pflug (2)
Fossil fungi Annella Sat. K. Srivast. (2)
incertae sedis Caenomyces E.W. Berry (Pyrenomycetes Schwein?) (1)
Dictyomykus R. Kar, Mand. & R.K. Kar (1)
Lithosporocarpia R. Kar, Mand. & R.K. Kar (1)
Mycokidstonia D. Pons & Locq. (1)
Mycozygosporangia R. Kar, Mand. & R.K. Kar (1)
Netothyrites C.M. Misra, S.N. Swamy, B. Prasad, B.S. Pundeer, R.S.
Rawat & K. Singh (2)
Palaeocercospora S. Mitra and Manju Banerjee (1)
Palaeocolletotrichum S. Mitra and Manju Banerjee (1)
Paleoblastocladia W. Remy, T.N. Taylor & Hass (1)
Palynomorphites L.R. Moore (1)
Pilula Harker, Sarjeant & Caldwell ex Harker & Sarjeant (2)
Protoascon L.R. Batra, Segal & R.W. Baxter (1)
Protocolletotrichum R. Kar, Mand. & R.K. Kar (1)
Reymanella Marcink. (1)
Sorosporonites X. Mu (1)
Stauromyca R. Kar, Mand. & R.K. Kar (1)
Tetradigita R. Kar, Mand. & R.K. Kar (1)
Tricellaesporonites Sheffy & Dilcher (3)

Table 6 Modern fungal genera to which fossil species have been assigned.

Phylum Order Family Modern genera Fossil species


Ascomycota Asterinales Asterinaceae Asterina Lév. A. eocenica Dilcher, A.
kosciuskensis Selkirk, A.
nodosaria Dilcher, A.
indodeightonii Vishnu, Khan
& Bera, A. mioconsobrina
Vishnu, Khan & Bera, A.
miosphaerelloides Vishnu,
Khan & Bera, A.
neocombreticola Vishnu,
Khan & Bera, A.
neoelaeocarpi Vishnu, Khan
& Bera, A. presaracae
Vishnu, Khan & Bera
Botryosphaeriales D. rodei Mahab. [Now:
Diplodites rodei (Mahab.)
Kalgutkar, Nambudiri &
Tidwell], D. sahnii Singhai
[Now: Diplodites sahnii
(Singhai) Kalgutkar,
Nambudiri & Tidwell]

1346
Table 6 Continued.

Phylum Order Family Modern genera Fossil species


Capnodiales Mycosphaerellaceae Ramularia Sacc. R. oblongispora Casp.
Chaetosphaeriales Chaetosphaeriaceae Chaetosphaeria C. elsikii M.J. Pound, J.M.K.
Tul. & C. Tul. O'Keefe, N.B. Nuñez Otaño,
J.B. Riding
Eurotiales Aspergillaceae Penicillium Link P. curtipes Berk.
Helotiales Mollisiaceae Trimmatostroma Trimmatostroma
Corda. intertrappea K.S. Patil &
Datar
Hypocreales Bionectriaceae Acremonium A. succineum Casp.
Link
Ceratostomatacea Gonatobotrys G. primigenius Casp.
Corda
Laboulbeniales Laboulbeniaceae Stigmatomyces Stigmatomyces succini W.
H. Karst. Rossi, Kotrba & Triebel
Lecanorales Sphaerophoraceae Sphaerophorus S. moniliformis Menge
Pers.
Meliolales Meliolaceae Meliola Fr. M. anfracta Dilcher [Now:
Meliolinites anfractus
(Dilcher) Kalgutkar &
Janson.], M. spinksii Dilcher
[Now: Meliolinites spinksii
(Dilcher) Selkirk]
Microthyriales Microthyriaceae Trichopeltina T. exporrecta Dilcher
Theiss.
Mycocaliciales Mycocaliciaceae Chaenothecopsis C. bitterfeldensis Rikkinen
Vain. & Poinar
Pleosporales Didymellaceae Epicoccum Link E. deccanense R. Srivast.,
Kapgate & S. Chatterjee
Pleosporaceae Alternaria Nees A. malayensis Trivedi &
ex Fr. C.L. Verma [Now:
Pluricellaesporites
malayensis (Trivedi & C.L.
Verma) Kalgutkar &
Janson.]
Torulaceae Torula Pers. ex T. globulifera Casp., T.
Fr. heteromorpha Casp., T.
mengeana Casp. & R. Klebs
in Casp.
Sporidesmiales Sporidesmiaceae Sporidesmium S. henryense Dilcher
Link ex Fr.
Taphrinales Protomycetaceae Protomyces P. protogenes W. Sm.
Unger
Trichosphaeriales Trichosphaeriaceae Brachysporium B. minutum Trivedi & C.L.
Sacc. Verma [Now:
Pluricellaesporites minutus
(Trivedi & C.L. Verma) ex
Kalgutkar & Janson.]
Dothideomycetes Vizellaceae Vizella Sacc. V. discontinua Selkirk, V.
family incertae memorabilis (Dilcher)
sedis Selkirk
Incertae sedis Incertae sedis Desmidiospora D. marginiconvoluta
Thaxt. Kalgutkar
Manginula G. M. maegdefraui Lange
Arnaud [Now: Entopeltacites
maegdefraui (Lange)
Selkirk], M. memorabilis

1347
Table 6 Continued.

Phylum Order Family Modern genera Fossil species


(Dilcher) Lange [Now: Vizella
memorabilis (Dilcher)
Selkirk], M. osbornii Lange
[Now: Entopeltacites osbornii
(Lange) Selkirk]
Sarcophoma S. deccani R.B. Singh & G.V.
Höhn. Patil
Tetracoccosporiu T. eocenum Biradar & Mahab.
m Szabó
Monotosporella S. M. doerfeltii Sadowski,
Hughes Beimforde, Gube & A.R.
Schmidt
Rhexoampullifera R. stogieana M.J. Pound,
(M.B. Ellis) P.M. J.M.K. O'Keefe, N.B. Nuñez
Kirk & C.M. Kirk Otaño, J.B. Riding, R. sufflata
M.J. Pound, J.M.K. O'Keefe,
N.B. Nuñez Otaño, J.B. Riding
Basidiomycota Agaricostilbales Chionosphaeraceae Stilbum Tode ex S. succini Casp.
Fr.
Boletales Sclerodermataceae Scleroderma S. echinosporites Rouse
Pers.
Cantharellales Hydnaceae Hydnum L. ex H. argillae R. Ludw.
Fr.
Geastrales Geastraceae Geastrum Pers. G. tepexense Magallon-Puebla
& Cevallos-Ferriz
Nidulariales Nidulariaceae Nidula V.S. White N. baltica Poinar
Cyathus Haller C. dominicanus Poinar
Polyporales Polyporaceae Fomes (Fr.) Fr. F. idahoensis R.W. Br.
Urocystidales Urocystidaceae Mundkurella M. mohgaoensis Chitaley &
Thirum. Yawale
Ustilaginales Ustilaginaceae Ustilago (Pers.) U. deccani Chitaley & Yawale
Roussel [Now: Inapertisporites
deccani (Chitaley & Yawale)
Kalgutkar & Janson.]
Chytridiomyco Chytridiales Chytriomycetaceae Entophlyctis A. E. willoughbyi W.H. Bradley
ta Fisch. [Now: Desmidiospora
willoughbyi (W.H. Bradley)
D.L.E. Glass, D.D. Br. &
Elsik]
Fungi incertae Patoullardiella P. imbricata Dilcher
sedis Speg.

Outline of fungus-like organisms


Obazoa Brown et al.
Opisthokonta Cavalier-Smith
Holomycota Liu et al. = Nucletmycea Brown et al.

Nucleariae Tedersoo et al.


Fonticulida Tedersoo et al.
Fonticulea Tedersoo et al.
Fonticulida Cavaler-Smith
Fonticulidae Worley, Raper & Hohl
Fonticula Worley, Raper & M. Hohl

1348
Diaphoretickes Adl et al.
S A R Burki et al. emend. Adl et al.
Rhizaria Cavalier-Smith
Endomyxa Cavalier-Smith
Phytomyxea Engler & Prantl
Plasmodiophorida Cook
Plasmodiophoridae Loeblich & Tappan
Ligniera Maire & A. Tison
Plasmodiophora Worona
Polymyxa Ledingham
Sorosphaerula Neuh. & Kirchm.
Spongospora Brunch.
Woronina Cornu

Phagomyxida Cavalier-Smith
Phagomyxidae Cavalier-Smith
Maullinia I. Maier, E.R. Parodi, Westermeier & D.G. Müll
Phagomyxa Karling

Cercozoa Cavalier-Smith
Sainouroidea Schuler et al.
Guttulinopsidae L.S. Olive
Guttulinopsis E.W. Olive

Straminipila M.W. Dick


Labyrinthulomycota Whittaker
Labyrinthulomycetes Dick
Labyrinthulales E.A. Bessey
Aplanochytriaceae Leander ex Cavalier-Smith
Aplanochytrium Bahnweg & Sparrow

Stellarchytriaceae Bennett et al. ad int.


Stellarchytrium FioRito & Leander

Labyrinthulaceae Haeckel
Labyrinthula Cienk.

Oblongichytridiales Bennett et al. ad int.


Oblongichytridiaceae Cavalier-Smith
Oblongichytrium R. Yokoy. & D. Honda

Thraustochytriales Sparrow
Althornidiaceae Jones and Alderman
Althornia E.B.G. Jones & Alderman

Thraustochytriacae Sparrow ex Cejp


Aurantiochytrium R. Yokoy. & D. Honda
Botryochytrium R. Yokoy., Salleh & D. Honda
Japanochytrium
Monorhizochytrium K. Doi & D. Honda
Parietichytrium R. Yokoy., Salleh & D. Honda
Schizochytrium S. Goldst. & Belsky ex Raghuk.

1349
Sicyoidochytrium R. Yokoy., Salleh & D. Honda
Thraustochytrium Sparrow
Ulkenia A. Gaertn. ex M.W. Dick

Amphitremida Gomaa et al.


Amphitremidae Poch
Amphitrema Archer
Archerella Loeblich & Tappan
Paramphitrema Valkanov

Diplophrydae Cavalier-Smith
Diplophrys J.S.F. Barker

Amphifilida Cavalier-Smith
Amphifilidae Cavalier-Smith
Amphifila Caval.-Sm.

Sorodiplophryidae Cavalier-Smith
Fibrophrys Takahashi et al.
Sorodiplophrys L.S. Olive & Dykstra

Hyphochytriomycota Whittaker
Hyphochytriomycetes Sparrow
Hyphochytriales Bessey ex Sparrow
Hyphochytriaceae Fischer
Canteriomyces Sparrow
Cystochytrium Ivimey Cook
Hyphochytrium Zopf

Rhizidiomycetaceae Karling ex Kirk, Cannon & David


Latrostium Zopf
Reessia Fisch
Rhizidiomyces Zopf

Oomycota Arx
Peronosporomycetes M.W. Dick
Albuginales Thines
Albuginaceae Schroet.
Albugo (Pers.) Roussel (40)
Pustula Thines (11)
Wilsoniana Thines (5)

Peronosporales A.N. Beketov


Peronosporaceae de Bary
Basidiophora Roze & Cornu (3)
Baobabopsis R.G. Shivas, Y.P. Tan, Telle & Thines (2)
Benua Constant. (1)
Bremia Regel (15)
Calycofera R. Bennett & Thines (2)
Eraphthora Telle & Thines (1)
Graminivora Thines (1)
Halophytophthora H.H. Ho & S.C. Jong (6)

1350
Hyaloperonospora Constant. (35)
Nothophytophthora T. Jung, Scanu, Bakonyi & M. Horta Jung (6)
Novotelnova Voglmayr & Constant. (1)
Paraperonospora Constant. (9)
Perofascia Constant. (2)
Peronospora Corda (350)
Peronosclerospora (S. Ito) Hara (15)
Phytophthora de Bary (150)
Phytopythium Abad, de Cock, Bala, Robideau, A.M. Lodhi & Lévesque (25)
Plasmopara J. Schröt. (150)
Plasmoverna Constant., Voglmayr, Fatehi & Thines (7)
Poakatesthia Thines (1)
Protobremia Voglmayr, Riethm., Göker, Weiss & Oberw. (1)
Pseudoperonospora Rostov. (9)
Sclerophthora Thirum., C.G. Shaw & Naras. (5)
Sclerospora J. Schröt. (2)
Viennotia Göker, Voglmayr, Riethm., M. Weiss & Oberw. (1)

Pythiaceae Schroet.
Elongisporangium Uzuhashi, Tojo & Kakish. (5)
Globisporangium Uzuhashi, Tojo & Kakish. (70)
Lagena Vanterp. & Ledingham (1)
Lagenidium Schenk (40)
Myzocytiopsis M.W. Dick (18)
Myzocytium Schenk (2)
Pilasporangium (Uzuhashi & Tojo) Uzuhashi, Tojo & Kakish. (1)
Pythiogeton Minden (16)
Pythium Pringsh. (200)

Salisapiliaceae
Salisapilia Hulvey, Nigrelli, Telle, Lamour & Thines (9)

Rhipidiales M.W. Dick


Rhipidiaceae Cejp
Aqualinderella Emerson & Weston (1)
Araiospora Thaxt. (4)
Mindeniella Kanouse (2)
Nellymyces A. Batko (1)
Rhipidium Cornu (6)
Sapromyces Fritsch (4)

Salispinaceae R. Bennett & Thines


Salispina Marano, A.L. Jesus & Pires-Zottar. (4)

Peronosporomycetes genera incertae sedis


Kawakamia Miyabe (4)
Paralagenidium Grooters, C.F.J. Spies, de Cock & Lévesque (2)
Trachysphaera Tabor & Bunting (1)

Saprolegniomycetes Thines & Beakes


Leptomitales Kanouse
Atkinsiellaceae Sparrow

1351
Atkinsiella Vishniac (1)
Bolbea Buaya & Thines (1)

Leptomitaceae Kütz
Apodachlya Pringsh. (5)
Apodachlyella Indoh (1)
Blastulidium Pérez (1)
Leptomitus C. Agardh (11)

Ectrogellaceae Cejp
Crypticola Humber, Frances & A.W. Sweeney (1)
Ectrogella Zopf (8)
Lagenisma Schnepf (1)

Saprolegniales K. Prantl
Achlyaceae ined.
Achlya Nees (80)
Brevilegnia Coker & Couch (16)
Dictyuchus Leitg. (9)
Thraustotheca Humphrey (4)

Saprolegniaceae Warm.
Aplanopsis Höhnk (1)
Calyptralegnia Coker (3)
Couchia W.W. Martin (3)
Isoachlya Kauffmann (9)
Newbya M.W. Dick & M.A. Spencer (13)
Protoachlya Coker (7)
Pythiopsis de Bary (7)
Saprolegnia Nees (80)
Scoliolegnia M.W. Dick (5)

Verrucalvaceae M.W. Dick


Aphanomyces de Bary (40)
Aquastella Glockling & D.P. Molloy (2)
Geolegnia Coker (4)
Leptolegnia de Bary (9)
Pachymetra B.J. Croft & M.W. Dick (1)
Plectospira Drechsler (4)
Verrucalvus P. Wong & M.W. Dick (1)

Saprolegniomycetes genera incertae sedis


Aphanomycopsis Scherff. (6)
Brevilegniella M.W. Dick (1)
Cornumyces M.W. Dick (8)
Clamydomycium M.W. Dick (7)
Ducellieria Teiling (1)
Eurychasmopsis Canter & M.W. Dick (1)
Leptolegniella Huneycutt (7)
Nematophthora Kerry & D.H. Crump (1)
Pythiella Couch (3)
Sommerstorffia Arnaudov (1)

1352
Synchaetophagus Apstein (1)

Oomycota orders incertae sedis


Anisolpidiales M.W. Dick
Anisolpidiaceae Karling
Anisolpidium Karling (7)

Diatomophthoraceae Buaya & Thines


Diatomophthora Buaya & Thines (3)

Eurychasmales Sparrow
Eurychasmataceae Petersen
Eurychasma Magnus (3)

Haliphthorales ined.
Haliphthoraceae Vishniac
Halioticida Muraosa & Hatai (1)
Halocrusticida K. Nakam. & Hatai (7)
Haliphthoros Vishniac (3)

Haptoglossales M.W. Dick


Haptoglossaceae M.W. Dick
Haptoglossa Drechsler (12)

Miraculales ined.
Miraculaceae Buaya, Hanic & Thines
Miracula Buaya, Hanic & Thines (2)

Olpidiopsidales M.W. Dick


Olpidiopsidaceae Sparrow
Olpidiopsis Cornu (12)

Pontismatales Thines
Postismataceae H.E. Petersen
Petersenia Sparrow (3)
Pontisma H.E. Petersen (10)
Sirolpidium H.E. Petersen (7)

Rozellopsidales M.W. Dick


Rozellopsidaceae M.W. Dick
Rozellopsis Karling (5)

Amorphea Adl et al.


Amoebozoa Lühe
Evosea Kang et al.
Eumycetozoa L.S. Olive
Dictyosteliomycetes Doweld
Acytosteliales S. Baldauf, S. Sheikh & Thulin
Acytosteliaceae Raper ex Raper & Quinlan
Acytostelium Raper
Heterostelium S. Baldauf, S. Sheikh & Thulin
Rostrostelium S. Baldauf, S. Sheikh & Thulin

1353
Cavenderiaceae S. Baldauf, S. Sheikh & Thulin
Cavenderia S. Baldauf, S. Sheikh & Thulin

Dictyosteliales L.S. Olive ex P.M. Kirk et al.


Dictyosteliaceae Rostaf. ex Cooke
Dictyostelium Bref.
Polysphondylium Bref.

Raperosteliaceae S. Baldauf, S. Sheikh & Thulin


Hagiwaraea S. Baldauf, S. Sheikh & Thulin
Raperostelium S. Baldauf, S. Sheikh & Thulin
Speleostelium S. Baldauf, S. Sheikh & Thulin
Tieghemostelium S. Baldauf, S. Sheikh & Thulin

Dictyosteliales genus incertae sedis


Coremiostelium S. Baldauf, S. Sheikh, Thulin & Spiegel

Dictyosteliomycetes genera incertace sedis


Coenonia Tiegh.
Synstelium S. Baldauf, S. Sheikh & Thulin

Ceratiomyxomycetes D. Hawksw., B. Sutton & Ainsw. in Leontyev et al. (2019)


Ceratiomyxales G.W. Martin ex M.L. Farr & Alexop.
Ceratiomyxaceae J. Schröt.
Ceratiomyxa J. Schröt.
Protosporangiaceae Leontyev, Stephenson, Schnittler, Shchepin, Novozhilov
Clastostelium L.S. Olive & Stoian.
Protosporangium L.S. Olive & Stoian.

Myxomycetes G. Winter
Lucisporomycetidae Leontyev, Schnittler, S.L. Stephenson, Novozhilov & Shchepin
Cribrariales T. Macbr.
Cribrariaceae Corda
Cribraria Pers.
Licaethalium Rostaf.
Lindbladia Fr.

Reticulariales Leontyev, Schnittler, S.L. Stephenson, Novozhilov & Shchepin


Reticulariaceae Chevall. ex Corda
Alwisia Berk. & Broome (6)
Lycogala Adans.
Reticularia Bull.
Tubifera J.F. Gmel.
Siphoptychium Rostaf.
Thecotubifera Leontyev, Schnittler, S.L. Stephenson & Novozh.

Liceales E. Jahn
Liceaceae Chevall.
Licea Schrad.
Listerella E. Jahn

1354
Trichiales T. Macbr.
Dianemataceae T. Macbr.
Calomyxa Nieuwl.
Dianema Rex
Dictydiaethalium Rostaf.
Prototrichia Rostaf.

Trichiaceae Chevall.
Arcyodes O.F. Cook
Arcyria F.H. Wigg.
Cornuvia Rostaf.
Hemitrichia Rostaf.
Metatrichia Ing
Oligonema Rostaf.
Perichaena Fr.
Trichia Haller

Lucisporomycetidae genera incertae sedis


Arcyriatella Hochg. & Gottsb.
Calonema Morgan
Minakatella G. Lister
Trichioides Novozh., Hoof & Jagers

Columellomycetidae Leontyev, Schnittler, S.L. Stephenson, Novozhilov & Shchepin


Echinosteliopsidales Shchepin, Leontyev, Schnittler, S.L. Stephenson, Novozhilov
Echinosteliopsidaceae L.S. Olive
Echinosteliopsis Reinhardt & L.S. Olive

Echinosteliales G.W. Martin


Echinosteliaceae Rostaf. ex Cooke
Barbeyella Meyl.
Echinostelium de Bary
Semimorula E.F. Haskins, McGuinn. & C.S. Berry

Clastodermatales Leontyev, Schnittler, S.L. Stephenson, Novozhilov & Shchepin


Clastodermataceae Alexop. & T.E. Brooks
Clastoderma A. Blytt.

Meridermatales Leontyev, Schnittler, S.L. Stephenson, Novozhilov & Shchepin


Meridermataceae Leontyev, Schnittler, S.L. Stephenson, Novozhilov & Shchepin
Meriderma Mar. Mey. & Poulain

Stemonitidales T. Macbr.
Amaurochaetaceae Rostaf. ex Cooke
Amaurochaete Rostaf.
Brefeldia Rostaf.
Comatricha Preuss
Enerthenema Bowman
Paradiacheopsis Hertel.
Stemonaria Nann.-Bremek., R. Sharma & Y. Yamam.
Stemonitopsis (Nann.-Bremek.) Nann.-Bremek.

1355
Stemonitidaceae Fr.
Macbrideola H.C. Gilbert
Stemonitis Gled.
Symphytocarpus Ing & Nann.-Bremek.

Physarales T. Macbr.
Lamprodermataceae T. Macbr.
Collaria Nann.-Bremek.
Colloderma G. Lister
Diacheopsis Meyl.
Elaeomyxa Hagelst.
Lamproderma Rostaf.

Didymiaceae Rostaf. ex Cooke


Diderma Pers.
Didymium Schrad.
Lepidoderma de Bary
Mucilago Battarra

Physaraceae Chevall.
Badhamia Berk.
Craterium Trentep.
Fuligo Haller
Kelleromyxa Eliasson
Leocarpus Link
Physarella Peck.
Physarina Höhn.
Physarum Pers.
Willkommlangea Kuntze

Columellomycetidae genera incertae sedis


Diachea Fr.
Leptoderma G. Lister
Paradiachea Hertel
Protophysarum M. Blackw. & Alexop.
Trabrooksia H.W. Keller

Variosea Cavalier-Smith et al.


Protosteliida Olive & Stoian. sensu Shadwick et Spiegel in Adl et al. 2012
Protosteliidae Olive & Stoian., emend Spiegel
Protostelium L.S. Olive & Stoian.

Fractovitellida Lahr et al. sensu Kang et al. 2017


Schizoplasmodiidae Shadwick & Spiegel in Adl et al.
Ceratiomyxella L.S. Olive & Stoian.
Nematostelium L.S. Olive & Stoian.
Schizoplasmodium L.S. Olive & Stoian.

Soliformoviidae Lahr & Katz


Soliformovum Spiegel

Cavosteliida Shadwick & Spiegel in Adl et al.

1356
Cavosteliidae S.L. Olive
Cavostelium S.L. Olive
Schizoplasmodiopsis S.L. Olive
Tychosporium Spiegel

Tubulinea Smirnov et al.


Elardia Kang et al.
Euamoebida Lepşi
Copromyxidae L.S. Olive & Stoian.
Copromyxa Zopf

Discosea Cavalier-Smith et al. sensu Smirnov et al. 2011


Flabellinea Smirnov et al.
Thecamoebida Schaeffer
Sappinia P.A. Dang.

Vannellida Smirnov et al.


Protosteliopsis L.S. Olive & Stoian.

Centramoebia Cavalier-Smith et al.


Acanthopodida Page
Acanthamoeba Volkonsky
Luapelamoeba Shadwick et al.

Pellitida Smirnov & Cavalier-Smith sensu Kang et al. 2017


Endostelium L.S. Olive, W.E. Benn. & Deasey
Discoba Simpson in Hampl et al.
Heterolobosea Page & Blanton
Tetramitia Cavalier-Smith
Eutetramitia Hanousková et al.
Acrasidae Poche
Acrasis Tiegh. (incl. Pocheina A.R. Loebl. & Tappan)

Discussion

Alternative classification of Leotiomycetes (Authors: A.H. Ekanayaka & K.D. Hyde)


The arrangement of Leotiomycetes in Ekanayaka et al. (2019) and Johnston et al et al.
(2019) are based on morphological interpretations and phylogenies using different data, however,
they are generally congruent. Johnston et al. (2019) used three phylogenetic reconstructions, one
based on 3156 single-copy genes for 49 taxa, the second based on 15 genes for 279 taxa, and the
third based on ITS alone for 568 taxa. Ekanayaka et al. (2019) used five genes with 482 taxa.
There are, however, some differences and therefore the outline from Ekanayaka et al. (2019) with
modifications is given below with notes. Understandably, if different taxa were used in the
Ekanayaka et al. (2019) and Johnston et al. (2019) phylogenies, different conclusions have been
drawn and therefore the classification of Leotiomycetes is not settled. We hope that by providing
an alternative outline, this will initiate positive discussion and further research with fresh
collections to resolve inconsistencies. However, with insufficient taxa in this class having been
sequenced it will take several years before the classification is stabilized.
Taxa with notes in this section are indicated by #.

Chaetomellales Crous & Denman


Chaetomellaceae Baral, P.R. Johnst. & Rossman

1357
Chaetomella Fuckel (26)
Corniculariella P. Karst. (1)
Pilidium Kunze (23)
Sphaerographium Sacc. (23)
Synchaetomella Decock & Seifert (3)
Xeropilidium Baral & Pärtel (1)

Cyttariales Luttr. ex Gamundí


Cordieritidaceae Sacc.#
Ameghiniella Speg. (2)
Annabella Fryar, Haelew., & D.E.A. Catches. (1)
Austrocenangium Gamundí (2)
Cordierites Mont. (5)
Diplocarpa Massee (2)
Diplolaeviopsis Giralt & D. Hawksw. (3)
Gelatinopsis Rambold & Triebel (8)
Ionomidotis E.J. Durand ex Thaxt. (13)
Llimoniella Hafellner & Nav.-Ros. (19)
Macroskyttea Etayo, Flakus, Suija & Kukwa (1)
Midotiopsis Henn. (2)
Phaeangella (Sacc.) Massee (11)
Rhymbocarpus Zopf (12)
Rhizocladosporium Crous & U. Braun (1)
Skyttea Sherwood, D. Hawksw. & Coppins (30)
Skyttella D. Hawksw. & R. Sant. (2)
Sabahriopsis Crous & M.J. Wingf. (1)
Thamnogalla D. Hawksw. (1)
Unguiculariopsis Rehm (25)

Cyttariaceae Lév.
Cyttaria Berk. (13)

Deltopyxidaceae Ekanayaka & K.D. Hyde#


Deltopyxis Baral & G. Marson (1)
Phaeopyxis Rambold & Triebel (1)

Erysiphales Gwynne-Vaughan#
Amorphothecaceae Parbery (= Myxotrichaceae Locq. ex Currah fide Ekanayaka et al. 2019)#
Amorphotheca Parbery (1)
Brefeldochium Verkley (1)
Byssoascus Arx (2)
Myxotrichum Kunze (12)
Oidiodendron Robak (ca. 30)
Polydesmia Boud. (7)
Hormoconis Arx & G.A. de Vries (2)

Erysiphaceae Tul. & C. Tul.


Arthrocladiella Vassilkov (1)
Blumeria Golovin ex Speer (1)
Brasiliomyces Viégas (6)
Bulbomicroidium Marm., S. Takam. & U. Braun (1)
Caespitotheca S. Takam. & U. Braun (1)

1358
Cystotheca Berk. &M.A. Curtis (9)
Erysiphe R. Hedw. ex DC. (478)
Golovinomyces (U. Braun) V.P. Heluta (66)
Leveillula G. Arnaud (49)
Microidium (To-anun & S. Takam.) To-anun & S. Takam. (3)
Neoerysiphe U. Braun (15)
Oidiopsis Scalia (ca. 12)
Oidium Link (ca. 200+)
Ovulariopsis Pat. & Har. (ca. 13)
Parauncinula S. Takam. & U. Braun (4)
Phyllactinia Lév. (117)
Pleochaeta Sacc. & Speg. (5)
Podosphaera Kunze (124)
Pseudoidium Y.S. Paul & J.N. Kapoor (ca. 80)
Queirozia Viégas & Cardoso (1)
Sawadaea Miyabe (10)
Takamatsuella U. Braun & A. Shi (1)
Typhulochaeta S. Ito & Hara (ca. 4)

Helotiales Nannf. ex Korf & Lizoň


Arachnopezizaceae Hosoya, J.G. Han & Baral
Arachnopeziza Fuckel (15)
Arachnoscypha Boud. (3)
Austropezia Spooner (1)
Eriopezia (Sacc.) Rehm (30)
Durella Tul. & C. Tul. (5)
Bryoglossaceae Ekanayaka & K.D. Hyde
Bryoclaviculus L. Ludw., P.R. Johnst. & Steel (1)
Bryoglossum Redhead (2)
Neocudoniella S. Imai (2)

Chlorociboriaceae Baral & P.R. Johnst.#


Chlorociboria Seaver ex C.S. Ramamurthi, Korf & L.R. Batra (22)

Chlorospleniaceae Ekanayaka & K.D. Hyde


Chlorosplenium Fr. (5)

Chrysodiscaceae Baral & Haelew.


Chrysodisca Baral, Polhorský & G. Marson (1)

Discinellaceae Ekanayaka & K.D. Hyde


Articulospora Ingold (ca. 5)
Acidea Hujslová & M. Kolařík (1)
Cladochasiella Marvanová (1)
Discinella Boud. (ca. 15)
Fontanospora Dyko (1)
Gyoerffyella Kol (10)
Lemonniera De Wild. (7)
Naevala B. Hein (6)
Margaritispora Ingold (1)
Pezoloma Clem. (ca. 15)
Tetrachaetum Ingold (3)

1359
Drepanopezizaceae Baral (=Drepanopezizaceae Bat. & H. Maia; Nom. inval., Arts 32.1(c), 36,
39.1 (Melbourne))
Blumeriella Arx (= Higginsia Nannf.; = Phloeosporella Höhn., = Microgloeum Petr.) (7)
Diplocarpon F.A. Wolf (7)
Drepanopeziza (Kleb.) Höhn. (= Gloeosporidiella Petr.) (5)
Felisbertia Viégas (7)
Leptotrochila P. Karst. (= Ephelina Sacc.; = Fabraea Sacc.; = Sporonema Desm.) (15)
Pseudopezicula Korf (2)
Spilopodia Boud. (= Holmiodiscus Svrček; = Melanodiscus Höhn.) (ca. 4)
Spilopodiella E. Müll. (1)
Thedgonia B. Sutton (6)

Gelatinodiscaceae S.E. Carp


Ascocoryne J.W. Groves & D.E. Wilson (= Didymocoryne Sacc. & Trotter) (8)
Ascotremella Seaver (2)
Chloroscypha Seaver (14)
Dimorphospora Tubaki (1)
Helicodendron Peyronel (3)
Neobulgaria Petr. (11)
Phaeangellina Dennis (1)
Skyathea Spooner & Dennis (1)

Godroniaceae Baral
Ascocalyx Naumov (4)
Atropellis Zeller & Goodd. (4)
Godronia Moug. & Lév. (ca. 30)
Gremmeniella M. Morelet (3)
Grovesiella M. Morelet (2)

Helotiaceae Rehm (= Roesleriaceae Y.J. Yao & Spooner fide Ekanayaka et al. 2019)
Amylocarpus Curr. (1)
Asterocalyx Höhn. (1)
Ascoconidium Seaver (3)
Bryoscyphus Spooner (19)
Calycella sensu (Sacc.) Sacc. (1)
Crocicreas Fr. (5)
Eubelonis Clem. (2)
Cudoniella Sacc. (31)
Cyathicula De Not. (30)
Dicephalospora Spooner (4)
Endoscypha Syd. (1)
Discorehmia Kirschst. (5)
Glarea Bills & Paláez (2)
Gloeotinia M. Wilson, Noble & E.G. Gray (2)
Helicocentralis Sri-indr., Chuaseehar., Boonyuen, K. Yamag., Suetrong & C.K.M. Tsui (1)
Hymenoscyphus Gray (170)
Hymenotorrendiella P.R. Johnst., Baral & R. Galán (9)
Lanzia Sacc. (55)
Muscicola Velen. (1)
Mytilodiscus Kropp & S.E. Carp. (1)
Ombrophila Fr. (11)

1360
Phaeohelotium Kanouse (41)
Pirottaea Sacc. (28)
Pithyella Boud. (8)
Pseudoniptera Velen. (25)
Roesleria Thüm. & Pass. (ca. 10)
Roeslerina Redhead (3)
Symphyosirinia E.A. Ellis (6)
Tatraea Svrcek (2)
Torrendiella Boud. & Torrend (3)
Xylogramma Wallr. (18)

Heterosphaeriaceae Rehm
Heterosphaeria Grev. (7)

Hyaloscyphaceae Nannf.
Ambrodiscus S.E. Carp. (1)
Aeruginoscyphus Dougoud (7)
Arbusculina Marvanová & Descals (3)
Clathrosphaerina Beverw. (2)
Curviclavula G. Delgado, F.A. Fernández & A.N. Mill. (1)
Dimorphotricha Spooner (1)
Echinula Graddon (1)
Glutinomyces Nor. Nakam. (1)
Graddonidiscus Raitv. & R. Galán (3)
Grahamiella Spooner (3)
Hegermila Raitv. (4)
Hyaloscypha Boud. (45)
Hyalodendriella Crous (1)
Hyphopeziza J.G. Han, Hosoya & H.D. Shin (1)
Incrupila Raitv. (10)
Meliniomyces Hambl. & Sigler (3)
Olla Velen. (2)
Parachnopeziza Korf (8)
Polaroscyphus Huhtinen (1)
Proprioscypha Spooner (1)
Protounguicularia Raitv. & R. Galán (10)
Pseudaegerita J.L. Crane & Schokn. (7)
Psilocistella Svrcek (10)
Rhizoscyphus W.Y. Zhuang & Korf (1)
Scytalidium Pesante (ca. 30)
Thindiomyces Arendh. & R. Sharma (1)
Unguiculariella K.S. Thind & R. Sharma (1)
Unguiculella Höhn. (17)

Hydrocinaceae Ekanayaka & K.D. Hyde


Clathrosporium Nawawi & Kuthub. (1)
Filosporella Nawawi (6)
Hydrocina Scheuer (1)
Varicosporium W. Kegel (11)
Xerombrophila Baral (1)

Lachnaceae Raitv.

1361
Albotricha Raitv. (19)
Asperopilum Spooner (1)
Belonidium Mont. & Dur. (1)
Brunnipila Baral (10)
Capitotricha (Raitv.) Baral (10)
Crucellisporiopsis Nag Raj (3)
Crucellisporium M.L. Farr (3)
Dasyscyphella Tranzschel (1)
Erioscyphella Kirschst. (10)
Incrucipulum Baral (6)
Lachnellula P. Karst. (40)
Lachnum Retz. (50)
Lachnopsis Guatim., R.W. Barreto & Crous (1)
Neodasyscypha Suková & Spooner (2)
Perrotia Boud. (19)
Proliferodiscus J.H. Haines & Dumont (8)
Tubolachnum Velen. (2)
Velebitea I. Kušan, Matočec & Jadan (1)

Loramycetaceae Dennis ex Digby & Goos


Acidomelania E. Walsh & N. Zhang (1)
Loramyces W. Weston (2)
Obtectodiscus E. Müll., Petrini & Samuels (2)

Mitrulaceae Rchb.
Mitrula Fr. (8)
Mollisiaceae Rehm
Bulbomollisia Graddon (1)
Cystodendron Bubák (2)
Discocurtisia Nannf. (12)
Mollisia (Fr.) P. Karst. (130)
Neotapesia E. Müll. & Hütter (3)
Niptera Fr. (10)
Nipterella Starbäck ex Dennis (2)
Phialocephala W.B. Kendr. (37)
Pseudonaevia Dennis & Spooner (2)
Pyrenopeziza Fuckel (3)
Sarconiptera Raitv. (1)
Scutobelonium Graddon (1)
Scutomollisia Nannf. (14)
Tapesia (Pers.) Fuckel (ca. 25)
Trimmatostroma Corda (30)
Variocladium Descals & Marvanová (1)

Ploettnerulaceae Kirschst.
Cadophora Lagerb. & Melin (15)
Collembolispora Marvanová & Pascoal (2)
Cylindrosporium Grev. (3)
Dennisiodiscus Svrcek (10)
Lasiomollisia Raitv. & Vesterh. (1)
Mastigosporium Riess (4)
Mycochaetophora Hara & Ogawa (2)

1362
Nothophacidium J. Reid & Cain (1)
Oculimacula Crous & W. Gams (6)
Peltigeromyces A. Möller (3)
Ploettnerula Kirschst. (1)
Pseudopeziza Fuckel (4)
Rhexocercosporidium U. Braun (2)
Rhynchosporium Heinsen ex A.B. Frank (5)
Ypsilina J. Webster, Descals & Marvanová (1)

Solenopeziaceae Ekanayaka & K.D. Hyde


Geniculospora Sv. Nilsson ex Marvanová & Sv. Nilsson (2)
Graddonia Dennis (7)
Halenospora E.B.G. Jones (1)
Lasiobelonium Ellis & Everh. (20)
Mycofalcella Marvanová, Om-Kalth. & J. Webster (2)
Solenopezia Sacc. (7)
Trichopeziza Fuckel (30)
Trichopezizella Dennis ex Raitv. (12)
Tricladium Ingold (25)

Vibrisseaceae Korf
Acephala Grünig & T.N. Sieber (2)
Cheirospora Moug. & Fr. (2)
Diplococcium Grove (30)
Fuscosclera Hern.-Restr., J. Mena & Gené (1)
Gorgoniceps (P. Karst.) P. Karst. (3)
Leucovibrissea (A. Sánchez) Korf (1)
Pocillum De Not. (1)
Strossmayeria Schulzer (= Pseudospiropes M.B. Ellis) (16)
Srinivasanomyces S. Rana & S.K. Singh (1)
Vibrissea Fr. (34)

Helotiales genera incertae sedis


Aquapoterium Raja & Shearer (1)
Arboricolonus S. Bien & Damm (1)
Barrenia E. Walsh & N. Zhang (2)
Brackelia Zhurb. (1)
Bulgariella P. Karst. (4)
Cecidioskyttea Etayo (1)
Chlorovibrissea L.M. Kohn (4)
Colipila Baral & Guy García (2)
Connersia Malloch (1)
Cryptocline Petr. (20)
Encoeliopsis Nannf. (4)
Gamarada D.J. Midgley & Tran-Dinh (1)
Larissia Raitv. (1)
Lemalis Fr. (3)
Libartania Nag Raj (2)
Merodontis Clem. (1)
Mitrulinia Spooner (1)
Mollisiopsis Rehm (7)
Muscia Gizhitsk (1)

1363
Patellariopsis Dennis (5)
Pestalopezia Seaver (3)
Phacidiella P. Karst. (1)
Pleuroascus Massee & E.S. Salmon (11)
Pseudomitrula Gamundí (1)
Sambucina Velen. (1)
Sarcomyces Massee (1)
Unguicularia Höhn. (7)

Lahmiales O.E. Erikss.


Lahmiaceae O.E. Erikss.
Lahmia Körb. (1)

Lauriomycetales Hern.-Restr., R.F. Castañeda & Guarro


Lauriomycetaceae Hern.-Restr., R.F. Castañeda & Guarro
Lauriomyces R.F. Castañeda (11)

Leotiales Korf & Lizoň


Cochlearomycetaceae Crous#
Cochlearomyces Crous (1)

Leotiaceae Corda
Geocoryne Korf (2)
Leotia Pers. (4)
Microglossum Gillet (ca. 10)
Thuemenidium Kuntze (1)
Mniaeciaceae Baral
Mniaecia Boud. (= Epiglia Boud.) (6)

Tympanidaceae Baral & Quijada


Aotearoamyces P.R. Johnst., J.A. Cooper & Quijada (1)
Claussenomyces Kirschst. (ca. 19)
Collophorina Damm & Crous (6)
Durandiella Seaver (15)
Gelatinosporium Peck (15)
Myriodiscus Boedijn (1)
Pragmopora A. Massal. (7)
Tympanis Fr. (ca. 27)

Leotiales genus incertae sedis


Gelatinomyces Sanoam., Sanoam., Jitjak, Rodtong & Whalley (2)

Lichinodiales M. Prieto, M. Schultz, Olariaga & Wedin


Lichinodiaceae M. Prieto, M. Schultz, Olariaga &Wedin
Lichinodium Nyl. (4)
Mycosymbioces J.L. Frank (1)

Lichinodiales genera incertae sedis


Epithamnolia Zhurb. (6)
Fulvoflamma Crous (1)

Marthamycetales P.R. Johnst. & Baral

1364
Marthamycetaceae Baral, Lantz, Hustad & Minter
Cyclaneusma DiCosmo, Peredo & Minter (2)
Marthamyces Minter (13)
Mellitiosporiella Höhn. (2)
Mellitiosporium Corda (11)
Naemacyclus Fuckel (6)
Phragmiticola Sherwood (1)
Propolina Sacc. (1)
Propolis (Fr.) Corda (ca. 9)

Medeolariales Korf#
Ascocorticiaceae J. Schröt.
Ascocorticiellum Jülich & B. de Vries (3)
Ascocorticium Bref. (8)
Ascosorus P. Henn. & Ruhland (1)

Ascodichaenaceae D. Hawksw. & Sherwood


Ascodichaena Butin (2)
Delpinoina Kuntze (2)

Dermateaceae Fr.
Arctomollisia Raitv. (2)
Cashiella Petr. (3)
Davidhawksworthia Crous (1)
Dermea Fr. (25)
Gelatinoamylaria Prasher & R. Sharma (1)
Neofabraea H.S. Jacks. (7)
Pezicula Tul. & C. Tul. (50)
Phlyctema Desm. (30)
Pseudofabraea Chen Chen, Verkley & Crous (1)
Rhizodermea Verkley & J.D. Zijlstra (1)
Schizothyrioma Höhn. (1)
Verkleyomyces Y. Marín & Crous (1)

Medeolariaceae Korf
Medeolaria Thaxt. (1)

Medeolariales genera incertae sedis


Coleophoma Höhn. (35)
Parafabraea Chen Chen, Verkley & Crous (2)

Phacidiales C.E. Bessey

Helicogoniaceae Baral
Eleutheromycella Höhn. (1)
Eleutheromyces Fuckel (1)
Gelatinipulvinella Hosoya & Y. Otani (2)
Geltingia Alstrup & D. Hawksw. (1)
Helicogonium W.L. White (19)
Humicolopsis Cabral & S. Marchand (2)

Phacidiaceae Fr. (= Bulgariaceae Fr.; = Phacidiostromataceae Höhn. fide Jaklitsch et al. 2016a)

1365
Allantophomopsiella Crous (1)
Allantophomopsis Petr. (8)
Bulgaria Fr. (2)
Calvophomopsis J.B. Tanney & Seifer (1)
Ceuthospora Grev. (ca. 100)#
Cornibusella J.B. Tanney & Seifer (1)
Darkera H.S. Whitney, J. Reid & Piroz. (5)
Gloeopycnis J.B. Tanney & Seifer (1)
Phacidiopycnis Potebnia (5)
Phacidium Fr. (ca. 200)
Potebniamyces Smerlis (1)#
Pseudophacidium P. Karst. (5)
Starbaeckia Rehm ex Starbäck (1)
Strasseria Bres. & Sacc. (1)#
#
Ekanayaka et al. (2019) accepted these genera in Phacidiaceae

Phacidiales genus incertae sedis


Coma Nag Raj & W.B. Kendr. (2)

Rhytismatales M.E. Barr ex Minter

Rhytismataceae Chevall. (= Hypodermataceae Rehm; = Cryptomycetaceae Höhn. nom. inval.


fide Jaklitsch et al. 2016a; = Cudoniaceae P.F. Cannon fide Ekanayaka et al. 2019)
Angelina Fr. (1)
Apiodiscus Petr. (1)
Bifusella Höhn. (11)
Bifusepta Darker (1)
Bivallum P.R. Johnst. (7)
Bonanseja Sacc. (1)
Canavirgella W. Merr, Wenner & Dreisbach (1)
Cavaraella Speg. (1)
Ceratophacidium J. Reid & Piroz. (1)
Cerion Massee (2)
Coccomyces De Not. (118)
Colpoma Wallr. (14)
Criella (Sacc.) Henn. (2)
Cryptomyces Grev. (1)
Cudonia Fr. (9)
Davisomycella Darker (10)
Didymascus Sacc. (1)
Discocainia J. Reid & A. Funk (4)
Duplicaria Fuckel (1)
Duplicariella B. Erikss. (1)
Elytroderma Darker (1)
Gelineostroma H.J. Swart (1)
Haplophyse Theiss. (10)
Heufleria Auersw. (1)
Hypoderma De Not. (1)
Hypodermella Tubeuf (3)
Hypodermellina Höhn. (1)
Hypohelion P.R. Johnst. (3)
Irydyonia Racib. (1)

1366
Laquearia Fr. (1)
Lasiostictella Sherwood (1)
Lirula Darker (3)
Lophodermella Höhn. (9)
Lophodermium Chevall. (145)
Lophophacidium Lagerb. (5)
Macroderma Höhn. (1)
Meloderma Darker (9)
Moutoniella Penz. & Sacc. (1)
Mycomelanea Velen. (1)
Myriophacidium Sherwood (6)
Nematococcomyces C.L. Hou, M. Piepenbr. & Oberw. (9)
Neococcomyces Y.R. Lin, C.T. Xiang & Z.Z. Li (3)
Neophacidium Petr. (2)
Nothorhytisma Minter, P.F. Cannon, A.I. Romero & Peredo (3)
Nymanomyces Henn. (1)
Parvacoccum R.S. Hunt & A. Funk (1)
Phaeophacidium P. Henn. & Lindau (4)
Ploioderma Darker (7)
Propolidium Sacc. (2)
Pseudorhytisma Juel (1)
Pseudotrochila Höhn. (2)
Pureke P.R. Johnst. (7)
Rhytisma Fr. (21)
Soleella Darker (7)
Spathularia Pers. (ca. 12)
Sporomega Corda (7)
Terriera B. Erikss. (26)
Therrya Sacc. (7)
Triblidiopsis P. Karst. (2)
Tryblidiopsis P. Karst. (3)
Virgella Darker (1)
Vladracula P.F. Cannon, Minter & Kamal (2)
Xyloschizon Syd. (2)
Zeus Minter & Diamandis (2)

Triblidiaceae Rehm
Huangshania O.E. Erikss. (2)
Pseudographis Nyl. (3)
Triblidium Rebent. (6)

Thelebolales P.F. Cannon


Thelebolaceae A. Engler (=Pseudeurotiaceae Malloch & Cain fide Ekanayaka et al. 2019)
Antarctomyces Stchigel & Guarro (1)
Ascophanus Boud. (20)
Ascozonus (Renny) E.C. Hansen (6)
Caccobius Kimbr. (6)
Cleistothelebolus Malloch & Cain (13)
Coprobolus Cain & Kimbr. (1)
Geomyces Traaen (ca. 10)
Gymnostellatospora Udagawa, Uchiy. & Kamiya (ca. 6)
Holwaya Sacc. (= Crinula Fr.) (2)

1367
Leptokalpion Brumm. (2)
Leuconeurospora Malloch & Cain (3)
Neelakesa Udaiyan & Hosag. (3)
Patinella Sacc. (1)
Pseudascozonus Brumm. (1)
Pseudeurotium J.F.H. Beyma (6)
Pseudogymnoascus Raillo (ca. 10)
Ramgea Brumm. (2)
Thelebolus Tode (13)

Thelebolales genera incertae sedis


Alatospora Ingold (4)
Gorgomyces M. Gönczöl & Révay (2)
Miniancora Marvanová & Bärl. (1)

Leotiomycetes families incertae sedis


Cenangiaceae Rehm#
Cenangium Fr. (30)
Moellerodiscus Henn. (7)
Piceomphale Svrcek (1)

Calloriaceae L. Marchand#
Aivenia Svrček (4)
Belonioscyphella Höhn. (4)
Calloria Fr. (5)
Chaetonaevia Arx (3)
Cistella Quél. (45)
Dactylaria Sacc. (ca. 100)
Diplonaevia Sacc. (23)
Duebenia Fr. (5)
Eupropolella Höhn. (7)
Hyalacrotes (Korf & L.M. Kohn) Raitv. (2)
Iridinea Velen. (2)
Laetinaevia Nannf. (15)
Leohumicola N.L. Nick. (7)
Loricella Velen. (2)
Micropodia Boud. (2)
Mycoarthris Marvanová & P.J. Fisher (1)
Naeviella (Rehm) Clem. (3)
Naeviopsis B. Hein (3)
Ploettnera Henn. (5)
Polyphilus D.G. Knapp, Ashrafi, W. Maier & Kovács (2)
Psilachnum Höhn. (28)
Rodwayella Spooner (3)
Rommelaarsia Baral & Haelew. (1)
Roseodiscus Baral (6)
Stamnaria Fuckel (5)
Tetracladium De Wild. (11)
Urceolella Boud. (24)

Hamatocanthoscyphaceae Ekanayaka & K.D. Hyde


Brachyalara Réblová & W. Gams (1)

1368
Chalara (Corda) Rabenh. (ca. 100)
Ciliolarina Svrček (1)
Gremmenia Korf (4)
Hamatocanthoscypha Svrček (3)
Infundichalara Réblová & W. Gams (2)
Microscypha Syd. & P. Syd. (6)
Pseudohelotium Fuckel (50)
Xenochalara M.J. Wingf. & Crous (1)
Xenopolyscytalum Crous (1)

Hemiphacidiaceae Korf#
Calycellinopsis W.Y. Zhuang (1)
Cenangiopsis Rehm (2)
Chlorencoelia J.R. Dixon (4)
Crumenulopsis J.W. Groves (1)
Didymascella Maire & Sacc. (5)
Encoelia (Fr.) P. Karst. (40)
Fabrella Kirschst. (1)
Heyderia Link (6)
Hysterostegiella Höhn. (10)
Korfia J. Reid & Cain (1)
Rhabdocline Syd. (4)
Sarcotrochila Höhn. (4)
Trochila Fr. (15)
Velutarina Korf (3)

Hyphodiscaceae Ekanayaka & K.D. Hyde


Fuscolachnum J.H. Haines (7)
Hyalopeziza Fuckel (15)
Hyphodiscus Kirschst. (16)
Soosiella Hujslová & M. Kolařík (1)
Scolecolachnum Guatim., R.W. Barreto & Crous (2)
Venturiocistella Raitv. (7)

Leptodontidiaceae Hern.-Restr., Crous & Gené#


Leptodontidium de Hoog. (11)

Neocrinulaceae Crous
Neocrinula Crous (2)

Neolauriomycetaceae Crous#
Exochalara W. Gams & Hol.-Jech. (3)
Lareunionomyces Crous & M.J. Wingf. (2)
Neolauriomyces Crous (1)

Pezizellaceae Velen. (= Bloxamiaceae Locq. ex Hern.-Restr., Gené, R.F. Castañeda, J. Mena,


Crous & Guarro fide Ekanayaka et al. 2019)#
Allophylaria (P. Karst.) P. Karst. (6)
Antinoa Velen. (8)
Bisporella Sacc. (19)
Bloxamia Berk. & Broome (19)
Calycellina Höhn (45)

1369
Calycina Nees ex Gray (= Pezizella Fuckel) (30)
Micropeziza Fuckel (12)
Mollisina Höhn. ex Weese (11)
Mollisinopsis Arendh. & R. Sharma (3)
Moserella Pöder & Scheuer (1)
Orbiliopsis (Sacc. & D. Sacc.) Syd. & P. Syd. (2)
Phaeoscypha Spooner (1)
Phialina Höhn. (ca. 13)
Triposporium Corda (14)
Poculinia Spooner (1)
Scleropezicula Verkley (6)
Velutaria Fuckel (1)
Xiambola Minter & Hol.-Jech. (1)
Zymochalara Guatim., R.W. Barreto & Crous (2)

Rutstroemiaceae Holst-Jensen, L.M. Kohn & T. Schumach. #


Bicornispora Checa, Barrasa, M.N. Blanco & A.T. Martínez (2)
Dencoeliopsis Korf (2)
Lambertella Höhn. (6)
Neometulocladosporiella Crous & M.J. Wingf. (1)
Rutstroemia P. Karst. (ca. 100)

Sclerotiniaceae Whetzel#
Amerosporium Speg. (31)
Amphobotrys Hennebert (1)
Banksiamyces G. Beaton (4)
Botrytis P. Micheli ex Pers. (3)
Ciboria Fuckel (21)
Ciborinia Whetzel (ca. 16)
Clarireedia L.A. Beirn, B.B. Clarke, C. Salgado & J.A. Crouch (4)
Coprotinia Whetzel (1)
Cristulariella Höhn. (5)
Cudoniopsis Speg. (1)
Dumontinia L.M. Kohn (5)
Elliottinia L.M. Kohn (1)
Grovesinia M.N. Cline, J.L. Crane & S.D. Cline (2)
Haradamyces Masuya, Kusunoki, Kosaka & Aikawa (1)
Kohninia Holst-Jensen, Vrålstad & T. Schumach. (1)
Martininia Dumont & Korf (1)
Monilinia Honey (30)
Mycopappus Redhead & G.P. White (4)
Myrioconium Syd. & P. Syd. (10)
Myriosclerotinia N.F. Buchw. (10)
Ovulinia Weiss (9)
Phaeosclerotinia Hori (1)
Poculum Velen. (ca. 22)
Pseudociboria Kanouse (1)
Pycnopeziza W.L. White & Whetzel (5)
Redheadia Y. Suto & Suyama (1)
Sclerencoelia Pärtel & Baral (3)
Scleromitrula S. Imai (6)
Sclerotinia Fuckel (15)

1370
Sclerotium Tode (100)
Seaverinia Whetzel (2)
Septotinia Whetzel ex J.W. Groves & M.E. Elliott (2)
Streptotinia Whetzel (3)
Stromatinia (Boud.) Boud. (15)
Valdensia Peyronel (3)
Valdensinia Peyronel (1)

Vandijckellaceae Sandoval-Denis
Vandijckella Sandoval-Denis (1)

Leotiomycetes family incertae sedis


Porodiplodiaceae Crous
Porodiplodia Crous (1)

Leotiomycetes genera incertae sedis


Adelodiscus Syd. (3)
Algincola Velen. (1)
Apiculospora Wijayaw., Camporesi, A.J.L. Phillips & K.D. Hyde (1)
Aquadiscula Shearer & J.L. Crane (2)
Ascluella DiCosmo, Nag Raj & W.B. Kendr. (1)
Ascoclavulina Otani (8)
Bagnisimitrula S. Imai (1)
Benguetia Syd. & P. Syd. (1)
Bioscypha Syd. (2)
Bulgariopsis Henn. (2)
Callerascus Whitton, K.D. Hyde & McKenzie (1)
Calloriopsis Syd. & P. Syd. (2)
Capillipes R. Sant. (1)
Capricola Velen. (1)
Cejpia Velen. (3)
Cenangiumella J. Fröhl. & K.D. Hyde (1)
Chloroepilichen Etayo (1)
Chlorospleniella P. Karst. (1)
Chondroderris Maire (1)
Ciliella Sacc. & P. Syd. (1)
Coleosperma Ingold (1)
Comesia Sacc. (3)
Cornuntum Velen. (1)
Coronellaria P. Karst. (4)
Criserosphaeria Speg. (1)
Crumenella P. Karst. (1)
Cryptohymenium Samuels & L.M. Kohn (1)
Cryptopezia Höhn. (1)
Dawsicola Döbbeler (1)
Dermateopsis Nannf. (2)
Didonia Velen. (5)
Discomycella Höhn. (1)
Echinodiscus Etayo & Diederich (2)
Epicladonia D. Hawksw. sensu stricto (3)
Episclerotium L.M. Kohn (2)
Erikssonopsis M. Morelet (1)

1371
Flagellospora Ingold (6)
Gloeopeziza Zukal (8)
Godroniopsis Diehl & E.K. Cash (3)
Grimmicola Döbbeler & Hertel (1)
Grovesia Dennis (1)
Helotiella Sacc. (1)
Hemiglossum Pat. (2)
Hymenobolus Durieu & Mont. (3)
Hyphoscypha Velen. (1)
Hysteronaevia Nannf. (12)
Hysteropezizella Höhn. (19)
Involucroscypha Raitv. (10)
Jacobsonia Boedijn (1)
Lasseria Dennis (1)
Livia Velen. (1)
Masseea Sacc. (4)
Melanopeziza Velen. (1)
Melanormia Körb. (1)
Metapezizella Petr. (1)
Micraspis Darker (3)
Microdiscus Sacc. (1)
Monochaetiellopsis B. Sutton & DiCosmo (2)
Mycosphaerangium Verkley (3)
Obconicum Velen. (2)
Obscurodiscus Raitv. (1)
Ocotomyces H.C. Evans & Minter (1)
Otwaya G.W. Beaton (12)
Pachydisca Boud. (1)
Parencoelia Petr. (4)
Patinellaria P. Karst. (1)
Pezolepis Syd. (2)
Pezomela Syd. (1)
Phaeofabraea Rehm (1)
Phragmonaevia Rehm (?1)
Phyllopezis Petr. (1)
Physmatomyces Rehm (1)
Pleoscutula Vouaux (3)
Podophacidium Niessl (2)
Polydiscidium Wakef. (7)
Polydiscina Syd. (1)
Potridiscus Döbbeler & Triebel (1)
Pseudolachnum Velen. (1)
Pseudopeltis L. Holm & K. Holm (1)
Pseudotryblidium Rehm (1)
Psilophana Syd. (1)
Psilothecium Clem. (1)
Pteromyces E. Bommer, M. Rousseau & Sacc. (1)
Pubigera Baral, Gminder & Svrček (1)
Radotinea Velen. (1)
Rhizocalyx Petr. (1)
Rhizothyrium Naumov (1)
Riedera Fr. (1)

1372
Sageria A. Funk (1)
Schnablia Sacc. & P. Syd. (1)
Sclerocrana Samuels & L.M. Kohn (4)
Scutulopsis Velen. (1)
Sinofavus W.Y. Zhuang (1)
Sorokina Sacc. (1)
Sorokinella J. Fröhl. & K.D. Hyde (2)
Spirosphaera Beverw. (8)
Stilbopeziza Speg. (1)
Themisia Velen. (8)
Tovariella Syd. (1)
Trichohelotium Killerm. (2)
Trizodia Laukka (1)
Trullula Ces. (15)
Waltonia Saho (1)
Woodiella Sacc. & P. Syd. (3)
Xeromedulla Korf & W.Y. Zhuang (3)
Zugazaea Korf, Iturr. & Lizoň (1)

Notes for alternative classification of Leotiomycetes

Amorphothecaceae (= Myxotrichaceae) – Ekanayaka et al. (2019) placed this family under


Erysiphales, considering its morphological similarity with other taxa in Erysiphales.
The authors accept the synonymy of Myxotrichaceae under Amorphothecaceae as the close
phylogenetic relatedness of these two families were shown in the phylogeny provided in
Ekanayaka et al. (2019) (A.H. Ekanayaka & K.D. Hyde).

Calloriaceae and Pezizellaceae – In other classification systems, Calloriaceae and Pezizellaceae


are members of Helotiales. Ekanayaka et al. (2019) placed Calloriaceae and Pezizellaceae under
Rhytismatales based on their phylogeny. Johnston et al. (2019) placed these families within
Helotiales. However, the placement of Calloriaceae is not clear. In addition, whereas
Pezizellaceae was retrieved as a monophyletic family in the “pezizelloid clade” of the 15-gene
tree, taxa from this family are polyphyletic in the ITS tree. In the present alternative classification
for Leotiomycetes we accept these families under Leotiomycetes families incertae sedis, until we
have more data to provide a stable classification (A.H. Ekanayaka & K.D. Hyde).

Cenangiaceae, Chlorociboriaceae, Hemiphacidiaceae, Rutstroemiaceae, Sclerotiniaceae –


Those families formed a well-supported distinct clade in Ekanayaka et al. (2019). Therefore, here
we keep them in Leotiomycetes families incertae sedis until we have more data (A.H. Ekanayaka
& K.D. Hyde).

Cochlearomycetaceae – Within the phylogeny of Ekanayaka et al. (2019), this family clustered
within Leotiales, and also Johnston et al. (2019) suggested its position should be within Leotiales
based on their ITS phylogeny. As a result, we placed this family in Leotiales (A.H. Ekanayaka &
K.D. Hyde).

Cordieritidaceae – Taxa of this family differ from other Helotiales by having a unique
ionomidotic reaction (solubility of excipular pigments in KOH). Hence, Jaklitsch et al. (2016)
suggested a separate phylogenetic position for this family away from other Helotiales. In Johnston
et al. (2019), this family clustered in the “sclerotinoid clade” within Helotiales in their 15-gene
tree. In the phylogeny of Ekanayaka et al. (2019), however, this family grouped sister to

1373
Deltopyxidaceae within Cyttariales. Therefore, here we place this family under Cyttariales until
we have more data to confirm its placement (A.H. Ekanayaka & K.D. Hyde).

Deltopyxidaceae – In the phylogeny of Ekanayaka et al. (2019), this family grouped sister to
Cordieritidaceae with strong statistical support within Cyttariales. We are unable to compare with
Johnston et al. (2019), as these authors did not include taxa from this family (A.H. Ekanayaka &
K.D. Hyde).

Erysiphales – In Ekanayaka et al. (2019), this order formed a distinct clade, while in Johnston et
al. (2019) its placement was within Helotiales both in the genomic-scale and 15-gene trees.
However, we believe that this order is distinct as it has a unique morphology, which differs from
taxa in Leotiomycetes. Further studies are needed to resolve this inconsistency (A.H. Ekanayaka &
K.D. Hyde).

Leptodontidiaceae – This family sat in a well-supported clade away from Helotiales in the
phylogeny provided by Ekanayaka et al. (2019). We cannot compare its placement with Johnston
et al. (2019) as their phylogenies did not include taxa from this family. Therefore, we placed this
family in Leotiomycetes families incertae sedis until more data are available to provide a stable
classification (A.H. Ekanayaka & K.D. Hyde).

Medeolariales – Ascocorticiaceae, Ascodichaenaceae, Dermateaceae, Medeolariaceae –


Ekanayaka et al. (2019) accommodated these families in Medeolariales based on morphological
similarities and phylogenetic analysis. In Johnston et al. (2019), Dermateaceae formed the basal-
most clade within Helotiales. However, Johnston et al. (2019) did not include Ascocorticiaceae,
Ascodichaenaceae and Medeolariaceae in their phylogeny, thus, we were unable to compare their
placements in Leotiomycetes. Future studies are needed to resolve the placement of these families
(A.H. Ekanayaka & K.D. Hyde).

Neolauriomycetaceae – This family produces a well-supported clade within “Sclerotiniales” in


the phylogeny of Ekanayaka et al. (2019). We cannot compare its placement with Johnston et al.
(2019) as their phylogenies did not include taxa from this family. Therefore, we placed this family
under Leotiomycetes families incertae sedis until more data become available to provide a stable
classification (A.H. Ekanayaka & K.D. Hyde).

Multi-locus phylogenies and comprehensive treatments of the Xylariales (Author: M. Stadler)


Wendt et al. (2018) provided a multigene genealogy of the stromatic families of the
Xylariales, including a significant number of representative species of the main lineages in the
Xylariaceae and four DNA loci. This study resulted in the segregation of the Xylariaceae sensu
lato in the traditional definition, and the Hypoxylaceae were resurrected and amended. Moreover,
the genera Biscogniauxia and Camillea were transferred to the Graphostromataceae. The
molecular phylogeny corresponds with the distribution of secondary metabolites and types of
conidiogenous structures, while the ascospore morphology, which had traditionally constituted the
salient feature to define the family, had to be abandoned. Notably, the Lopadostomaceae also
contains genera with similar ascospores that had previously been accommodated in the
Xylariaceae sensu lato and some genera like Whalleya and Jumillera were transferred to
Lopadostomataceae.
Wendt et al. (2018) also proposed to exclude several genera of which no information on the
asexual morph and no molecular data are available, from the Xylariaceae and place them at
interim in Xylariales incertae sedis. Moreover, they resurrected the genus Pyrenonopolyporus for
some species formerly placed in Hypoxylon with massive stromata and long tubular perithecia and
segregated the new genus Jackrogersella from Annulohypoxylon as an outcome of their
polyphasic study. Concurrently with the study by Wendt et al. (2018), Daranagama et al. (2018)

1374
provided a very comprehensive overview on the families of the “stromatic” Xylariales, which
roughly comprise the genera that had traditionally been classified in the Xylariaceae according to
the traditional concept of the late 1990s, which was based on ascospore morphology. In the first
comprehensive study of this type, they provided illustrations of most of the type species or other
representative species and revised the history of their taxonomy. A molecular phylogeny was also
presented, using data from additional taxa that represented genera that were not included in the
paper by Wendt et al. (2018). Therefore, certain deviations from the concept were observed. In
addition Daranagama et al. (2018) retained some genera in the Xylariaceae that were expelled
from the family in the concurrent study. Helaly et al. (2018) reviewed the taxonomy, ecology and
in particular the secondary metabolite production of the Xylariales. Even though no taxonomic
novelties are provided in this paper, the records of endophytic and marine strains of Xylariales
that had previously been reported in the literature to produce novel bioactive compounds were
revised and numerous incongruities between the data published in chemistry journals and the
current taxonomy were found. The authors gave an overview on the current taxonomic status of
the taxonomy of the order in relation to the numbers of known metabolites and pointed out some
genera and families that deserve further study as they seem to be hitherto unexplored. Recently,
the genus Dematophora has been resurrected and divided from Rosellinia based on a comparison
of molecular data, morphology of the asexual morphs and chemotaxonomic evidence (Wittstein et
al. 2020), and all the serious pathogens like D. necatrix and D. bunodes now no longer belong to
Rosellinia, which mainly comprises saprotrophic and endophytic species. Finally, Samarakoon et
al. (2020) have recently found out that the sexual morph of the economically important
endophytic genus Muscodor is close to Induratia, which was erected earlier and takes preference
over the latter genus. They provided a polyphasic study involving a multi-locus phylogeny,
divergence time estimations, morphological studies and comparison of volatile secondary
metabolite profiles, which resulted in the recgnisiton of a new family Induratiaceae in the order
Xylariales. This family is sister to the Xylariaceae sensu stricto and also includes the genus
Emarcea. All names in Muscodor were newly combined in Induratia, and two new species were
described with their full life cycle, comprising both the asexual and sexual morph.

Placement and phylogenies of Laboulbeniomycetes (Author: D. Haelewaters)


The placement of Laboulbeniomycetes has been traditionally problematic (Blackwell et al.
2020). First considered as cuticle hairs or even parasitic worms, life history studies and molecular
phylogenetic analyses have helped in placing these fungi among filamentous Ascomycota. The
first phylogenetic analysis of the class by Weir & Blackwell (2001) was based on SSU rDNA and
supported the placement of Laboulbeniales–Pyxidiophora as a separate clade within Ascomycota,
sister to Sordariomycetes although without statistical support for this sister relationship. Schoch et
al. (2009) were the first ones to obtain support for the sister relationship between
Laboulbeniomycetes and Sordariomycetes based on a six-locus dataset. Their results were later
confirmed by Goldmann & Weir (2018) as well as Haelewaters et al. (2019b) who proposed the
informal taxon ‘laboulbeniomyceta’ as a descriptor of the most recent common ancestor of both
classes.
To date, limited sequence data are available for members of Laboulbeniomycetes. An NCBI
GenBank search for ‘Laboulbeniomycetes’ resulted in 727 sequences only, the majority of which
are SSU sequences (17 February 2020). An SSU-based phylogeny by Goldmann & Weir (2018)
including 65 isolates resulted in evidence for multiple clades in the class. Haelewaters et al.
(2019b), based on a three-locus dataset with 61 isolates, described the third order in the class,
Herpomycetales, to accommodate the genus Herpomyces. Blackwell et al. (2020) presented a
phylogenetic reconstruction of the Laboulbeniomycetes from a concatenated SSU–LSU dataset of
75 isolates and found high support for 5 clades: orders Herpomycetales, Laboulbeniales and
Pyxidiophorales, in addition to two unnamed clades, Chantransiopsis clade (Chantransiopsis sp.,
Tetrameronycha spp., Subbaromyces splendens) and Laboulbeniopsis clade (with Laboulbeniopsis
termitarius). Finally, several recent studies have pointed at the polyphyly of accepted higher taxa

1375
in the Laboulbeniales order (Goldmann & Weir 2018, Haelewaters et al. 2018b), and so future
integration of molecular data will undoubtedly change the classification of the order and by
extension the whole class.

Structural and functional organization of Microsporidia (Author: Yuri S. Tokarev)


Microsporidia is a monophyletic group of highly specialized intracellular parasites which
are ultimately dependent upon and being able to develop only within the host cell (Vavra & Lukes
2013). Microsporidia infect Metazoa and some protists: Gregarina, Ciliata, Paramyxea (Fokin et
al. 2008, Sokolova et al. 2013, Larsson 2014, Cali et al. 2017, Stentiford et al. 2017). Host
switching between representatives of different families (Ghani et al. 2013, Malysh et al. 2018a,
2019), orders (Schuld et al. 1999, Ovcharenko et al. 2017, Malysh et al. 2018b), classes (Hinney
et al. 2016) and phyla may occur (Franzen et al. 2006, Ironside et al. 2008, Nylund et al. 2010,
Choudhary et al. 2011, Meissner et al. 2012). Parasite-host interactions are diverse, ranging from
asymptomatic presence in the form of a latent infection to devastating epizootics especially in
arthropods and fishes (Bader et al. 1998, Stentiford et al. 2013, Sokolova et al. 2015, Yu et al.
2019). Routes of transmission include alimentary and transovarial/ transplacental infection,
transfer from host to host by parasites etc. (Becnel & Andreadis 1999, Dunn & Smith, 2001,
Didier et al. 2004, Wang-Peng et al. 2018).
Typical life cycle includes merogonial (for parasite multiplication) and sporogonial
sequence (for multiplication and spore formation). The cells multiply either by binary or multiple
fission. The merogonial developmental stages are single cells or plasmodia delineated with a
plasma membrane (Dunn & Smith 2001). In early sporogonial stages, cytoplasm becomes more
condensed, electron dense material is deposited on the cell surface which further transforms into
the spore wall. Other primordial spore structures emerge in late sporogonial stages (Issi et al.
2012a, Sokolova et al. 2015, Cali & Takvorian 2014). The nuclear apparatus is mono- or
diplokaryotic, the number of nuclei varies from one to many (in plasmodia). In some species the
transformation of the nuclear apparatus from one type to another one takes place (sometimes
accompanied by meiosis) in the course of life cycle (Lee et al. 2014). The parasites develop either
in direct contact with the host cell cytoplasm or within a parasitophorous vacuole which is derived
from the membranes of endoplasmic reticulum, Golgi complex or nuclear envelope of the host
cell (Bohne et al. 2011, Issi et al. 2012b, Vavra & Lukes 2013).
The spore is a specialized parasite cell which serves for invasion into the host cell,
dissemination over the host organism and spreading within the host population (Vavra & Lukes
2013). The spore wall is a complex chitin- and protein-rich structure consisting of inner electron-
translucent endospore layer with an underlying plasma membrane and outer electron-dense
exospore layer (Bigliardi & Sacchi 2001). The main distinctive feature of microsporidia spore is
the extrusion apparatus, which includes polar tube (polar filament), polar cap (polar sac –
anchoring disc complex), polaroplast and posterior vacuole (Fig. 2) (Vavra & Larsson 2014). The
polar tube is attached to the apical pole of the spore by the polar cap (Franzen 2004, Xu & Weiss
2005). The length, diameter and structure of polar tube vary greatly among species. Most often,
the polar tube is thin and long, exceeding the length of the spore, with proximal and distal parts
straight and coiled, respectively (Delbac & Polonais 2008, Issi et al. 2012a).

1376
Figure 2 – Spore structure of Microsporidia with different types of polar tube. A Metchnikovella
incurvata with a manubrial polar tube. B Mrazekia macrocyclopis with a manubrial polar tube
containing a coiled distal part. C Helmichia lacustris with an uncoiled polar tube (UPT).
D Neoperezia sp. with numerous coils of isofilar polar tube. E Crepidulospora beklemishevi with
anterior (PTC1) and posterior (PTC2) polar tube coils of different structure and size. AD –
anchoring disc; M – manubrium; PS – polar sac; PTC – polar tube coils. Figs 2A, 2E are
reproduced with a kind permission of Yuliya Y. Sokolova (Institute of Cytology RAS, St.
Petersburg, Russia) and Anastasia V. Simakova (Tomsk State University, Tomsk, Russia),
respectively.

The number of coils ranges from 3 to 30. In isofilar polar tube, the diameter is the same
throughout the tube length. It is either coiled (Fig. 2D) or uncoiled (Fig. 2C), as its length exceeds
or do not exceed the spore length, respectively (Issi et al. 2012a, Tokarev et al. 2012). In anisofilar
polar tube, several posterior coils are of lesser diameter and sometimes of different electron
density (Tokarev et al. 2010). In heterofilar polar tube, the anterior coils are remarkably bigger as
compared to the posterior ones and their structure is different (Fig. 2E) (Vavra & Larsson 2014).
The manubrium is thick and short, not exceeding the spore length, cylindrical, usually thicker at
central or distal ‘bulbal’ part, which is interconnected with manubrial cisternae associated with
vesicles and/or short tubules (Fig. 2A) (Issi et al. 2010, Sokolova et al. 2013). Another type of
polar tube is clavate, which is thick and short, not exceeding the spore length, thicker at distal
‘bulbar’ part, which is continued into thin polar tube with several coils (Fig. 2B). This clavate
polar tube is sometimes referred to as “manubrium” (Issi et al. 2010, which is not correct. The
manubrium is present in Rudimicrosporea (Sokolova et al. 2013), being different in its structure
and supposed to be a primitive form of the canonical polar tube of higher Microsporidia (Vivier
1975). The polaroplast is a lamellar or vesicular (or combined), occupying the anterior part of the
spore and surrounding the straight part of the polar tube (Vavra & Larsson 2014). Its function is
unclear and participation in extrusion process is supposed. The spores are formed as separate units
(“free spores”) or by packets enclosed within a common envelope of parasite origin, so called
sporophorous vesicle (Dunn & Smith 2001, Sokolova et al. 2015).

1377
Figure 3 – Alternative hypotheses of phylogenetic and taxonomic relationships within Fungs-like
representatives of Aphelida (Aphelidiomycota) – Rozellomycota (Rozellida/ Cryptomycota/
Rozellosporidia) – Microsporidia (ARM) clade and its relations to Fungi. The schematic
phylogenetic trees follow the recent phylogenetic reconstructions by Karpov et al. (2014),
Tedersoo et al. (2018), Bass et al. (2018), Corsaro et al. (2019, 2020). Three alternative schemes
are presented as nowadays there is no consensus among the specialists. A Karpov et al. (2014)
proposed the superphylum Opisthosporidia for the ARM clade, which was considered as a sister
to Fungi. Several new genera related to Rozella and Microsporidia have been added to the tree
more recently (Corsaro et al. 2014a, b, 2016, 2019, 2020, Haag et al. 2014). B In the interpretation
of Bass et al. (2018) the boundaries of Microsporidia should be expanded to include all these
genera except Rozella. C According to Tedersoo et al. (2018); Wijayawardene et al. (2018) ARM
clade should be included into kingdom Fungi, and phylum Microsporidia should be merged with
Rozellomycota. To further conserve Microsporidia as a monophyletic taxon within Rozellomycota
it has been redefined as a subphylum. The labels for taxon ranks and phyla abbreviations are
explained in the right lower corner. The synonymic names for the phyla are indicated:
Aphelidiomycota Tedersoo et al 2018 = Aphelida Karpov, Aleoshin & Mikhailov 2014 =
Rozellida Lara, Moreiro, López-García, 2010 = Rozellomycota James & Berbee, 2011 emend.
Corsaro & Michel 2014 = Cryptomycota Jones & Richards 2011 emend. Karpov & Aleoshin 2014
= Rozellosporidia Karpov, Torruella, Moreira, Mamkaeva, López-García 2017.

Many of the basic physiological functions and biochemical processes in Microsporidia are
reduced as they mostly rely on the host cell. As a result of deep adaptation to intracellular
parasitism, structural organization of prespore stages is simplified, and many biochemical

1378
pathways are absent. Basic nutrients are imported from the host cell including ATP (Tsaousis et
al. 2008). The physiological processes in the host cell are altered and can be managed by the
actively proliferating parasites to provide the latter with enough nutrients including sugars, amino-
and fatty acids (Cuomo et al. 2012, Senderskiy et al. 2014). Exploitation of host cell metabolic
pathways results in reorganization of host cell organelles: mitochondria, endoplasmatic reticulum
and Golgi complex, which come into contact with the parasite cells (Simakova et al. 2005,
Tokarev et al. 2010, Issi et al. 2012b, Vavra & Lukes 2013). Other adaptations to survival within
the host cell include alteration of host cell cycle, suppression of host cell apoptosis (Martin-
Hernandez et al. 2017, Sokolova et al. 2019) and renewal of infected epithelial cells (Issi 1986).
Mitochondria are reduced to mitosomes which lack genome (Williams et al. 2002).
Vesicular transport of cargo proteins is absent (Beznoussenko et al. 2007). The genomes of
Microsporidia are reduced and compact, many genes and gene families are lost and the sequences
of ribosomal RNA and protein-coding genes are shortened and highly derived (Peyretaillade et al.
2015). The genome size is ranged from 2.3 to ~50 Mb, in the vast majority of sequenced species it
does not exceed 15 Mb (Keeling et al. 2014).

Phylogenetic approaches and current status of Rozellomycota (Author: Yuri S. Tokarev)


With the advent of molecular phylogeny studies, major groups within Microsporidia Tree of
Life were primarily recognized and polyphyletic nature of traditional high rank taxa was
demonstrated (Fig. 3) (Vossbrinck & Debrunner-Vossbrinck 2005, Vossbrinck et al. 2014). Later
on, some basal or sister groups of parasites were examined using molecular phylogenetic and
phylogenomic approaches. In particular, relationships between Aphelida, Rozellomycota and
Microsporidia were established (Karpov et al. 2014). Several new representatives of
Rozellomycota were discovered and examined (Haag et al. 2014, Corsaro et al. 2014a, b, 2016,
Quandt et al. 2017). Genome surveys of Metchnikovellida were provided (Mikhailov et al. 2016,
Galindo et al. 2018) confirming basal position of this lineage within Microsporidia and
demonstrating shared genomic signatures with other Microsporidia. Notably, the order of
Metchnikovellida was always referred to as “primitive Microsporidia” (class Rudimicrosporea) on
the basis of ultrastructural features, such as poorly developed or absent polaroplast and absent
posterior vacuole as well as specific structure of polar tube, so called manubrium. Finally,
phylogenetic position of Chytridiopsis typographi (order Chytridiopsidea) was also resolved
(Corsaro et al. 2019), showing more basal position than Metchnikovellida. In this parasite species,
only the sequences of rRNA genes are available, showing less compact organization as compared
to other Microsporidia. In addition, spore structure of C. typographi is described as “unique” as
compared to other Microsporidia (Purrini & Weiser 1985) and essentially, the developmental
sequence of C. typographi includes a unique budding stage, unknown for other species of
Microsporidia, which multiply by fission of cells only (Tonka et al. 2010). Thus, ultrastructural
and developmental traits support the basal position of Chytridiopsida inferred in the phylogenetic
reconstructions. Here the order Chytridiopsida includes the composition of the families and
genera suggested by Larsson (2014) with exclusion of Burkeidae, which is less similar to the
chytridiopsids and was attributed to this group only temporaly. The mentioned taxon is
provisionally placed to Microsporidia genera incertae sedis as possessing more canonical
ultrastructure.
Recently, Bass et al. (2018) proposed taxonomic expansion of Microsporidia to include
unclassified genera of Cryptomycota (Rozellomycota). This assumption was based upon
monophyletic arrangement of Microsporidia and sister taxa, though the vast majority of sequences
used in that analysis were short (below 400 bp). We prefer to preserve Microsporidia as a more
compact group unless more robust phylogenies are available. Thus, such representatives of
Rozellomycota as Nucleophaga, Paramicrosporidium, Morellospora and Mitosporidium are not
considered here as Microsporidia.
Upon following the ranking of Tedersoo et al. (2016) who proposed Rozellomycota
(including Microsporidia) and Aphelidiomycota as phyla in Kingdom Fungi, the phylum

1379
Microsporidia becomes obsolete. To recognize this group as a monophyletic lineage of highly
specialized intracellular parasites with specific developmental, structural and genetic features (see
above, Structural and functional organization of Microsporidia), we propose an intermediate rank
of subphylum for this taxon. The superphylum Opisthosporidia proposed by Karpov et al. (2014)
is also retained to include Aphelidiomycota and Rozellomycota. However recent transcriptomic
data provide poor support for the monophyly of Opisthosporidia (Toruella et al. 2018), and in
future the aphelids may be excluded and the higher ranks may be again reshuffled.
The Microsporidia Tree of Life consists of five major clades (Vossbrinck & Debrunner-
Vossbrinck 2005, Vossbrinck et al. 2014). To accommodate these clades within a formal class-
order-family system, the following changes to the taxonomy of Microsporidia are proposed.
Orders Dissociodihaplophasida and Meiodihaplophasida are suppressed as polyphyletic. Orders
Glugeida and Nosematida are redefined for the purposes of the current revision. Three new orders,
Amblyosporida ord. nov., Neopereziida ord. nov. and Ovavesiculida ord. nov. are introduced in
this study. All these orders are referred to as distinct monophyletic clades of the Rozellomycota
Tree of Life, each presented by the type family: Glugeidae, Nosematidae, Amblyosporidae,
Neopereziidae and Ovavesiculidae, respectively, and related taxa. Order Chytridiopsidea is
excluded from Microsporidia (see the outline) and family Enterocytozoonidae is transferred to
Nosematida. Allocation of order Metchnikovellida to class Rudimicrosporea is also supported.
Each phylogenetic lineage corresponding to the taxa of class, order or family rank contains
representatives with diverse features. This makes it difficult to define these taxa using
developmental, structural or ecological features, so that primary taxonomic allocation of species is
based upon phylogeny.

Classification of Glomeromycota (Authors: B.T. Goto & N. Wijayawardene)


Outline of arbuscular mycorrhizal fungi (AMF) was initially organized in Gerdemann &
Trappe (1974) and updated by Schenck & Peréz (1990). Initially, AMF fungi have been included
in one order (i.e. Glomerales) in Zygomycota (Morton & Benny 1990). However, Schüßler et al.
(2001) established new phylum, Glomeromycota to accommodate AMF. Since Schüßler et al.
(2001), taxonomy of AMF has been rapidly updating over the last years (Oehl et al. 2011a, b, c, d,
e, f, Goto et al. 2012, Błaszkowski et al. 2015, 2018a, b, 2019a, b, Corazon-Guivin et al. 2019a, b,
Jobim et al. 2019). Oehl et al. (2008) introduced Dentiscutataceae, Racocetraceae and
Scutellosporaceae and later Oehl et al. (2011b) introduced Gigasporales, to include these families
along with Gigasporaceae. At the same time, Oehl et al. (2011b) introduced two new classes (i.e.
Archaeosporomycetes and Paraglomeromycetes) thus currently Glomeromycota comprises three
classes.
Morton & Msiska (2010) did not recognize families introduced in Oehl et al. (2008) and
Gigasporales was not accepted in Wijayawardene et al. (2018b). Nevertheless, phylogenetic
reconstructions using different genes (TUB2, RPB1, ITS, SSU and LSU rRNA) (e.g. Goto et al.
2012, Mello et al. 2012, Silva et al. 2012, Marinho et al. 2014, de Souza et al. 2018, Tedersoo et
al. 2018) supported the monophyletic nature of the families and or genera proposed by Oehl et al
(2008). Hence, we conclude that higher level classification of families in Glomeromycota is
debatable. Therefore, in this study, we include an alternative classification for Glomeromycota
which was included in Wijayawardene et al. (2018b).

Alternative classification for Glomeromycota

Glomeromycota C. Walker & A. Schüßler


Archaeosporomycetes Sieverd., G.A. Silva, B.T. Goto & Oehl
Archaeosporales C. Walker & A. Schüßler
Ambisporaceae C. Walker, Vestberg & A. Schüßler
Ambispora C. Walker, Vestberg & A. Schüßler (10)

1380
Archaeosporaceae J.B. Morton & D. Redecker
Archaeospora J.B. Morton & D. Redecker (= Intraspora Oehl & Sieverd.) (3)
Palaeospora Oehl, Palenz., Sánchez-Castro & G.A. Silva (1)

Geosiphonaceae Engl. & E. Gilg


Geosiphon F. Wettst. (1)

Glomeromycetes Caval.-Sm.
Diversisporales C. Walker & A. Schüßler
Acaulosporaceae J.B. Morton & Benny
Acaulospora Gerd. & Trappe (= Kuklospora Oehl & Sieverd.) (57)

Diversisporaceae C. Walker & A. Schüßler


Corymbiglomus Błaszk. & Chwat (2)
Desertispora Błaszk., Kozłowska, Ryszka, Al-Yahya’ei & Symanczik (1)
Diversispora C. Walker & A. Schüßler (18)
Otospora Oehl, Palenz. & N. Ferrol (1)
Redeckera C. Walker & A. Schüßler (3)
Tricispora Oehl, Sieverd., G.A. Silva & Palenz. (1)

Gigasporaceae J.B. Morton & Benny


Bulbospora Oehl & G.A. Silva (1)
Cetraspora Oehl, F. A. Souza & Sieverd. (6)
Dentiscutata Sieverd., F.A. Souza & Oehl (7)
Gigaspora Gerd. & Trappe (7)
Intraornatospora B.T. Goto, Oehl & G.A. Silva (1)
Paradentiscutata B.T. Goto, Oehl & G.A. Silva (2)
Racocetra Oehl, F.A. Souza & Sieverd. (14)
Scutellospora C. Walker & F.E. Sanders (21)

Pacisporaceae C. Walker, Błaszk., A. Schüßler & Schwarzott


Pacispora Sieverd. & Oehl (6)

Sacculosporaceae Oehl, Sieverd., G.A. Silva, B.T. Goto, Sánchez-Castro & Palenz.
Sacculospora Oehl, Sieverd., G.A. Silva, B.T. Goto, I.C. Sánchez & Palenz. (2)

Glomerales J.B. Morton & Benny


Claroideoglomeraceae C. Walker & A. Schüßler
Claroideoglomus C. Walker & A. Schüßler (8)

Glomeraceae Piroz. & Dalpé


Dominikia Błaszk., Chwat & Kovács (13)
Funneliformis C. Walker & A. Schüßler (16)
Glomus Tul. & C. Tul. (55)
Halonatospora Błaszk., Niezgoda, B.T. Goto & Kozłowska (1)
Kamienskia Błaszk., Chwat & Kovács (1)
Oehlia Błaszk., Kozłowska, Niezgoda, B.T.Goto & Dalpé (1)
Rhizoglomus Sieverd., G.A. Silva & Oehl (19)
Rhizophagus P.A. Dang. (ca. 19)
Sclerocystis Berk. & Broome (10)
Sclerocarpum B.T. Goto, Błaszk., Niezgoda, Kozłowska & Jobim (1)
Septoglomus Sieverd., G.A. Silva & Oehl (10)

1381
Simiglomus Sieverd., G.A. Silva & Oehl (1)

Glomeromycetes genus incertae sedis


Entrophospora R.N. Ames & R.W. Schneid. (3)

Paraglomeromycetes Oehl, G.A. Silva, B.T. Goto & Sieverd.


Paraglomerales C. Walker & A. Schüßler
Paraglomeraceae J.B. Morton & D. Redecker
Paraglomus J.B. Morton & D. Redecker (1)
Innospora Błaszk., Kovács, Chwat & Kozłowska (7)

Pervetustaceae Błaszk., Chwat, Kozłowska, Symanczik & Al-Yahya’ei


Pervetustus Błaszk., Chwat, Kozłowska, Symanczik & Al-Yahya’ei (1)

Oomycota (Author: M. Thines, with contributions from H.P. Grossart)


Traditionally, the Oomycota (from Greek ωάριο (egg) and μύκης (fungus) and were treated
with other osmotrophic aseptate organisms in the polyphyletic “Phycomycetes”. However, already
at the turn from the 19th to the 20th century, it was known that the oomycetes differed in various
aspects, and chlor-zinc-iodine solution, which gives a blue staining with cellulose, was widely
used to identify oomycete thalli in their hosts (e.g. Petersen 1905). Already in 1939, just one year,
after the first electron microscope became commercially available, Vlk (1939) reported that the
zoospores of Saprolegnia did not resemble motile spores of fungi, but rather of heterokont algae.
Since then, it was widely recognised that oomycetes belonged within heterokonta, which were
suggested as an independent kingdom by Leedale (1974), largely congruent with the kingdom
Straminipila, introduced by Dick (2001). By emphasizing periplastid protein targeting,
cytoskeletal and periplastid evolution, Cavalier-Smith 2018 proposed a new classification,
assuming that Heterokonta, Alveolata, Rhizaria, and Hacrobia (cryptophytes and haptophytes) are
a monophylum which could be recognized as a re-circumscribed kingdom Chromista. However,
there is still some controversy on the precise phylogenetic relationships between the different
groups, in particular haptophytes and cryptomonads may not belong together with the heterokonts
or the SAR clade, but be associated with the Archaeplastida (Burki et al. 2007, 2016).
Thus, oomycetes are currently best placed in the kingdom Straminipila and its phylum
Oomycota.The kingdom Straminipila forms a major line of eukaryotes (e.g. van den Hoek et al.
1995) currently containing more than 25,000 known species, most of which belong to the two
major phyla with photosynthetic members, Phaeophyta and Bacillariophyta. The osmotrophic
Oomycota are, with about 1700 recognised species, the next largest phylum of the Straminipila.
The main chracterisitic of the Straminipila is the formation of stiff hairs, one of the two flagella,
which strengthens and reverses its thrust, rendering oomycete spores excellent swimmers. These
stiff “straw hairs” gave their name to the whole group, derived from “stramen” (straw) and “pilus”
(hair). The majority of the species of the Oomycota are parasitic (Beakes & Thines 2017), with
Phytophthora infestans, the organism that triggered the Irish Potato Famine, being its most
prominent member (Yoshida et al. 2013). Most Oomycota show an asexual reproduction via
zoospores formed in zoosporangia or, in case of some genera of the obligate biotrophic downy
mildes, such as Peronospora, Hyaloperonospora, and Bremia, via multinucleate, wind-dispersed
conidiosporangia. Apart from the holocarpic oomycetes, in which the entire thallus is converted
into a zoosporangium, asexual sporangia and conidiosporangia are formed by specialized parts of
the mycelium, i.e. sporogenous hyphae, sporangiophores or sporangiophores (Beakes & Thines
2017). The holocarpic nature is the ancestral form for the whole group, but likey derived in
members of Lagenidiales (Buaya & Thines 2020). Sexual reproduction in most of the species of
the crown classes, Peronosporomycetes and Saprolegniomycetes, is by oogamy
(gametangiogamy), whereby the zygote often converts into a thick-walled resting spore. However,
in many species of the Leptomitales sexual reproduction is cryptic, e.g. by zoomeiospore fusion in

1382
Lagenisma coscinodisci (Schnepf et al. 1977). Leptomitales are also unusual in that some have a
significant amount of chitin in addition to glucans and cellulose in their cell walls (Lin & Aronson
1970). The orders of the oomycetes that branch before the crown classes are, with the exception of
Haliphthorales not forming an extended hyphal network but, apart from the fragmenting thalli in
some members of Pontismatales (Buaya et al. 2019), only simple globose to sacculate thalli that
convert into zoosporangia. The oospore-like resting structures formed by Olpidiopsidales are
likely reflecting convergent evolution to the crown groups. The oomycetes have likely origninated
in the marine environment more than 500 million years ago, have colonised limnic and terrestrial
habitats several times independently and are now found in almost any environment in which
eukaryotic organisms strive (Marano et al. 2016). Most of the known species belong to the
probably monophyletic downy mildews (Sharma et al. 2015), which are obligate biotrophic plant
parasites that are similar to other obligate parasite groups, have diversified by host jumps,
radiation and subsequent speciation (Choi & Thines 2015, Thines 2019). However, it can be
assumed that across all oomycete groups only a small fraction of the existing species are known.

Myxomycetes (Author: D. Leontyev)


Even the limited phylogenetic data currently available support the conclusion that some very
conspicuous characters traditionally considered as important for the classification of myxomycetes
have evolved several times independently. This is the case for both the formation and reduction of
the stalk and the capillitium, the crystallization of lime deposits, and the evolution of compound
fruiting bodies (Leontyev et al. 2014, Leontyev & Schnittler 2017). All of these characters were
used in the traditional system of classification to delimit genera, families and even orders (as was
the case for the order Liceales s. l., which was characterized by the absence of capillitium).
However, in the light of the phylogenetic data, other characters appear to be better predictors of
evolutionary relationships than the traditional criteria. These are, for example, the attachment of
the capillitium to the peridium, details of stalk formation, color of the spore mass, or the presence
of spore-like cells within the stalk. These new criteria were used to provide emended descriptions
for some of the traditional myxomycete taxa in the recently published classification by Leontyev
et al. (2019).

Dictyostelids (Author: D. Leontyev)


The new system of classification for dictyostelids works well for the genus level; however,
it relies on 18S gene markers to distinguish species, and this does not always separate out some
morphologically similar but genetically different taxa. Moreover, Coremiostelium and Synstelium
still occupy a problematic position in the entire phylogeny (Sheikh et al. 2018). A consideration of
morphological characteristics is essential for evaluating new species of dictyostelids. Identifying
new genetic markers to use for distinguishing dictyostelids at the species level would be
exceedingly worthwhile.

Hidden taxa of the fungal tree of life (Authors: F.Q. Brearley & D. Haelewaters)
Whereas four to five million species of fungi are estimated to be found across the globe, we
have only described around 144,000 of these (i.e. less than 2%) (Willis et al. 2018). The question
remains where are all these fungi yet to be described can be found? asa Lücking (2017) suggest
that these missing fungi will be found in: i) habitats that are naturally diverse yet poorly explored,
e.g., tropical forests; ii) cryptic taxa, those that are morphologically indistinguishable; and iii) in
fungal collections that might contain cryptic or new species hidden under current names. These
three categories are not mutually exclusive and, with the rapid rise of next-generation sequencing,
we can also add iv) molecular novelties. A final category that is often neglected is the study of
natural history collections of organisms other than fungi (plant herbaria, dried insect collections).
Large collections of dried, pinned insects have been dubbed “treasure troves” for the study
of ectoparasitic fungi, such as the Laboulbeniales (Laboulbeniomycetes). During a study of
different systematic insect collections around the world, Laboulbeniales fruit bodies were

1383
discovered on 1,937 of 45,785 specimens (Haelewaters & Rossi 2017). This and other insect
collections-based works have led to the description of multiple new species (e.g., Santamaria et al.
2016, Haelewaters & Rossi 2017), a better understanding of host usage patterns and global
number of Laboulbeniales species (Weir & Hammond 1997), and estimates of parasite
prevalences on a given host through time (Haelewaters et al. 2017). Likewise, plant herbaria can
be screened for fungal “hitchhikers” inadvertently sampled along with the plant host (Lang et al.
2019). Examples are the causal agent of potato late blight (Phytophthora infestans), rust fungi,
downy mildews, etc. Herbarium specimens allow for identification of fungal strains based on
morphology and DNA sequence data, which can be coupled with host plant studies to provide a
complete overview of host-pathogen dynamics.
Given that fungi are often microscopic, morphologically similar among species, occupy
hidden habitats, and are often recalcitrant in axenic culture, it is difficult to find them, but perhaps
they are simply ‘hiding’ in material already collected. Lücking et al. (2014) found that a single
species of Cora was actually more than one-hundred species following more careful
morphological observation and sequencing. This restudy/resequencing of currently known taxa is
likely to increase the number of known species. An increasing number of examples of cryptic
diversity is coming to light with the advances of molecular phylogenetic studies, in different
groups, e.g., Eurotiomycetes (Pringle et al. 2005), Laboulbeniomycetes (Haelewaters et al. 2018a),
Lecanoromycetes (Singh et al. 2015) and Leotiomycetes (Grünig et al. 2008) in Ascomycota;
Agaricomycetes (Stefani et al. 2014, Accioly et al. 2019), and Ustilaginomycetes (Li et al. 2017)
in Basidiomycota.
Because of the decreasing price and increasing availability, large-scale sequencing studies
offer the opportunity to mined data for new species. These large-scale sequencing studies offer the
opportunity to find new species as such studies regularly have a sizeable proportion of sequences
that can only assigned at a phylum or even kingdom level. Indeed, the rate of accumulation of
novel/unassigned sequences is massively outpacing the rate with which taxonomists can describe
new species, especially given the lack of taxonomic expertise among, for example, the basal
fungal clades.
Whilst some have called for a sequence-based taxonomy (Hibbett et al. 2016) we recognise
that this is a controversial subject and, instead, note that these sequences can be used in a
‘taxonomic feedback’ capacity to identify new taxa. For example, Rosling et al. (2011) and Jones
et al. (2011) described the class of ubiquitous soil-inhabiting fungi, the Archaeorhizomycetes, and
the basal fungal phylum the Cryptomycota, respectively. Capitalising on the global sequencing of
soil samples by Tedersoo et al. (2014), Tedersoo et al. (2017) re-analysed samples and found
nearly 40 strongly supported new clades (roughly equivalent to orders) in the fungal tree of life.
About half of these were in the basal clades (particularly Rozellomycota) and about half were
found in tropical habitats. This study looked at soil samples only but fungi associated with other
habitats such as aquatic fungi or endophytic fungi would benefit from further study – aquatic
habitats may be particularly rich in new clades and taxa given the association of basal clades with
aquatic habitats. The rich resource of DNA extracted from the millions of plant leaves stored in
herbaria can also be fruitfully mined to find new plant-associated fungi (e.g. Datlof et al. 2017,
Daru et al. 2018).
Another possibility is that the molecular markers we are currently using are not sufficient to
delineate the full range of ‘species’ (OTUs, RSVs etc.) – this is certainly the case for
Glomeromycota where the ITS region is not sufficiently phylogenetically informative and
development and comparison of suitable primers has occupied a lot of energy (e.g. Kohout et al.
2014).
Whilst some have called for a molecular taxonomy, we recognise that this is controversial
and, as yet, not incorporated into relevant taxonomic codes. Therefore, discovery of new physical
specimens is required and the work of Henkel et al. (2012) and Hyde et al. (2018b) illustrate how
dedicated work of mycologists over a number of years can lead to the discoveries and descriptions
of numerous new fungal species. Indeed, Truong et al. (2017) show how this can be done with

1384
straightforward short collecting expeditions of underexplored habitats (Nothofagus forest of South
America in this case) combined with taxonomic work and sequencing of relevant material to
advance our knowledge of fungal diversity in a comprehensive way.

Notes

Abrothallus De Not.
The phylogenetic reconstruction by Suija et al. (2015) showed that Abrothallus
accommodates the monotypic Epinephroma Zhurb. and made a new combination based on that.
Pérez-Ortega et al. (2011) showed that Vouauxiomyces Dyko & D. Hawksw. represents an asexual
stage of Abrothallus. Synonymization with Abrothallus was done in Rossman et al. (2016) (A.
Suija).

Abrothallaceae Pérez-Ort. & Suija


Diederich et al. (2018) proposed to synonymize Licheniconiaceae with Abrothallaceae as
several studies beforehand (e.g. Liu et al. 2017) have revealed close relationships of these two
monotypic families. However, Hngsanan et al. (2020) accepted Licheniconiaceae over
Abrothallaceae (A. Suija & N. Wijayawardene).

Acrodictyaceae J.W. Xia & X.G. Zhang


Xia et al. (2017) established Acrodictyaceae and Junewangiaceae in Sordariomycetes (J.
Ma).

Acrogenosporaceae Jayasiri & K.D. Hyde


Based on a multi-gene phylogeny, Jayasiri et al. (2018) showed that this is a distinct family
within the Minutisphaerales (S. Fryar).

Aculeata W. Dong, H. Zhang & K.D. Hyde


Dong et al. (2018) introduced this genus in Herpotrichiellaceae based on phylogenetic
analyses and distinct morphological characters (Q. Tian).

Adelolecia Hertel & Hafellner


Kistenich et al. (2018) transferred this genus from Ramalinaceae to Lecanoraceae (E.
Timdal).

Aeruginoscyphus Dougoud
Dougoud (2012) introduced this genus within Hyaloscyphaceae to accommodate Peziza
sericea (D. Haelewaters).

Agyriaceae Corda
Kraichak et al. (2018a) regarded that Miltideaceae Hafellner as a synonym of Agyriaceae
(N. Wijayawardene).

Alanomyces Roh. Sharma


Sharma et al. (2017) showed that this genus is a separate lineage in Aplosporellaceae and
introduced a new genus with a single species (A.J.L. Phillips).

Alleppeysporonites Ramanujam & K.P. Rao (fossil).


This genus shows striking similarity to the dematiaceous fungus Grallomyces F. Stevens
(Barnett 1956, Ellis 1971, Subramanian 1971) (R.K. Saxena).

Allographa Chevall.

1385
Lücking & Kalb (2018) resurrected Allographa from synonymy of Graphis (M. Kukwa).

Alternaria Nees
Currently, Alternaria has about with 589 legitimate species epithets (MycoBank 2019).
More than 150 of them have been synonymized or assigned as “indistinguishable as unique” by
Simmons (2007) since their descriptions and illustrations are scanty while the type material is
unavailable. Remaining list of species should be reduced by sixty five synonymic names due to
revisions of Woudenberg et al. (2014, 2015). Thirteen species should be transferred to Alternaria
from abolished genera Ulocladium and Nimbya (Gannibal 2018, Gannibal & Lawrence 2018).
Thus the genus Alternaria at the moment contains 366 accepted and recognizable species. Not all
of them have been subjected to molecular phylogenetic studies (P.B. Gannibal).

Amarenomyces O.E. Erikss.


Rossman et al. (2015) treated Amarenomyces as a synonym of Amarenographium. Based on
molecular analyses coupled with morphological characteristic, Wijayawardene et al. (2016)
introduced Amarenographium ammophilae Wanas.et al.; whereas, Hyde et al. (2017) introduced
Amarenomyces dactylidis Mapook et al. Multigene phylogenetic analyses revealed that these two
genera were not congeneric, thus Hyde et al. (2017) reinstated Amarenomyces (R. Phookamsak).

Amazonotheca Bat. & H. Maia


Phookamsak et al. (2016) treated the genus in Dothideomycetes, genera incertae sedis based
on herbarium study from Dr. Lima’s collection (P-4. Serra do Veado-Serra do Navio, 24 August
1961, Lima J.A. (Leg.), A.C. Batista A.C. & Xavier Filho L. (det.), URM 28927). The type
specimen was deposited in the Universidade Federal de Pernambuco (URM), Brazil but it could
not be loaned (R. Phookamsak).

Amorphothecaceae Parbery
Amorphothecaceae is a monotypic family that was previously placed in Leotiomycetes
genera incertae sedis (Baral 2016). Ekanayaka et al. (2019) retrieved it as sister to Erisyphaceae
but this placement was without support. Strong support came with the 15-gene tree of Johnston et
al. (2019), in which Amorphothecaceae was placed in the pezizelloid clade of Helotiales (D.
Haelewaters).

Anatolinites Elsik, V.S. Ediger & Bati (fossil)


Spores of Anatolinites have a tendency to break apart along the septa. Kalgutkar &
Jansonius (2000) considered Cupulisporonites Z.C. Song & Liu Cao a junior taxonomic synonym
of Anatolinites (R.K. Saxena).

Annabella Fryar, Haelew., & D.E.A. Catches.


Fryar et al. (2019) introduced this genus for A. australiensis Fryar, Haelew., & D.E.A.
Catches. from mangrove wood with perithecioid hyaline to yellowish apothecia. The genus is
confirmed as a member of Cordieritidaceae based on the molecular phylogenetic analysis of a
concatenated dataset of three ribosomal nuclear loci (D. Haelewaters).

Antennopsis R. Heim
Guswenrivo et al. (2018) used the partial 18S rRNA gene to place this genus within
Ascomycota and found that it is positioned within Sordariomycetes. It has a sister relationship to
Graphium euwallaceae (Graphiaceae), but we are reluctant in formally placing Antennopsis in this
family, because the phylogeny was based only on 18S (D. Haelewaters).

Anthopsis Fil. March. et al.

1386
Phylogenetic analyses placed this genus in Cyphellophoraceae, Chaetothyriales. The type
species, A. deltoidea, clusters within a large Cyphellophora clade and therefore Anthopsis might
be a synonym of Cyphellophora. The synonymy, however, has not yet been properly proposed.
Anthopsis currently includes three species, A. deltoidea, A. catenata and A. microspora, but DNA
sequence data proved that A. catenata is not a member of Chaetothyriales (Moussa et al. 2017).
No DNA sequence data is available for A. microspora (H. Madrid).

Aotearoamyces P.R. Johnst., J.A. Cooper & Quijada


Quijada et al. (2018a) introduced this genus for A. nothofagi, a species from fallen wood in
Nothofagaceae forests in New Zealand. It was placed in the family Tympanidaceae within the
order Phacidiales based on the phylogenetic analysis of three ribosomal nuclear loci. Based on a
larger 15-locus analysis, Johnston et al. (2019) placed Aotearoamyces within Leotiales genera
incertae sedis with high support (D. Haelewaters).

Aplosporella Speg.
Slippers et al. (2013) suggested that this genus is the sexual morph of Aplosporella, but they
declined to make a formal synonymy. Phillips et al. (2019) formally placed Bagnisiella as a
synonym of Aplosporella (A.J.L. Phillips).

Appendicisporonites R.K. Saxena & S. Khare (fossil).


Appendicisporonites appears to be related to the setose pycnidia found in some
Coelomycetes Grove (R.K. Saxena).

Aquacidia Aptroot
Aptroot et al. (2018) introduced this genus and confirmed its placement in Pilocarpaceae
(N. Wijayawardene).

Aquimonospora J. Yang & K.D. Hyde


Yang et al. (2019) introduced this monotypic genus to accommodate the new species
Aquimonospora tratensis. A new genus and species collected from freshwater habitat in Thailand
(S. Tibpromma).

Arthrocladium Papendorf
This genus comprises four species which form a monophyletic group in Trichomeriaceae.
They are associated mainly with plant material and occasionally also with opportunistic infections
in humans (Nascimento et al. 2016) (H. Madrid).

Arthrophiala (D.J. Soares, R.W. Barreto & U. Braun) W.S. Lisboa et al.
This monotypic genus was recently erected to accommodate the plant pathogenic fungus
Pseudocercospora arthrospora. DNA sequence data revealed that this fungus is a member of
Chaetothyriales (Crous et al. 2016), while true Pseudocercospora species belong in Capnodiales.
In the protologue, Arthrophiala was considered as a member of Chaetothyriaceae, but its
definitive phylogenetic placement may require further study (H. Madrid).

Ascochytites Barlinge & Paradkar (fossil)


The genus name Ascochytites was first used by Teterevnikova-Babaian & Taslakhchian
(1973), but not then validly published due to lack of illustration of the spores (R.K. Saxena).

Ascodesmisites Trivedi, Chaturv. & C.L. Verma (fossil)


The fossil specimen shows close affinity with family Pezizaceae but differs from the latter
by the absence of exciple. Minute fruiting bodies and various sexual stages somewhat resemble

1387
those of Ascodesmis Tiegh. In fossil specimen, the male and female sex organs are found growing
on two different hyphae indicating that the fungus could be dioecious (R.K. Saxena).

Asterinites Doub. & D. Pons ex Kalgutkar & Janson. (fossil)


Jansonius & Hills (1976) remarked: “Two species described; until one of these is selected as
type species, the genus is not validly published. The genus seems to be intended for mycelia,
rather than definite fruiting bodies”. The lectotype was designated by Kalgutkar & Jansonius
(2000) (R.K. Saxena).

Asterothyrites Cookson emend. Kalgutkar & Janson. (fossil)


The lectotype was selected by Jansonius & Hills (1976). Kalgutkar & Jansonius (2000)
emended the generic diagnosis. Paramicrothallites K.P. Jain & R.C. Gupta is a junior taxonomic
synonym of Asterothyrites (Kalgutkar & Jansonius 2000) (R.K. Saxena).

Astragalicola Jaklitsch & Voglmayr


Jaklitsch et al. (2018) proposed this new genus based on morphological and molecular data
(S. Fryar).

Atractium Syd. & P. Syd.


For many years, this genus was considered as a synonym of Fusarium, but a multilocus
phylogenetic study by Gräfenhan et al. (2011) proved that Atractium represents a distinct genus in
Nectriaceae (H. Madrid).

Axisporonites Kalgutkar & Janson. (fossil)


Kalgutkar and Jansonius (2000) designated Multicellaesporites indicus P. Kumar (Now:
Axisporonites indicus (P. Kumar) Kalgutkar & Janson.) as type of this monotypic genus (R.K.
Saxena).

Bacidia De Not.
Kistenich et al. (2018) treated Bacidiopsora Kalb as synonym of Bacidia (E. Timdal).

Bacillicladium Hubka, Réblová & Thureborn


This monotypic genus, typified by B. lobatum, forms a distinct lineage in Chaetothyriales
which might represent an undescribed family (Réblová et al. 2016) (H. Madrid).

Baeomycetales Lumbsch et al.


Kraichak et al. (2018a) regarded that Arctomiales S. Stenroos et al., Hymeneliales S.
Stenroos et al and Trapeliales B.P. Hodk. & Lendemer as synonyms of Baeomycetales (N.
Wijayawardene).

Bahusandhika Subram.
Liu et al. (2018a) transferred this genus to the new family, Lentimurisporaceae, based on a
multi-gene phylogeny (S. Fryar).

Barrenia E. Walsh & N. Zhang


Johnston et al. (2019) accepted this genus in Mollisiaceae (D. Haelewaters).

Barrmaeliaceae Voglmayr & Jaklitsch


Voglmayr et al. (2018) recognised the family Barrmaeliaceae with the genera Barrmaelia
and Entosordaria from phylogenetic analyses of a combined DNA data matrix containing ITS,
LSU, rpb2 and tub2 sequences of representative Xylariales including the type species of both
genera. They also established that the morphologically similar genus Clypeosphaeria belongs to

1388
Xylariaceae sensu stricto. The other genera of the family Clypeosphaeraceae aside from
Clypeosphaeria, however, still need to be studied by methods of polyphasic taxonomy to assure
their affinities to the Xylariaceae. The study by Vogmayr et al. (2018) also revealed that some
DNA sequence data of the genera Alloanthostomella, Neoanthostomella, and
Pseudoanthostomella that are available in the public domain were erroneous, which gave rise to
some serious doubt as to their familiar affinities. These genera are therefore placed in Xylariales
Incertae sedis until the phylogenetic position has been verified based on new DNA sequence data
(M. Stadler).

Basidiobolales Jacz. & P.A. Jacz.


Basidiobolales Caval.-Sm., Biological Reviews Cambridge 73: 246 (1998) is an isonym (K.
Bensch).

Basidiobolomycetes Doweld
Basidiobolomycetes Humber, Mycotaxon 120: 484 (2012) is an isonym, Doweld was
published already in 2001; see IF and MB (K. Bensch).

Basidiosporites Elsik (fossil)


Kalgutkar & Jansonius (2000) considered Amepiospora Sal.-Cheb. & Locq. a later synonym
of this genus (R.K. Saxena).

Bellicidia Kistenich, Timdal, Bendiksby & S.Ekman


Kistenich et al. (2018) introduced this genus in Ramalinaceae (E. Timdal).

Biatora Fr.
Kistenich et al. (2018) reduced Ivanpisutia S.Y. Kondr., Lőkös & Hur and Myrionora R.C.
Harris under this genus (E. Timdal).

Bibbya J.H.Willis
Kistenich et al. (2018) resurrected this genus in Ramalinaceae (E. Timdal).

Bifusisporella R.M.F. Silva et al.


Silva et al (2019) placed this new genus in the family Magnaporthaceae based on molecular
and morphological characters. This genus shown the phylogenetic inference related to
Omnidemptus but differ in by having an asexual morph with sporodochial conidiomata (Cannon &
Alcorn 1994) (S. Tibpromma).

Biporipsilonites Kalgutkar & Janson. (fossil)


Spores in this genus can be differentiated from Diporisporites Hammen by having pore
chambers. Kalgutkar & Jansonius (2000) designated Diporicellaesporites belluloides Z.C. Song
(Now: Biporipsilonites belluloides (Z.C. Song) Kalgutkar & Janson.) as type of this genus (R.K.
Saxena).

Botryosphaeriaceae Theiss. & Syd.


See under Endomelanconiopsis E.I. Rojas & Samuels Tao (A.J.L. Phillips).

Botryozyma Shann & M.T. Sm. emend. Lachance & Kurtzman


Kurtzman & Boekhout (2017) include Ascobotryozyma J. Kerrigan, M.T. Sm. & J.D. Rogers
(under Botryozyma) to comply with the Melbourne Code (W.P. Pfliegler & E. Horváth).

Boubovia Svrček

1389
This genus and Coprotus Korf & Kimbr. represents an independent clade of Pezizales, not
included in Ascodesmidaceae, as shown by Perry et al. (2007) and also Hansen et al. (2013) (P.
Alvarado).

Brachysporisporites R.T. Lange & P.H. Sm. (fossil)


Granatisporites Elsik & Janson. is a junior taxonomic synonym of Brachysporisporites
(Kalgutkar & Jansonius 2000). Spores of Brachysporisporites are usually compared to the conidia
of the modern Brachysporium Sacc. (R.K. Saxena).

Bradymyces Hubka et al.


This genus currently includes with three species which form a monophyletic group within
Trichomeriaceae (Réblová et al. 2016) (H. Madrid).

Brefeldiellites Dilcher (fossil)


The fossil form of this monotypic genus is similar to the Brefeldiella Speg. but cannot be
placed in this extant genus because the spores are not known (R.K. Saxena).

Bresadolia Speg.
Motato-Vásquez et al. (2018) reinstated this genus and accepted as in Polyporaceae (V.
Papp).

Brigantiaeaceae Hafellner & Bellem.


Kraichak et al. (2018a) regarded Letrouitiaceae as a synonym of Brigantiaeaceae (N.
Wijayawardene).

Brunneofusispora S.K. Huang & K.D. Hyde


Brunneofusispora was introduced in Phookamsak et al. (2019) to accommodate massarina-
like species and species identified as Massarina rubi (Fuckel) Sacc. Multi-gene phylogenetic
analyses revealed that the genus formed a distinct clade within Occultibambusaceae (Phookamsak
et al. 2019) (R. Phookamsak).

Brunneomurispora Phook. et al.


Phookamsak et al. (2019) introduced a monotypic genus Brunneomurispora to
accommodate B. lonicerae. Multi-gene phylogenetic analyses showed that the genus formed a
distinct clade basal to Neosetophoma in Phaeosphaeriaceae. However, the sexual morph of
Neosetophoma has phragmosporous ascospores; whereas, Brunneomurispora has dictyosporous
ascospores. Asexual morph of Brunneomurispora is undetermined (R. Phookamsak).

Bryobilimbia Fryday et al.


Fryday et al. (2014) introduced Bryobilimbia to accommodate six species, including Lecidea
hypnorum and some related taxa based on morphological characters such as bacilliform conidia,
slightly swollen paraphyses and thallus minutely squamulose without marginal lobes.
Phylogenetic analyses revealed Bryobilimbia as a distinct monophyletic lineage within
Lecideaceae, Lecideales, Lecanoromycetes (P. Rodriguez-Flakus).

Bulbomicroidium Marm. et al.


Marmolejo et al. (2018) introduced this monotypic genus and confirmed its placement in
Erysiphaceae based on molecular phylogenetic analysis of the LSU (S. Takamatsu).

Byssonectria P. Karst.
The genus Kotlabaea Svrček has been reduced under Byssonectria by Lindemann et al.
(2015) (P. Alvarado).

1390
Callimothallus Dilcher (fossil)
Callimothallus lacks any central dehiscence and is characterized by numerous pores. Elsik
(1978) pointed out that the porate condition in Callimothallus is required for at least in a number
of the cells to separate it from Phragmothyrites W.N. Edwards and that if the porate nature is well
represented, even fragments of the fructification are recognizable. Kalgutkar & Jansonius (2000)
considered Pseudosphaerialites Venkatach. & R.K. Kar and Siwalikiathyrites R.K. Saxena & H.P.
Singh as junior synonyms of Callimothallus (R.K. Saxena).

Calongea Healy, Bonito & Trappe


This genus was erected to accommodate Pachyphloeus prieguensis (Healy et al. 2009),
reflecting a separate lineage within Pezizaceae (I. Kušan, N. Matočec & P. Alvarado).

Calyptosphaeria Réblová & A.N. Mill.


Réblová et al. (2018) introduced this genus to accommodate two new species and two new
combinations based on a multi-gene phylogeny and morphological characters (S. Fryar).

Camptophora Réblová & Unter.


This genus, originally proposed to accommodate a ‘Cyphellophora’ species clustering
outside the Cyphellophoraceae clade. It currently includes two species, viz. C. hylomeconis, the
generic type and C. schimae (Réblová et al. 2013, Yang et al. 2018a) (H. Madrid).

Candida Berkhout
Transfer of several Candida species to various yeast genera (including novel genera) is
expected. But Candida is expected to be the retained genus name for the C. tropicalis clade
instead of Lodderomyces Van der Walt in Debaryomycetaceae to comply with the Melbourne
Code. Transferring the hundreds of Candida species to monophyletic genera is one of the main
current challenges in yeast taxonomy (Kurtzman & Boekhout 2017) (W.P. Pfliegler & E. Horváth).

Castanedospora G. Delgado & A.N. Mill.


Delgado et al. (2018) introduced this hyphomycetous genus to accommodate Sporidesmium
pachyanthicola R.F. Castañeda & W.B. Kendr. In the phylogenetic analyses, it grouped in
Extremaceae (Capnodiales, Dothideomycetes) distant from Sporidesmiaceae sensu stricto in
Sordariomycetes (G. Delgado & S. Fryar).

Catinaria Vain.
Kistenich et al. (2018) transferred this genus from Ramalinaceae to the Lecanorales genera
incertae sedis (E. Timdal).

Cenangiaceae Rehm
In their 15-gene phylogeny, Johnston et al. (2019) found high support for a sclerotinioid
clade including Cenangiaceae within the Helotiales. Here, we follow the emended concept of
Pärtel et al. (2017) for the family Cenangiaceae, including the following genera: Cenangiopsis,
Cenangium, Chlorencoelia, Crumenulopsis, Encoelia, Fabrella, Heyderia, Rhabdocline,
Sarcotrochila, Trochila, and Velutarina (D. Haelewaters).

Cephaliophora Thaxt.
This genus is firmly nested as a separate lineage within Ascodesmidaceae, Pezizales (Kušan
et al. 2018) (I. Kušan & N. Matočec).

Cercosporites E.S. Salmon (fossil)

1391
Aspergillites Trivedi & C.L. Verma ex Janson., Hills & Hartk.-Fröd. is a taxonomic latter
synonym of Cercosporites (Kalgutkar & Jansonius 2000) (R.K. Saxena).

Chaetosphaerites Félix (fossil)


This genus is characterized by spores having two middle cells being dark brown and two
end cells pale brown. The shape of the sporidia is strongly obtuse spindle-shaped, almost like that
of a cylinder with rounded ends. Cannanorosporonites Ramanujam & K.P. Rao is a latter
taxonomic synonym of Chaetosphaerites (R.K. Saxena).

Cheirospora Moug. & Fr.


Johnston et al (2019) accepted this genus in Mollisiaceae (D. Haelewaters).

Chlorociboriaceae Baral & P.R. Johnst.


Based on Johnston et al.’s (2019) 15-gene phylogenetic analysis, Chlorobiboriaceae is part
of the sclerotinioid clade within Helotiales (D. Haelewaters & N. Wijayawardene).

Chroogomphus (Singer) O.K. Mill.


Scambler et al. (2018) revised the infrageneric classification of the genus, introduced three
subgenera (Chroogomphus, Floccigomphus and Siccigomphus) and five sections/clades within
subg. Chroogomphus (Chroogomphus, Confusi, Filiformis, Fulminei and the informal Vinicolores
clade). Chroogomphus subfulmineus from Cyprus, Finland and the UK, and C. pakistanicus and
C. pruinosus from Pakistan were recently described bringing the number of recognized species to
25 (Scambler et al. 2018, Kiran et al. 2020) (D. Haelewaters).

Chrysodiscaceae Baral & Haelew.


Baral & Polhorský (2019) introduced this family to accommodate Chrysodisca peziculoides,
a broadly distributed European discomycete resembling Pezicula but phylogenetically separated
from previously recognized families within Helotiales (D. Haelewaters).

Chrysonectria Lechat & J. Fourn


Lechat et al (2018a) introduced this genus and showed that it is phylogenetically belongs to
Bionectriaceae (N. Wijayawardene)

Cinereomycetella Zmitr.
Justo et al. (2017) showed that Diplomitoporus overholtsii (Pilát) Gilb. & Ryvarden forms a
distinct phylogenetic lineage in the family Polyporaceae. Hence, Zmitrovich (2018) introduced
the monotypic genus Cinereomycetella in Gelatoporiaceae to accommodate Diplomitoporus
overholtsii (V. Papp).

Cladoniaceae Zenker
Kraichak et al. (2018a) revised Lecanoromycetes and showed that Squamarinaceae
Hafellner, Stereocaulaceae Chevall are synonyms of Cladoniaceae Zenker (in Lecanorales) based
on temporal-based classification (N. Wijayawardene).

Cladophialophora Borelli
The genus Cladophialophora is polyphyletic within Chaetothyriales. The generic type, C.
carrionii and most species of this genus belong in Herpotrichiellaceae, but a few species are
related to other chaetothyrialean families, such as Epibryaceae and Trichomeriaceae (Madrid et al.
2016) (H. Madrid).

Cladorrhinum Sacc. & Marchal

1392
Phylogenetic studies by Cai et al. (2006), Madrid et al. (2011) and Carmarán et al. (2015)
proved that this genus belongs in Lasiosphaeriaceae. Nevertheless, Hyde et al. (2020) accepted
this genus in Podosporaceae. The genus, however, is polyphyletic within this fungal family (H.
Madrid).

Cladosporites Félix (fossil)


The conidia greatly resemble those of the genera Cephalothecium Corda and Cladosporium
Link. (R.K. Saxena).

Clarireedia L.A. Beirn


Salgado-Salazar et al. (2018) proposed this new genus with C. homoeocarpa as the type
species (Rutstroemiaceae) based on molecular and morphological characters. The genus
comprises three other species except the type species viz. C. bennettii, C. jacksonii, and C.
monteithiana (Salgado-Salazar et al. 2018) (S. Fryar & S. Somrithipol).

Claussenomyces Kirschst.
According to Quijada et al (2018b) and Species Fungorum (2020), the genus
Claussenomyces currently contains 16 species. However, the genus is “heterogeneous” (Jaklitsch
et al. 2016a) and polyphyletic. As a result, it is currently under taxonomic revision.
“Claussenomyces” prasinulus is treated as Leotiales genera incertae sedis based on its placement
in Johnston et al.’s (2019) 15-gene tree. (D. Haelewaters, I. Kušan & N. Matočec).

Cochlearomyces Crous
Crous et al. (2017) established this genus to accommodate a synnematous fungus on leaf
litter of Eucalyptus. The same authors also introduced a new family (Cochlearomycetaceae) for
the genera Cochlearomyces and Satchmopsis, which was retrieved in Leotiales based on the
phylogenies of both Ekanayaka et al. (2019) and Johnston et al. (2019) (D. Haelewaters).

Coleophoma Hohn.
Using a polyphasic approach, Crous & Groenewald (2016) established the teleomorph-
anamorph connection between Coleophoma and Parafabraea Chen Chen, Verkley & Crous, thus
reducing Parafabraea to synonymy. Coleophoma is placed in Dermateaceae (Johnston et al. 2019)
(D. Haelewaters).

Colligerites K.P. Jain & R.K. Kar (fossil).


Involutisporonites Elsik broadly resembles Colligerites but in the former, coiling is not
perfect and it has a hyaline cell at the tip (R.K. Saxena).

Compsocladium I.M. Lamb


Kistenich et al. (2018) transferred this genus from Ramalinaceae to the Lecanorales
incertae sedis (E. Timdal).

Coprotus Korf & Kimbr.


See under Boubovia Svrček (P. Alvarado).

Coprinites Poinar & Singer (fossil)


The fossil mushroom has affinity with the present day genus Coprinus Pers (R.K. Saxena).

Corticifraga D. Hawksw. & R. Sant.


The genus Corticifraga, comprising 7 species and has been recently transferred to
Gomphilaceae (Pino-Bodas et al. 2017, Suija et aĺ. 2018), but so far only the type species has been
sequenced (M. Kukwa & A. Suija).

1393
Craspedodidimella F.R. Barbosa, R.F. Castañeda & Gusmão
Barbosa et al. (2017) introduced this genus and showed that it belongs in Ascomycota genera
incertae sedis (F.R. Barbosa).

Crassiclypeus A. Hashim. et al.


Hashimoto et al. (2018) proposed this genus with Crassiclypeus aquaticus as the type
species, which was previously included in the Lophiostoma bipolare species complex (S. Fryar).

Crepatura C.L. Zhao


Zhao et al. (2019) proposed this new genus with Crepatura ellipsospora as the type
species based on molecular and morphological characters. While, this genus phylogenetically
closely related to Pirex concentricus but morphology is different (S. Tibpromma).

Crustospathula Aptroot
Kistenich et al. (2018) transferred this genus from the Ramalinaceae to the Malmideaceae
(E. Timdal).

Cryptodidymosphaerites Currah, Stockey & B.A. LePage (fossil)


This genus from the Princeton chert presents sufficient characters such as the presence of a
pseudothecium, ascospores morphology, orientation and the mycoparasitic habit to place it close
to Didymosphaeria Fuckel. (Aptroot 1995) (R.K. Saxena).

Cryptodiscus Corda
Pino-Bodas et al. (2017) regarded that Lettauia D. Hawksw. & R. Sant. as a synonym of
Cryptodiscus, however, 3 species are still orphaned under that name, but so far have not been
relocated (M. Kukwa & A. Suija).

Cryptophyllachora L. Kiss, Kovács & R.G. Shivas


Kiss et al. (2018) proposed Cryptophyllachora to accommodate C. eurasiatica (the type
species) collected from common ragweed (Ambrosia artemisiifolia) in Hungary, and C. ambrosiae
that was transferred from Phyllachora ambrosiae (S. Somrithipol).

Cucitella Jaklitsch & Voglmayr


Jaklitsch et al. (2018) proposed this new genus based on morphological and molecular data
(S. Fryar).

Cucurbitariaceites R.K. Kar, R.Y. Singh & S.C.D. Sah (fossil)


Cucurbitariaceites is easily distinguishable from all the fossil genera of Microthyriales G.
Arnaud. by its circular to subcircular shape, darker outer layer and thin inner layer, in the absence
of true paraphyses and the presence of cylindrical asci. Cucurbitariaceites closely resembles the
extant family Cucurbitariaceae G. Winter in all the characters and Cucurbitaria Gray is the
widely known genus of this family (Bessey 1950) (R.K. Saxena).

Culbersonia Essl.
This genus is considered part of the Caliciaceae by Aptroot et al. (2019) (P. Alvarado).

Cylindriaceae Crous & L. Lombard


Crous et al. (2018) introduced this new family to accommodate Cylindrium, which was
previously in incertae sedis. Hyde et al. (2020) accepted this family in Amphisphaeriales (S.
Fryar).

1394
Cylindrosporium Grev.
Baral (2016) and Ekanayaka et al. (2019) accepted this genus in Ploettnerulaceae (D.
Haelewaters & N. Wijayawardene).

Cyphellophora G.A. de Vries


Species of Cyphellophora are characterized by slow-growing colonies, dematiaceous
mycelium and phialidic conidiogenous cells usually with conspicuous collarettes.
Phylogenetically, they form a distinct clade in Chaetothyriales for which the family
Cyphellophoraceae was erected (Réblová et al. 2013) (H. Madrid).

Deccanodia Singhai (fossil)


Deccanodia resembles Diplodia Fr. in its pycnidium and 2-celled, brown and mostly
ellipsoid conidia (Barnett 1960, Gilman 1959). However, in Diplodia, black pycnidia and equal 2-
celled conidia are present thus showing a distinct difference from the fossil fungus which has
slightly brown pycnidium and unequally divided 2-celled conidia. Deccanodia also resembles
Apiocarpella Syd., belonging to Fungi Imperfecti (Barnett 1960) (R.K. Saxena).

Dekkera Van der Walt


Expected transfer of species to Brettanomyces Kufferath & van Laer to comply with the
Melbourne Code (Kurtzman & Boekhout 2017) (W.P. Pfliegler and E. Horváth).

Dendrostoma X.L. Fan & C.M. Tian


Fan et al. (2018a) placed this new genus in the family Erythrogloeaceae with three species
viz. D. mali (the type species), D. quercinum, and D. osmanthi. The genus comprises 19 species
associated wood canker disease (Jaklitsch et al. 2019, Jiang et al. 2019b, Zhu et al. 2019). (S.
Fryar, X. Fan & S. Somrithipol).

Densocarpa Gilkey
According to recently shown phylogenies (e.g. Kumar et al. 2017), this genus should not be
considered as a synonymy of Stephensia as it represents separate lineage in Geopyxis clade (I.
Kušan & N. Matočec).

Densospora McGee
Single-locus and multigene phylogenetic reconstructions depict Densosporaceae as a distant
sister clade of Endogonaceae within the order Endogonales (Desirò et al. 2017) (K. Bensch).

Dentocorticium (Parmasto) M.J. Larsen & Gilb.


Liu et al. (2018b) treated Dendrodontia Hjortstam & Ryvarden and Fuscocerrena Ryvarden
as synonyms of Dentocorticium (V. Papp).

Dextrinoporus H.S. Yuan


Yuan & Qin (2018) introduced this monotypic genus to accommodate the new species D.
aquaticus H.S. Yuan and showed that it forms a distinct phylogenetic lineage in the family
Polyporaceae (V. Papp).

Diademosa Shoemaker & C.E. Babc.


This genus has been originally considered as a genus of Diademaceae (Shoemaker &
Babcock 1992). Later it was transferred to Pleosporaceae (Ariyawansa et al. 2014) but molecular
phylogeny has not still used for its replacement (P.B. Gannibal).

Diaporthosporellaceae C.M. Tian & Q. Yang

1395
Yang et al. (2018b) introduced Diaporthosporellaceae for Diaporthosporella cercidicola, a
new genus and species collected from diseased branches of Cercis chinensis in China (S.
Somrithipol & S.S.N. Maharachchikumbura).

Diaporthostoma X.L. Fan & C.M. Tian


Diaporthostoma machili is the type species of recently introduced genus Diaporthostoma by
Fan et al. (2018a) (S.S.N. Maharachchikumbura).

Diaporthostomataceae X.L. Fan & C.M. Tian


This monotypic family was introduced by Fan et al. (2018a) based on morphology and the
analysis of partial ITS, LSU, rpb2 and tef1-α gene sequences. This family belongs to the order
Diaporthales (S.S.N. Maharachchikumbura and S. Fryar).

Dictyoceratosporella Y.R. Ma & X.G. Zhang


Ma et al. (2016) established this hyphomycetous genus. Sequence data are lacking thus
taxonomic placement is uncertain (J. Ma).

Dictyosporites Félix (fossil)


These spores are comparable to the conidia of some modern genera like Dictyosporium
Corda, Stemphylium Wallr., Septosporium Corda and Alternaria Nees, and the ascospores of
Pleospora Rabenh. ex Ces. & De Not. Arbusculites Paradkar, Dactylosporites Paradkar,
Pleosporonites R.T. Lange & P.H. Sm. and Ravenelites Ramanujam & Ramachar are latter
taxonomic synonyms of Dictyosporites (Kalgutkar & Jansonius 2000) (R.K. Saxena).

Digitiseta Gordillo & Decock


Gordillo & Decock (2018) proposed Digitiseta and transferred two Myrothecium (M.
setiramosum and M. dimorphum) into the genus as D. setiramosa (the type species) and D.
dimorpha. Digitiseta parvidigitata and D. multidigitata were described as new and also included.
Digitiseta differs from the two closely related genera, Inaequalispora and Parvothecium, in
having short apical branches of the setoid extension (S. Somrithipol).

Diphymyces I.I. Tav.


Diphymyces was introduced by Tavares (1985) to accommodate Laboulbeniales species
with thalli that have a septum vertically separating cells II and VI, four tiers of perithecial wall
cells, and (sub-) apical perithecial outgrowths. The separation of this genus from Corethromyces
Thaxt. was primarily based on the position of cells II and VI, which is a variable character
(discussed in De Kesel & Haelewaters 2019). Also the genera Asaphomyces Thaxt. and
Euphoriomyces Thaxt. might be synonymous. These four genera are placed in three different
subtribes (Tavares 1985), but several higher taxa (subtribes and tribes) from the Tavares (1985)
classification are polyphyletic following molecular phylogenetic treatments (Goldmann & Weir
2018, Haelewaters et al. 2018b). Molecular phylogeny is needed to resolve these taxonomic and
systematic problems (D. Haelewaters).

Diplodites D.N. Babajan & Tasl. ex Kalgutkar, Nambudiri & Tidwell (fossil)
Kalgutkar et al. (1993) validated the name Diplodites to encompass fossil taxa that are
morphologically similar to the extant fungi Diplodia Fr., Botryodiplodia (Sacc.) Sacc. and other
related genera such as Dothiorella Sacc. and Macrophoma (Sacc.) Berl. & Voglino.
Palaeodiplodites Kyoto Watanabe, H. Nishida & Tak. Kobay. is a junior taxonomic synonym of
Diplodites (Kalgutkar & Jansonius 2000). (R.K. Saxena).

Diploneurospora K.P. Jain & R.C. Gupta (fossil)

1396
This monotypic genus accommodates two-celled (cells unequal), uniseriate, elliptical
ascospores with uneven margin; upper cell prominent, dark brown, thick-walled, wall sculptured
with longitudinal ribs; lower cell hyaline, appendage-like, small in size, rib sculpture faint. It
closely resembles to the single celled ascospores of extant genus Neurospora Shear & B.O.
Dodge (R.K. Saxena).

Diporisporites Hammen (fossil)


Kalgutkar & Jansonius (2000) considered Scabradiporites Y.K. Mathur a latter taxonomic
synonym of Diporisporites (R.K. Saxena).

Discinellaceae Ekanayaka & K.D. Hyde


Ekanayaka et al. (2019) introduced this family within Helotiales. Johnston et al. (2019)
found high support for what they referred to as the Discinella-Pezoloma lineage (sensu Baral
2016), placed as sister clade to Gelatinodiscaceae in the discinelloid clade of Helotiales. A
number of genera that were previously placed elsewhere are now considered members of
Discinellaceae: Cladochasiella, Fontanospora, Margaritispora, Naevala, Pseudopezicula and
Tetrachaetum (D. Haelewaters).

Dothiorella Sacc.
Yang et al. (2017) considered that Spencermartinsia should no longer be considered a
separate genus and placed it as a synonym of Dothiorella. Currently 389 names are listed in
MycoBank, but only 38 are known from culture (A.J.L. Phillips).

Drepanopezizaceae Baral
Introduced in Johnston et al. (2019) as sister family to Ploettnerulaceae within Helotiales
(D. Haelewaters).

Ducatina Ertz & Søchting


Ertz et al. (2017b) introduced this genus which belongs in Trapeliaceae (N.
Wijayawardene).

Durella Tul. & C. Tul.


Baral (2016) treated this genus in his “Strossmayeria lineage”, which was confirmed by the
ITS tree of Johnston et al. (2019). However, two species, “Durella” macrospora and “D.”
melanochlora, are phylogenetically distinct from the type (D. connivens) and are here placed
under Helotiales genera incertae sedis (D. Haelewaters).

Durotheca Læssøe et al.


De Long et al. (2019) described two new species of Durotheca from China and provided a
new molecular phylogeny that proved the affinities of the genus to the Hypoxylaceae. This had
already been indicated by morphological data, such as the presence of a nodulisporium-like
asexual morph in some species (M. Stadler).

Dyadosporites Hammen ex R.T. Clarke (fossil)


Jansonius & Hills (1976) remarked that “although van der Hammen (1954) gave a diagnosis
and the name of the type species, the latter was never described (or illustrated)”. Therefore,
Dyadosporites was not a validly published name of a taxon, but as merely proposed in
anticipation of future acceptance of the group. Clarke (1965) was the first to have validly
published this genus name and also the first to assign a species (Dyadosporites ellipsus R.T.
Clarke) to it. Dyadosporonites Elsik and Psidimobipiospora Sal.-Cheb. & Locq. are latter
taxonomic synonyms of Dyadosporites (Kalgutkar & Jansonius 2000) (R.K. Saxena).

1397
Efibulella Zmitr.
Justo et al. (2017) showed that Phlebia deflectens (P. Karst.) Ryvarden forms a distinct
phylogenetic lineage in the family Phanerochaetaceae. Hence, Zmitrovich (2018) introduced the
monotypic genus Efibulella to accommodate Phlebia deflectens (V. Papp).

Elaphroporia Z.Q. Wu & C.L. Zhao


Wu et al. (2018) introduced this monotypic genus to accommodate the new species E.
ailaoshanensis Z.Q. Wu & C.L. Zhao, and accepted it as a genus in the residual polyporoid clade
based on phylogenetic analyses (V. Papp).

Emergomyces Dukik et al.


This genus of thermally dimorphic clinical fungi belongs in the family Ajellomycetaceae,
according to multilocus phylogenetic studies by Dukik et al. (2017) (H. Madrid).

Endomelanconiopsis E.I. Rojas & Samuels Tao


Yang et al. (2017) considered that this genus warrants a separate family and thus introduced
Endomelanconiopsisaceae to accommodate it. Phillips et al. (2019) took into account phylogeny
(ITS, LSU), morphology and evolutionary divergence times and concluded that
Endomelanconiopsis resides within Botryosphaeriaceae. Hence, Phillips et al. (2019) regarded
Endomelanconiopsisaceae as a synonym of Botryosphaeriaceae (A.J.L. Phillips).

Endophoma Tsuneda & M.L Davey


This is a monotypic genus in Didymellaceae, typified by E. elongata. The fungus is an
atypical coelomycete with endogenous conidiogenesis and its taxonomic position has been
elucidated by DNA sequence analyses (Tsuneda et al. 2011a) (H. Madrid).

Enterodictyon Müll. Arg.


It is an autonomous genus in Wijayawardene et al. (2017a), but the type species, E. indicum
Müll. Arg., was transferred to Diorygma (Joseph et al. 2018). However, some species still need to
be studied and relocated (M. Kukwa).

Entrophospora Ames & Schneider


The fungus was treated as an incertae sedis in the last classification (Wijayawardene et al.
2018). Nevertheless, all partial rDNA sequences published within the last years, suggest that their
type fungus (E. infrequens) belongs to Claroideoglomus clade (Oehl et al 2011e,f), justifying the
use of Entrophosporaceae instead of Claroideoglomeraceae.

Epicladonia D. Hawksw.
According to Pino-Bodas et al. (2017), the generic type, E. sandstedei (Zopf) D. Hawksw.
belongs to Leotiomycetes while other species are placed in Ostropales (A. Suija).
Epithamnolia Zhurb.
Suija et al. (2017) showed that lichenicolous species of Hainesia form a distinct
phylogenetic lineage within Phacidiales, and provisionally transferred lichenicolous species to the
morphologically similar genus Epithamnolia. According to molecular analysis by Quijada et al.
(2018a), the genus was placed in the poorly known Mniaecia lineage, which is now family
Mniaeciaceae within Leotiales (Johnston et al. 2019). In contrast, Ekanayaka et al. (2019)
regarded this genus as a member of the Epicladonia-Epithamnolia clade in Lichinodiales,
however this placement had no statistical support (D. Haelewaters, A. Suija & N. Wijayawardene).

Eudarluca Speg.
Rossman et al. (2015) treated Eudarluca as a synonym of Sphaerellopsis based on a study of
Trakunyingcharoen et al. (2014). The congeneric status of Eudarluca and Sphaerellopsis was

1398
clarified based on Eudarluca caricis and Sphaerellopsis filum. However, Phookamsak et al.
(2014) examined the isotype of Eudarluca australis and compared the morphology with E.
caricis. Based on morphological examination, Phookamsak et al. (2014, 2019) mentioned that
these two genera were not congeneric and suggested to instate Eudarluca in Phaeosphaeriaceae
for pending further studies (R. Phookamsak).

Eutiarosporella Crous
Based on ITS and LSU sequence phylogeny, Crous et al. (2015a) introduced
Eutiarosporella as a new genus for tiarosporella-like fungi with long-necked conidiomata and
holoblastic conidiogenesis (A.J.L. Phillips).

Exesisporites Elsik (fossil)


The centrally located pore in Exesisporites is generally surrounded by a dark circular patch
which is interpreted as a thickened wall. Ethridge Glass et al. (1986) cited possible affinity of
Exesisporites to the extant fungus Nigrospora Zimm. (R.K. Saxena).

Exochalara W. Gams & Hol.-Jech.


Crous et al (2018b) treat this genus as a member in Neolauriomycetaceae (K. Bensch).

Exophiala J.W. Carmich.


This is a species-rich genus of black yeast-like fungi, polyphyletic within Chaetothyriales.
The type species, E. salmonis, and most members of this genus belong in Herpotrichiellaceae, but
some species are related to other chaetothyrialean families, such as Trichomeriaceae (Madrid et
al. 2016) (H. Madrid).

Extremaceae Quaedvl. & Crous


Paradevriesiaceae was introduced by Crous et al. (2019) and comprises Paradevriesia
compacta (CBS 118294), P. Americana (CBS 117726), and P. pseudoamericana (CPC 16174).
However, Crous et al. (2019) did not include sequence of Extremaceae in their phylogenetic tree.
In Hongsanan et al. (in prep), Paradevriesia strains form a distinct lineages within Extremaceae.
Hence, Hongsanan et al. (in prep.) regarded Paradevriesiaceae as a synonym of Extremaceae (S.
Hongsanan & N. Wijayawardene).

Flabellascoma A. Hashim. et al.


Hashimoto et al. (2018) proposed this genus to accommodate two species which were
previously in the Lophiostoma bipolare species complex (S. Fryar).

Flagellospora Ingold
Ekanayaka et al. (2019) retrieved this genus as a distinct phylogenetic lineage without
support. We propose it here as Leotiales genera incertae sedis based on the placement of F.
curvula, the type species, in the 15-gene tree of Johnston et al. (2019) (D. Haelewaters).

Flavignomonia N. Jiang et al.


Jiang et al. (2019c) proposed this new genus based on morphological and molecular data
with synnemata similar to Synnemasporella but differ in orange synnematal tips and hyaline
conidia (S. Tibpromma).

Fomitopsidaceae Jülich
Zmitrovich (2018) treated Adustoporiaceae Audet, Amyloporiaceae Audet, Fibroporiaceae
Audet, Lentoporiaceae Audet, Pycnoporellaceae Audet, Rhodoniaceae Audet and
Sarcoporiaceae Audet as synonyms of Fomitopsidaceae (V. Papp).

1399
Fonsecaea Negroni
This genus currently includes eight species, most of which are associated with infections in
humans. Phylogenetically, Fonsecaea belongs in Herpotrichiellaceae (Arzanlou et al. 2007,
Madrid et al. 2016, Dong et al. 2018) (H. Madrid).

Foveodiporites C.P. Varma & Rawat emend. Kalgutkar & Janson. (fossil)
Punctodiporites C.P. Varma & Rawat is a later synonym of Foveodiporites as emended by
Kalgutkar & Jansonius (2000) (R.K. Saxena).

Frasnacritetrus Taug. emend. R.K. Saxena & S. Sarkar (fossil)


Taugourdeau (1968) originally described this genus under Acritarcha incertae sedis from
the Late Devonian (Frasnian) sediments of France. Kendrick & Carmichael (1973) published a
list of staurosporous genera and their illustrations which strongly suggests that Frasnacritetrus is
a fossil representative of Tetraploa Berk. & Broome, hence its placement under Acritarcha
incertae sedis by Taugourdeau (1968) does not seem justified. All the seven assemblages studied
by Saxena & Sarkar (1986), wherefrom the present microfossils have been recovered, contain
poaceous pollen as well. Since Tetraploa mainly grows on Poaceae, the association of
Frasnacritetrus with poaceous pollen is considered a supporting evidence for the affinity of
Frasnacritetrus with Tetraploa (R.K. Saxena).

Frutidella Lalb
Kistenich et al. (2018) transferred this genus from the Ramalinaceae to the Lecanoraceae
(E. Timdal).

Fuscosclera Hern.-Restr. et al.


Johnston et al (2019) accepted this genus in Mollisiaceae (D. Haelewaters).

Fusichalara S. Hughes & Nag Raj


Fusichalara minuta clustered in the family Sclerococcaceae (Réblová et al. 2016, Yu et al.
2018), but not type species (H. Zhang).

Fusiformisporites Rouse (fossil)


Striadyadosporites Dueñas is a later taxonomic synonym of Fusiformisporites (R.K.
Saxena).

Galbinothrix Frisch et al.


This genus was recently introduced from Japan and Korea by Frish et al. (2018). Multi-gene
phylogenetic analyses revealed that the monotypic genus formed a distinct clade together with
Chrysothrix, Melarthonis, and Arthonia (P. Rodriguez-Flakus).
Geesterania Westphalen, Tomšovský & Rajchenberg
Westphalen et al. (2018) introduced this genus in Meruliaceae to accommodate Junghuhnia
carneola (Bres.) Rajchenb. and Geesterania davidii Westphalen & Rajchenberg. (V. Papp).

Gelatinoamylaria Prasher & R. Sharma


Prasher et al. (2016) introduced Gelatinoamylaria as a new genus to accommodate a species
with gelatinous apothecia and amyloid ascospores. It is tentatively placed in Dermateaceae but no
sequences are available. It should be analyzed phylogenetically to determine its true systematic
position (I. Kušan & N. Matočec).

Gelatinosporium Peck
This genus belongs to Tympanidaceae following the family concept of Baral (2016) and
Quijada et al. (2020) (N. Wijayawardene).

1400
Geonectria Lechat & J. Fourn.
This is newly described genus (Lechat et al. 2018b) that belongs to Bionectriaceae (I. Kušan
& N. Matočec).

Gloeoporellus Zmitr.
Justo et al. (2017) showed that Tyromyces merulinus (Berk.) G. Cunn. forms a distinct
phylogenetic lineage in the family Incrustoporiaceae. Hence, Zmitrovich (2018) introduced the
monotypic genus Gloeoporellus to accommodate Tyromyces merulinus (V. Papp).

Gloniales Jayasiri & K.D. Hyde


Jayasiri et al. (2018) introduced this order as a distinct sister clade to the Mytinidiales (S.
Fryar).

Grovesiella M. Morelet
Previously placed in Tympanidaceae. However, G. abieticola, the type species of the genus,
was retrieved as a member of Godroniaceae in the ITS tree of Johnston et al. (2019) (D.
Haelewaters).

Gymnostellatospora Udag. et al.


Previously regarded as a member of Myxotrichaceae (Baral 2016), this genus is now placed
in Pseudeurotiaceae, Thelebolales based on the ITS phylogeny of Johnston et al. (2019). This
placement agrees with the suggestion of Minnis & Lindner (2013) to consider the genera
Geomyces, Gymnostellatospora, Leuconeurospora, Pseudeurotium and Pseudogymnoascus as
members of the family Pseudeurotiaceae (D. Haelewaters).

Hanseniaspora Zikes
This teleomorph genus has a priority over its anamorphic counterpart, Kloeckera. Kloeckera
species have recently been transferred here (Kurtzman & Boekhout 2017, Čadež et al. 2019) (W.P.
Pfliegler & E. Horváth).

Harmoniella V.N. Boriss.


This enigmatic genus currently includes two species, the generic type H. chrysocephala, and
H. campanaensis. They have been collected on plant material in Ukraine and Chile, respectively.
Both are apparently non-culturable fungi and so far it has been impossible to generate DNA
sequence data of them. No sexual morph is known for Harmoniella (H. Madrid).

Hemigraphaceae D.Q. Dai & K.D. Hyde


Dai et al. (2018) introduced this family in Asterinales to accommodate Hemigrapha (Müll.
Arg.) D. Hawksw. (N. Wijayawardene).

Hemiphacidiaceae Korf
Ekanayaka et al. (2019) reinstated this family but careful analysis of previous work shows
that this decision is flawed. Pärtel et al. (2017) included in their phylogenetic analysis the type
species of the genus Cenangium, C. ferruginosum, to find that it is positioned in the clade that
Ekanayaka et al. (2019) refer to as Hemiphacidiaceae but truly is Cenangiaceae. The clade named
“Cenangiaceae” by Ekanayaka et al. (2019) is regarded as Helotiales genera incertae sedis by
other studies, including Zhao et al. (2016) Pärtel et al. (2017), Johnston et al. (2019). This clade
includes “Cenangium” acuum, Piceomphale bulgarioides, and P. pinicola (basionym
Moellerodiscus pinicola). In this outline, we follow Pärtel et al.’s (2017) emended concept of
Cenangiaceae including members of the previous Hemiphacidiaceae (see Cenangiaceae) (D.
Haelewaters).

1401
Hemipholiota (Singer) Bon
Bon (1986) elevated Pholiota subgen. Hemipholiota to genus level and typified by
Hemipholiota populnea (Singer) Bon as the type species. Molecular data analyses confirmed that
Hemipholiota is a separate genus from Pholiota with a unique and uncertain phylogenetic
position, distinct from Strophariaceae (Moncalvo et al. 2002, Gulden et al. 2005) (B. Dima).

Henssenia Ertz, R.S. Poulsen & Søchting


Ertz et al. (2017a) introduced this genus and showed that it belongs in Koerberiaceae (N.
Wijayawardene).

Hermanssonia Zmitr.
Justo et al. (2017) showed that Phlebia centrifuga P. Karst. forms a distinct phylogenetic
lineage in the family Meruliaceae. Hence, Zmitrovich (2018) introduced the monotypic genus
Hermanssonia to accommodate Phlebia centrifuga (V. Papp).

Herpomycetales Haelew. & Pfister


Haelewaters et al. (2019b) introduced this new order to accommodate the genus
Herpomyces, which was previously placed in Laboulbeniales. Blackwell et al. (2020) found high
support for the polyphyly of the thallus-forming Laboulbeniomycetes, i.e. orders Herpomycetales
and Laboulbeniales (D. Haelewaters).

Hesperomyces Thaxt.
Wijayawardene et al. (2017a) mentioned eleven species for this genus. This number will be
much higher, after Haelewaters et al. (2018a) found that H. virescens Thaxt. is a complex of
multiple species, each with their own host. At least 10 species can be recognized within H.
virescens sensu lato (D. Haelewaters, unpubl. data) (D. Haelewaters).

Heufleria Auersw.
Baral (2016) and Ekanayaka et al. (2019) accepted this genus in Rhytismataceae (N.
Wijayawardene & D. Haelewaters).

Hilidicellites Kalgutkar & Janson. (fossil)


Kalgutkar & Jansonius (2000) opined that Dicellaesporites appendiculatus Sheffy &
Dilcher is a misfit in Dicellaesporites Elsik and therefore they proposed Hilidicellites to
accommodate it (R.K. Saxena).

Hoffmannoscypha Stielow, Göker & Klenk


This genus, erected for a single species (Geopora pellita), was recently shown to represent a
separate phylogenetic lineage (Van Vooren et al. 2017) (I. Kušan & N. Matočec).

Hortaea Nishim. & Miyaji


This genus currently includes two species, H. werneckii and H. thailandica. Hortaea
werneckii is a well-known clinically relevant fungus, causing superficial skin infections in humans
(Marchetta et al. 2018) (H. Madrid).

Hyaloscyphaceae Nannf.
This family is currently polyphyletic. Based on a four-gene phylogenetic analysis, Han et al.
(2014) found that Hyaloscyphaceae sensu lato is a heterogeneous assemblage of 10
hyaloscyphaceous taxa. In Johnston et al. (2019), three highly supported clades were retrieved,
Han Clade 4 (Gamarada, Hyphodiscus, Hyphopeziza, Venturiocistella), Han Clade 7 (Amicodisca,
Dematioscypha), and Hyaloscyphaceae (“Chalara” longipes, Hyalopeziza, type genus

1402
Hyaloscypha, Meliniomyces, Olla, Rhizoscyphus). More taxa and sequence data are needed to
resolve the classification and taxonomy within Hyaloscyphaceae sensu lato (D. Haelewaters).

Hydnocystis Tul.
Stephensia Tul. & C. Tul. has been treated as a synonym of Hydnocystis by Kumar et al.
(2017) (P. Alvarado).

Hydnophanerochaete Sheng H. Wu & C.C. Chen


Chen et al. (2018) introduced this monotypic genus in Meruliaceae to accommodate
Phanerochaete odontoidea Sheng H. Wu (V. Papp).

Hypomontagnella Sir et al.


Lambert at al. (2019) erected the new genus Hypomontagnella Sir et al. (Hypoxylaceae)
based on a comparison of molecular and morphological data for Hypoxylon monticulosum and
similar species. Interestingly, all strains so far cultured produce the selective antifungal
polyketides of the sporothiolide type, which can therefore be regarded as a chemotaxonomic
marker (M. Stadler).

Hypoxylonites Elsik. (fossil)


Hypoxylonsporites P. Kumar is a later taxonomic synonym of Hypoxylonites Elsik (Elsik
1990) (R.K. Saxena).

Immotthia M.E. Barr


Hyde et al. (2017) accommodated Immotthia in Roussoellaceae based on morphological
characteristics resembling the genus Roussoella. However, phylogenetic affinity of the genus
needs to be confirmed by molecular data (R. Phookamsak).

Inapertisporites Hammen (fossil)


Triporisporonites Sheffy & Dilcher is a later taxonomic synonym of Inapertisporites.
Inapertisporites “Hammen ex Rouse” is illegitimate name being a later homonym (and later
taxonomic synonym) of Inapertisporites Hammen (R.K. Saxena).

Intralichen D. Hawksw. & M.S. Cole


This genus was introduced by Hawksworth & Cole (2002) to accommodate dematiaceous
hyphomycetes occurring in lichens, some of which were previously placed in Bispora or
Trimmatostroma. The only available DNA sequence is labelled ‘Bispora’ christiansenii IMI
227584, a sequence of 18S rDNA from a study by Sert et al. (2007), with the GenBank number
AM279680. BLAST searches with this sequence, however, revealed that it shows 99% identity to
yeasts of the genera Candida, Debaryomyces and Meyerozyma, indicating that probably a
contaminant was sequenced (H. Madrid).

Jansoniisporites Kalgutkar (fossil)


Brachysporisporites endophragmia Kalgutkar & Sigler is a misfit in Brachysporisporites
R.T. Lange & P.H. Sm. and therefore Kalgutkar (1997) proposed Jansoniisporites to
accommodate it (R.K. Saxena).

Japewia Tønsberg
Kistenich et al. (2018) transferred this genus from the Ramalinaceae to the Lecanoraceae
(E. Timdal)

Junewangiaceae J.W. Xia & X.G. Zhang


See under Acrodictyaceae (J. Ma).

1403
Kellermania Ellis & Everh.
Based on phylogenetic information and morphology, Minnis et al. (2012) placed Alpakesa,
Piptarthron, Planistroma, and Planistromella as synonyms of Kellermania (A.J.L. Phillips).

Khaleijomyces Abdel-Wahab
Abdel-Wahab et al (2018) erected this new genus for a new marine species within
Juncigenaceae (S. Fryar).

Kiliasia Hafellner
Kistenich et al. (2018) resurrected this genus in the Ramalinaceae (E. Timdal).

Knufia L.J. Hutchison & Unter.


The molecular phylogeny and taxonomy of this genus has been studied by Tsuneda et al.
(2011b), Isola et al. (2016) and Mehrabi et al. (2018) among others, revealing its placement in
Trichomeriaceae (H. Madrid).

Kumarisporites Kalgutkar & Janson. (fossil)


Imprimospora ramanujamii Kumar is a misfit in Imprimospora and therefore Kalgutkar &
Jansonius (2000) proposed Kumarisporites to accommodate it (R.K. Saxena).

Kusaghiporia J. Hussein et al.


Hussein et al. (2018) described this genus with K. usambarensis Hussein J., Tibell S. &
Tibuhwa as the type species and showed that it belongs to Laetiporaceae (V. Papp).

Kutchiathyrites R.K. Kar (fossil)


Kalgutkar & Jansonius (2000) opined that Kutchiathyrites is a multicellular spore/ conidium
showing a clear attachment area, scar or pore. Kutchiathyrites eccentricus R.K. Kar demonstrates
close similarity to the conidia of the hyphomycetous fungus Mycocenterolobium platysporum
Goos (R.K. Saxena).

Laboulbenia Mont. & C.P. Robin


The most recent number of species in this genus is 633 (Song et al. 2019). This number will
likely increase in the future; based on a preliminary concatenated ITS-LSU rDNA dataset,
Haelewaters et al. (2019a) showed that one of the most cosmopolitan taxa in the genus, L.
flagellata Peyr., is a complex of species. A number of taxa were recently lectotypified in
Haelewaters et al. (2015, 2019a) (D. Haelewaters).

Lachnopsis Guatimosim et al.


Lachnopsis was introduced in Lachnaceae to accommodate two species that are only
distinguishable from Lachnum based on DNA sequence data (Guatimosim et al. 2016) (D.
Haelewaters).

Lasionectriella Lechat & J. Fourn.


Lechat et al. (2016a) described this genus to accommodate two species within the family
Bionectriaceae (I. Kušan & N. Matočec).

Lecanoraceae Körb. (= Carbonicolaceae Bendiksby & Timdal)


Kraichak et al. (2018a) regarded Carbonicolaceae as a synonym of Lecanoraceae (N.
Wijayawardene).

Lentimurispora N.G. Liu et al.

1404
Liu et al. (2018a) introduced this genus and Lentimurisporaceae which has a distinct lineage
in Pleosporales (S. Fryar).

Lentistoma A. Hashim.
Hashimoto et al. (2018) proposed this genus to accommodate Lentistoma bipolare, which
was transferred from Lophiostoma based on molecular and morphological characters (S. Fryar).

Lentithecium K.D. Hyde et al.


Lentithecium aquaticum Yin. Zhang, J. Fourn. & K.D. Hyde, L. arundinaceum (Sowerby)
K.D. Hyde, J. Fourn. & Yin. Zhang, and L. lineare (E. Müll. ex Dennis) K.D. Hyde, J. Fourn. &
Yin. Zhang do not group with L. fluviatile (Aptroot & Van Ryck.) K.D. Hyde, J. Fourn. & Yin.
Zhang, the type species of the genus Lentithecium in molecular phylogenetic analysis (MycoBank:
see under Lentithecium carbonneanum J. Fourn., Raja & Oberlies) (H. Raja).

Leptoparies A. Hashim.
Hashimoto et al. (2018) proposed this genus to accommodate a single species within the
Lophiostomataceae (S. Fryar).

Leptosillia Höhn.
Voglmayr et al (2019) demonstrated that the genus Leptosillia belongs to the Xylariales
based on a multi locus DNA sequence analyses of SSU-ITS-LSU rDNA, rpb1, rpb2, tef1 and
tub2. They also established the genera Cresporhaphis and Liberomyces are congeneric with
Leptosillia and erected the new family Leptosilliaceae. A number of taxa were epi-or
lectotypified, and the new genus Furfurella was erected in family Delonicicolaceae (M. Stadler).

Lichinodium Nyl.
Lichinodium was previously placed in class Lichinomycetes. However, Prieto et al. (2019)
showed that this genus is placed within Leotiomycetes as a previously unrecognized lineage and
introduced a new order (Lichinodiales) and family (Lichinodiaceae) (M. Prieto).

Lithophila Selbmann & Isola


This monotypic genus, typified by L. guttulata, occurs on marble and was placed in
Trichomeriaceae based on multilocus phylogenetic analyses (Isola et al. 2016) (H. Madrid).

Liua Phook. & K.D. Hyde


Phookamsak et al. (2019) introduced a monotypic genus Liua to accommodate
camarosporium-like species and is typified by L. muriformis Phookamsak, H.B. Jiang & K.D.
Hyde. Multi-gene phylogenetic analyses showed that Liua formed a sister lineage with Cycasicola
in Thyridariaceae (R. Phookamsak).

Lodderomyces Van der Walt


Expected transfer of species to Candida Berkhout to comply with the Melbourne Code
(Kurtzman & Boekhout 2017) (W.P. Pfliegler and E. Horváth).

Lonicericola Phook. et al.


Phookamsak et al. (2019) introduced the new genus Lonicericola based on DNA sequence
analyses. The genus formed a distinct clade closely related with Pseudomonodictys and
Paratrimmatostroma in Parabambusicolaceae (R. Phookamsak).

Lotinia Pérez-Butrón et al.


This genus, erected for a single species (L. verna), represents a separate phylogenetic
lineage (Van Vooren et 2017) (I. Kušan & N. Matočec).

1405
Manglicola Kohlm. & E. Kohlm.
Manglicola consists of two species, but molecular data supports only M. guatemalensis
Kohlm. & E. Kohlm. showing phylogenetic affiliations to Manglicolaceae, while M. samuelsii
Huhndorf, was placed in the Hypsostromataceae based on morphological data (Huhndorf 1994).
Sequence data from M. samuelsii is necessary to place it within a phylogenetic framework and
assess if the genus is polyphyletic within the Dothideomycetes (H. Raja).

Marasasiomyces Crous
Based on ITS and LSU sequence phylogeny, Crous et al. (2015a) introduced
Marasasiomyces as a new genus for tiarosporella-like fungi with long-necked conidiomata
covered in brown setae (A.J.L. Phillips).

Marthamycetales P.R. Johnst. & Baral


Introduced in Johnston et al. (2019) to accommodate the phylogenetically isolated family
Marthamycetaceae within Leotiomycetes (D. Haelewaters).

Masonhalea Kärnefelt
Thell et al. (2018) proposed to resurrect the genus (A. Tsurykau).

Massariosphaeria (E. Müll.) Crivelli


This genus was firstly recognized as a section of Leptosphaeria (Müller 1950). However,
Massariosphaeria was introduced by Crivelli (1983). Wang et al. (2007) showed that
Massariosphaeria is polyphyletic. However, the type species M. phaeospora is closely related to
the type species of Cyclothyriellaceae, Cyclothyriella rubronotata. Therefore, Massariosphaeria
is placed in Cyclothyriellaceae (Jaklitsch et al. 2016a, Wijayawardene et al. 2018a) (S.
Hongsanan).

Mathurisporites Kalgutkar & Janson. (fossil)


Kalgutkar & Jansonius (2000) opined that Pluricellaesporites ellipticus Y.K. Mathur & K.
Mathur is a misfit in Pluricellaesporites Hammen and therefore they proposed Mathurisporites to
accommodate it (R.K. Saxena).

Megalaria Hafellner
Kistenich et al. (2018) regarded that Catillochroma Kalb and Lopezaria Kalb & Hafellner as
synonyms of this genus (E. Timdal).

Melaspileellaceae D.Q. Dai & K.D. Hyde


Dai et al. (2018) introduced this family in Asterinales to accommodate Melaspileella (P.
Karst.) Vain (N. Wijayawardene).

Metschnikowiaceae T. Kamienski
Family status is expected to change following the phylogenetic relationships of the genera of
Debaryomycetaceae and Metschnikowiaceae (Shen et al. 2016) (W.P. Pfliegler & E. Horváth).

Micraspidales Quijada & Tanney


Previously, the genus Micraspis was placed in different families (Cryptomycetaceae,
Helotiaceae, Phacidiaceae, Tympanidaceae). Because of its isolated phylogenetic position within
Leotiomycetes, Quijada et al. (2020) established the family Micraspidaceae and order
Micraspidales to accommodate the genus (D. Haelewaters).

Microcaliciaceae Tibell

1406
Kraichak et al. (2018a) accepted the family as in Pertusariales (N. Wijayawardene).

Microkamienskia Corazon-Guivin et al.


Corazon-Guivin et al. (2019) introduced Microkamienskia with two new combinations. This
genus is an arbuscular mycorrhizal fungus and similar to Kamienskia in spore but differ in size
(Błaszkowski et al. 2015) (S. Tibpromma).

Minimelanolocus R.F. Castañeda & Heredia


This genus is currently considered a putative member of Herpotrichiellaceae, although no
DNA sequence data is available for the type species, M. navicularis (H. Madrid).

Mniaeciaceae Baral
Introduced in Johnston et al. (2019), elevating the Mniaecia lineage sensu Baral (2016) to
family level. Mniaeciaceae is sister to Leotiaceae within Leotiales (D. Haelewaters).

Monoporisporites Hammen (fossil)


Polyporisporites Hammen, Psiammopomopiospora Sal.-Cheb. & Locq. and
Reticulatisporonites Elsik are later taxonomic synonyms of Monoporisporites Hamme Dong n
(Kalgutkar & Jansonius 2000) (R.K. Saxena).

Mortierella Coem.
There are 247 records of Mortierella species in the Species Fungorum, but according to
Yadav et al. (2015) nearly 100 of validated species have been described (A. L. C. M. de A.
Santiago).

Mossopisporites Kalgutkar & Janson. (fossil)


Kalgutkar & Jansonius (2000) opined that Triporicellaesporites multicellulus Ke & Shi is a
misfit in Triporicellaesporites Ke & Shi and therefore they proposed Mossopisporites to
accommodate it (R.K. Saxena).

Mucoharknessia Crous et al.


Based on ITS and LSU sequence phylogeny, Crous et al. (2015a) introduced
Mucoharknessia as a new genus for tiarosporella-like fungi that resembles Harknessia
(Harknessiaceae, Diaporthales), but are distinguished by having pycnidia that lack furfuraceous
tissue around the ostiole and conidia with a mucoid apical appendage (A.J.L. Phillips).

Mucor Fresen.
The greatest number of mucoralean species described to date belongs to Mucor with more
than 300 species cited in literature (Jacobs & Botha 2008, Álvarez et al. 2011). Although the exact
number of valid taxa is unknown, Gherbawy et al. (2010) stated the number of species may have
ranged from 50 to 75 at the time of the study. Based on morphological characteristics, maximum
growth temperature and mating experiments, Schipper (1973, 1975, 1976, 1978) monographed the
genus and described 39 species, four varieties and 11 forms. Knowledge of the genus was
subsequently expanded with a description of 26 new taxa [Mehrotra & Mehrotra (1979), Mirza et
al. (1979), Subrahamanyam (1983), Chen & Zheng (1986), Schipper & Samson (1994), Watanabe
(1994), Zalar et al. (1997), Pei (2000), Alves et al. (2002), Jacobs & Botha (2008), Hermet et al.
(2012), Madden et al. (2012), Voglmayr & Clémençon (2016), Li et al. (2016), Lima et al. (2017),
Wanasinghe et al. (2018), De Souza et al. (2018), de Lima et al. (2018)]. (A. L. C. M. de A.
Santiago)

Muellerella Müll. Arg.

1407
Muggia et al. (2015) hypothesized that the genus represents sexual stage of Lichenodiplis
(A. Suija).

Multicellaesporites Elsik emend. P. Kumar (fossil)


Kumar (1990) emended the generic diagnosis. Warkallisporonites Ramanujam & K.P. Rao
is a later taxonomic synonym of Multicellaesporites Elsik (R. K. Saxena).

Mycobilimbia Rehm
Kistenich et al. (2018) transferred this genus from the Lecideaceae to the Ramalinaceae (E.
Timdal).

Mycocarpon S.A. Hutch. (fossil)


Hutchinson (1955) opined that Sporocarpon pachydermum Will. is a misfit in Sporocarpon
Will. and therefore he proposed Mycocarpon to accommodate it (R.K. Saxena).

Mycoceros D. Magyar & Z. Merényi


This genus was recently described to accommodate a species parasitizing Pinaceae grain
pollens. Based on an ITS+LSU phylogeny, it was clearly placed within Orbiliomycetes by Magyar
et al. (2018) (I. Kušan & N. Matočec).

Mycomicrothelia Keissl.
All the tropical species have been placed in Bogoriella (Trypetheliaceae, Trypetheliales),
but type and other temperate taxa are still in Mycomicrothelia (Aptroot & Lücking 2016) (A.
Aptroot).

Mycopappus Redhead & G.P. White


Baral (2016) and Ekanayaka et al. (2019) accepted this genus as in Sclerotiniaceae (N.
Wijayawardene).

Mycosphaerellaceae Lindau
We accept only 111 genera which have been confirmed as well-established genera in
Mycosphaerellaceae by phylogenetic analyses. Hongsanan et al. (2020) lists doubtful genera in
Mycosphaerellaceae based on Videira et al. (2017) (N. Wijayawardene & R. Phookamsak).

Myelorrhiza Verdon & Elix


Kistenich et al. (2018) transferred this genus from the Cladoniaceae to the Ramalinaceae
(E. Timdal).

Myochroidea Printzen et al.


Printzen et al. (2008) accommodate Myochroidea including four species of the Lecidea
leprosula group such as M. leprosula (Arnold) Printzen, T. Sprib. & Tønsberg, M. porphyrospoda
(Anzi) Printzen, T. Sprib. & Tønsberg, M. rufofusca (Anzi) Printzen, T. Sprib. & Tønsberg and M.
minutula Printzen, T. Sprib. & Tønsberg based on morphological key characters. Although no
phylogenetic analyses have been performed, the authors suggested to include this genus either in
Psoraceae, Pilocarpaceae, or Ramalinaceae but to confirm the final placement of Myochroidea in
the system, molecular analyses are needed (P. Rodriguez-Flakus).

Myrmecocystis Harkn.
This genus was recently resurrected by Alvarado et al. (2018) (P. Alvarado).

Myxotrichaceae Currah

1408
Johnston et al. (2019) found high support for the placement of this family in the pezizelloid
clade of Helotiales, placed sister to Amorphothecaceae. In the ITS tree of Johnston et al. (2019),
Amorphothecaceaea consisting of Byssoascus, "Malbranchea" flavorosea, Myxotrichum and
Oidiodendron was retrieved in a maximum supported branch sister to Amorphotheca resinae (D.
Haelewaters).

Neocelosporium Crous
Crous et al. (2018) introduced a new family Neocelosporiaceae and new order
Neocelosporiales to accommodate the genus Neocelosporium, which represents a distinct lineage
in Dothideomycetes. However, Hongsanan et al. (in prep.) found that Neocelosporiales is placed
within the order Dothideales. As a result, Neocelosporiales is here regarded as a synonym of
Dothideales and Neocelosporiaceae is accommodated in Dothideales (N. Wijayawardene & S.
Hongsanan).

Neodendryphiella Iturrieta-González et al.


Iturrieta-González et al. (2018) introduced this new genus with three new species in
Dictyosporiaceae. Currently, the genus comprises three species viz. N. mali, N. michoacanensis
and N. tarraconensis (the type species). Neodendryphiella differs from Dendryphiella in lacking
of nodulose conidiophores bearing conidiogenous cells with pores surrounded by a thickened and
darkened wall (S. Fryar & S. Somrithipol).

Neoeutypella M. Raza et al.


Neoeutypella was introduced as a monotypic genus in Phookamsak et al. (2019) to
accommodate N. baoshanensis M. Raza et al. and the strains identified as “Eutypella caricae
(strains EL51C and GL08362)”. Based on phylogenetic analyses, Neoeutypella formed a distinct
lineage, clustered with Diatrypella species but the genus differs from Diatrypella in having large
entostromata, 8-spored, spindle-shaped asci and allantoid ascospores (Phookamsak et al. 2019) (R.
Phookamsak).

Neolauriomycetaceae Crous
Neolauriomycetaceae was introduced within Helotiales by Crous et al. (2018) to
accommodate three genera: Exochalara, Lareunionomyces, and Neolauriomyces (D. Haelewaters).

Neomelanconiella Crous
Crous et al. (2018) introduced this genus and showed that it has a distinct lineage in
Diaporthales. A new family Neomelanconiellaceae is introduced (N. Wijayawardene).

Neomelanconiella Crous
Crous et al. (2018) introduced this genus and showed that it has a distinct lineage in
Diaporthales. Hence introduced the new family Neomelanconiellaceae (N. Wijayawardene).

Neoscytalidium Crous & Slippers


Two species of Neoscytalidium were frequently reported from opportunistic infections in
humans, i.e. N. dimidiatum and N. hyalinum. The latter species, however, is currently considered a
hyaline mutant of N. dimidiatum (Huang et al. 2016). Neoscytalidium currently includes three
species (H. Madrid).

Neoseptorioides Crous et al.


This genus was introduced for species morphologically distinct from Septorioides (Crous et
al. 2015a) (A.J.L. Phillips).

Neostagonosporella C.L. Yang et al.

1409
Yang et al. (2019) introduced a holomorph genus Neostagonosporella to accommodate
massarina-like taxon collected from living bamboo culms from China. Multi-gene phylogenetic
analyses revealed the genus in Phaeosphaeriaceae (R. Phookamsak).

Neptunomyces M. Gonçalves et al.


This monotypic genus was recently introduced by Gonçalves et al. (2019). Based on
phylogenetic analysis this genus is closest to Xenocamarosporium but conidial morphology is
distinct (Gonçalves et al. 2019) (S. Tibpromma).

Noosia Crous et al.


Recent phylogenetic analyses indicated that Noosia belongs to Periconiaceae (Tanaka et al.
2015, Thambugala et al. 2017) (D. Wanasinghe).

Nothomitra Maas Geest.


According to Hustad et al. (2013) and Hustad & Miller (2015), this genus is
phylogenetically close to Sarcoleotia in a basal clade of Geoglossomycetes (I. Kušan & N.
Matočec).

Oblongocollomyces Tao Yang & Crous


This genus was introduced by Yang et al. (2017) to accommodate Sphaeropsis variablis
(A.J.L. Phillips).

Odontoefibula C.C. Chen & Sheng H. Wu


Chen et al. (2018) introduced this monotypic genus in Phanerochaetaceae to accommodate
the new species Odontoefibula orientalis C.C. Chen & Sheng H. Wu (V. Papp).

Odoria V. Papp & Dima


Papp & Dima (2018) introduced this new monotypic genus in Meruliaceae to accommodate
the threatened old-growth forest polypore, Aurantiporus alborubescens (Bourdot & Galzin) H.
Jahn (V. Papp).

Ophiobolopsis Phook. et al.


Phookamsak et al. (2017) introduced Ophiobolopsis to accommodate ophiobolus-like
species in Phaeosphaeriaceae based on multi-gene phylogenetic analyses (R. Phookamsak).

Opilionomyces Santam. et al.


Santamaria et al. (2017) introduced this monotypic genus and confirmed its placement in
family Laboulbeniaceae, subfamily Laboulbenioideae, tribe Laboulbenieae based on
morphological characters (D. Haelewaters).

Ornasporonites Ramanujam & K.P. Rao (fossil)


This monotypic genus differs from Fusiformisporites because the latter possesses only a
single septum and is longitudinally ribbed and inaperturate (R.K. Saxena).

Oscarbrefeldia Holterm.
Doubtful genus not treated by Kurtzman (2011) (W.P. Pfliegler & E. Horváth).

Ovadendron Sigler & J.W. Carmich.


This fungal genus is listed as a member of Onygenales in de Hoog et al. (2015), but its
family placement needs to be thoroughly assessed (H. Madrid).

Pachydisca Boud.

1410
Dumont (1975) proposed to exclude the genus from Sclerotiniaceae and place it in
Helotiales based on morphological study of the type species, P. guernisacii. Jaklitsch et al. (2016a)
treated this genus as a synonym of Discinella Boud. However, Species Fungorum (2020) lists 32
accepted species. In this outline, Pachydisca is regarded as Helotiales genera incertae sedis (D.
Haelewaters).

Pachyphlodes Zobel
Scabropezia Dissing & Pfister has been regarded as a synonym of Plicariella or
Pachyphlodes Zobel in Healy et al. (2018) (P. Alvarado).

Palaeoamphisphaerella Ramanujam & Srisailam (fossil)


Imprimospora Norris is considered as a later synonym of Palaeoamphisphaerella (R.K.
Saxena).

Palaeomycites Mesch. (fossil)


Phycomycites Ellis, Palaeomyces Renault ex Kidston & Lang, Rhizophagites E.J. Butler ex
Rosend., Propythium Elias, Aplanosporites R.K. Kar, Glomites T.N. Taylor, W. Remy, Hass &
Kerp are later taxonomic synonyms of Palaeomycites (R.K. Saxena).

Palaeopericonia Ibañez & Zamuner (fossil)


The material is made up of only asexual structures represented by conidia produced on
single conidiophores. The closely related genera are Periconia Tode, Torula Pers., Stachybotrys
Corda, Humicola Traaen, Thermomyces Tsikl. and Chlamydomyces Bainier (Ibañez & Zamuner
1996) (R.K. Saxena).

Palaeophoma Singhai (fossil)


One-celled hyaline, bent or curved or lunate conidia, and a spherical and brown pycnidium
have been shared by the living genus Selenophoma Maire (Barnet 1960). But the fossil fungus
also possesses spherical conidia which are not present in Selenophoma. Moreover, Palaeophoma
has non-ostiolate pycnidium whereas Selenophoma possesses a definite ostiole (R.K. Saxena).

Palaeosclerotium G.W. Rothwell (fossil)


Dennis (1976) opined that Palaeosclerotium represents an intermediate evolutionary stage
between Ascomycetes and Basidiomycetes. Pirozynski & Weresub (1979) stated that
Palaeosclerotium is neither an ascomycete nor a basidiomycete, but an early dikaryotic fungus
and a representative of a group that links Basidiomycota with extinct, probably symbiotic, lichen-
like nematophytes (R.K. Saxena).

Paleoslimacomyces Kalgutkar & Sigler (fossil)


Conidia of this monotypic genus show some morphological similarity with those of extant
Slimacomyces monospora (W.B. Kendr.) Minter, which was originally described by Kendrick
(1958) in Helicoma Corda (R.K. Saxena).

Pappia Zmitr.
Papp & Dima (2018) showed that Aurantiporus fissilis (Berk. & M.A. Curtis) H. Jahn ex
Ryvarden forms a distinct phylogenetic lineage in the family Meruliaceae. Hence, Zmitrovich
(2018) introduced the new monotypic genus Pappia in Meruliaceae to accommodate
Aurantiporus fissilis (V. Papp).

Papulosporonites Schmied. & A.J. Schwab (fossil)


In Polyadosporites Hammen, the individual cells are less tightly appressed into a spherical
mass than those in Papulosporonites (R.K. Saxena).

1411
Paracladophialophora Crous
This genus currently includes two species, P. carceris and P. cyperacearum. They form a
distinct clade in Chaetothyriales for which the family Paracladophialophoraceae was recently
proposed (H. Madrid).

Parafenestella Jaklitsch & Voglmayr


Jaklitsch et al. (2018) proposed this new genus in the Cucurbitariaceae based on
morphological and molecular characters. (S. Fryar)

Parallopsora Kistenich et al.


Kistenich et al. (2018) introduced this genus in Ramalinaceae (E. Timdal).

Paraophiobolus Phook. et al.


Phookamsak et al. (2017) introduced Paraophiobolus to accommodate ophiobolus-like
species in Phaeosphaeriaceae based on multi-gene phylogenetic analyses (R. Phookamsak).

Pararoussoella Wanas. et al.


Wanasinghe et al. (2018) introduced a monotypic genus to accommodate roussoella-like
species in Thyridariaceae. However, Phookamsak et al. (2019) accommodated the genus in
Roussoellaceae based on multi-gene analyses and this concurred with Jiang et al. (2019a) and
Karunarathna et al. (2019). (R. Phookamsak)

Paratrimmatostroma Jayasiri et al.


Phookamsak et al. (2019) introduced the new genus Paratrimmatostroma based on multi-
gene phylogenetic analyses coupled with morphological characteristic. Paratrimmatostroma is
sister to Pseudomonodictys in Parabambusicolaceae based on phylogenetic analyses of a
combined SSU, ITS, LSU and TEF1-α sequence dataset (Phookamsak et al. 2019).
Paratrimmatostroma can be distinguished from Pseudomonodictys in forming sporodochia on
host substrate and having branched, straight or flexuous conidia, with variable conidial shape such
as helicoid, cylindrical, sigmoid, or reniform (Phookamsak et al. 2019) (R. Phookamsak).

Parazalerion Madrid et al.


Phookamsak et al. (2019) introduced a monotypic conidial genus to accommodate zalerion-
like taxon and is typified by P. indica Madrid, Gené, Cano & Guarro. The genus was isolated
from soil in India and is characterized by having irregularly coiled, dematiaceous, multiseptate
conidia which often form knots of cells. Phylogenetic analysis revealed that the genus formed a
sister lineage with Spirosphaera minuta in Microthyriales (R. Phookamsak & H. Madrid).

Parmulariales D.Q. Dai & K.D. Hyde


Dai et al. (2018) introduced this order to accommodate Parmulariaceae (N. Wijayawardene).

Patinella Sacc.
Patinella hyalophaea, the type species of the genus, was retrieved near Holwaya mucida and
its anamorph Crinula caliciiformis in Johnston et al.’s (2019) ITS tree, as Leotiomycetes genera
incertae sedis. In their 15-gene tree, the Holwaya–Crinula clade was placed sister to Thelebolales
(Pseudeurotiaceae, Thelebolaceae) with high support (D. Haelewaters).

Patinella Sacc.
The type species of Patinella was nested near the order Thelebolales (Hyde et al. 2017) by
two phylogenies (based on ITS and LSU) together with Ramgea Brumm. and Holwaya Sacc. as a
weakly supported sister clade to the family Thelebolaceae. More phylogenetic information is

1412
needed to ascertain the true position of these three genera. It is best to place this genus into
Phacidiales incertae sedis at the moment (I. Kušan & N. Matočec).

Peltigeraceae Dumort.
Kraichak et al. (2018a) regarded that Lobariaceae Chevall. and Nephromataceae Wetm. ex
J.C. David & D. Hawksw. are synonyms of Peltigeraceae (N. Wijayawardene).

Perennicordyceps Matočec & I. Kušan


The genus Perennicordyceps was erected by Matočec et al. (2014) to segregate a
monophyletic clade of four species aside from Polycephalomyces based on both molecular and
non-molecular evidence. According to Crous et al. (2017), this genus is phylogenetically placed
within Ophiocordycipitaceae as a sister clade to Polycephalomyces (I. Kušan & N. Matočec).

Pertusaria DC.
Several species were combined into other genera (Wei et al. 2017), but as no recent
taxonomic revision of the genus is available, the exact number of species is obscure (M. Kukwa).

Petrophila de Hoog & Quaedvl.


This monotypic genus, typified by P. incerta, was placed in the family Extremaceae in a
phylogenetic study by Isola et al. (2016) (H. Madrid).

Phacidiales Bessey
The order Phacidiales includes two families, Helicogoniaceae and Phacidiaceae (Johnston
et al. 2019). Two lineages that were previously recognized in Phacidiales (sensu Baral 2016),
Mniaeciaceae (referred to as Mniaecia lineage in Baral (2016)) and Tympanidaceae, are now
recognized within Leotiales (D. Haelewaters).

Phaeopoacea Thambug. et al.


Thambugala et al. (2017) introduced Phaeopoacea to accommodate phaeosphaeria-like taxa
in Phaeosphaeriaceae and is typified by Phaeopoacea festucae. Thambugala et al. (2017) also
transferred Phaeosphaeria phragmiticola Leuchtm. to Phaeopoacea as P. phragmiticola
(Leuchtm.) Thambugala & K.D. Hyde based on molecular data. The sexual and asexual morph
connection of this genus is well-resolved (R. Phookamsak).

Phialina Höhn.
Baral (2016) and Ekanayaka et al. (2019) accepted this genus in Pezizellaceae (N.
Wijayawardene).

Phialocephala W.B. Kendr.


Phialocephala is currently placed in Mollisiaceae based on the phylogenetic reconstruction
of a 15-gene dataset. It should be noted that consensus remains unclear about the systematic
position of several mollisioid genera, including Mollisia and Phialocephala (Tanney & Seifert
2020) (D. Haelewaters).

Phialophora Medlar
This genus historically included a heterogeneous assemblage of phialidic dematiaceous
hyphomycetes usually with poorly developed conidiophores, producing phialides with
conspicuous collarettes. It comprised members of several families, orders and classes of
ascomycetes (Gams 2000). The current concept of the genus, however, only includes phialidic
members of Herpotrichiellaceae with or without a yeast phase in culture (Li et al. 2017) (H.
Madrid).

1413
Phragmonaevia Rehm
Kirk et al. (2008) regarded this name as doubtful, but Baral (2016) listed it among
Helotiales genera incertae sedis. No sequences are currently available for any member of this
genus (N. Wijayawardene).

Phragmothyrites W.N. Edwards (fossil)


Microthallites Dilcher is a later taxonomic synonym of Phragmothyrites (R.K. Saxena).

Phyllopsora Müll. Arg.


Kistenich et al. (2018) regarded that Crocynia (Ach.) A. Massal. as a synonym of this genus
(E. Timdal).

Piricauda Bubák
Da Silva et al. (2016) showed that Piricauda paraguayensis could be accommodated in
Mycosphaerellaceae. However, it should be pointed out that Piricauda paraguayensis is not the
type species, thus, we tentatively keep this genus in Mycosphaerellaceae (N. Wijayawardene, S.
Hongsanan & R. Phookamsak).

Picoa Vittad.
The genus Phaeangium Pat. Has been regarded as a synonym of Picoa, as discussed in
Zitouni-Haouar et al. (2015) (P. Alvarado).

Pilatotrama Zmitr.
Zmitrovich (2018) introduced this new monotypic genus in Polyporaceae to accommodate
Trametes ljubarskyi Pilát. (V. Papp).

Placocrea Syd.
Boonmee et al. (2017) treated this genus as a member in Teratosphaeriaceae (S. Boonmee).

Plagiosphaera Petr.
Voglmayr in Song et al. (2019) demonstrated that the taxon P. immersa Petr. belongs to the
family Magnaporthaceae (Magnaporthales) based on a multi-locus phylogenetic study of ITS-
LSU-rpb1-tef1 DNA sequence analyses. The other members of the genus are listed as
Sordariomycetes incertae sedis (Huhndorf et al. 2004, Index Fungorum 2019) (D. Haelewaters).

Pleuromyces Dima, P.-A. Moreau & V. Papp


Crous et al. (2018) introduced this monotypic genus to accommodate the new species P.
hungaricus V. Papp, Dima & P.-A. Moreau, and accepted it as a genus in Tubariaceae based on
phylogenetic analyses (V. Papp).

Plicariella (Sacc.) Rehm


See under Pachyphlodes Zobel (P. Alvarado).

Pluricellaesporites Hammen (fossil)


Piriurella Cookson & Eisenack is a later taxonomic synonym of Pluricellaesporites
Hammen (R.K. Saxena).

Polycellaesporonites Anil Chandra et al. (fossil)


Capsular, muriform fungal spores with a hilum, and distally with an elongated, knob-like or
beaked, extension as that in the modern Alternaria (R.K. Saxena).

Polycephalomyces Kobayasi

1414
After segregation of the genus Perennicordyceps (Matočec et al. 2014) and the description
of several new species in the genus, Polycephalomyces currently includes 18 species (Xiao et al.
2018) (D. Haelewaters).

Prathoda Subram.
Simmons (2007) resurrected Prathoda which is distinct from Alternaria (Pleosporaceae). In
MycoBank and Species Fungorum, Prathoda is mentioned as a synonym of Alternaria, but its
molecular phylogeny has not been recovered and closest relationship with Alternaria has not been
settled. Therefore we left Prathoda as a separate genus (P.B. Gannibal).

Proliferophorum Wang et al.


Phookamsak et al. (2019) introduced a monotypic genus Proliferophorum to accommodate
hyphomycetous species, P. thailandicum G.N. Wang et al. in Diaporthomycetidae based on
phylogenetic analysis. The genus is characterized by having mononematous, caespitose
conidiophores, polyblastic, terminal, sympodial, pale brown or subhyaline, with minute, truncate
conidiogenous cells, sometimes percurrently proliferating 1–2 times at broken ends of
conidiogenous cells and fusiform to cylindrical, pigmented, septate conidia. The genus was
collected from decaying submerged wood in Thailand (R. Phookamsak).

Protofenestella Jaklitsch & Voglmayr


Jaklitsch et al. (2018) proposed this new genus in the Cucurbitariaceae based on
morphological and molecular characters (S. Fryar).

Protothelenellaceae Vezda et al.


Kraichak et al. (2018a) regarded that Thrombiaceae Poelt & Vezda ex J.C. David & D.
Hawksw. as a synonym of Protothelenellaceae. Further, the family has been transferred to
Baeomycetales from Ostropales (N. Wijayawardene).

Pseudaegerita J.L. Crane & Schokn.


This aero-aquatic hyphomycete genus was shown to be a member of Hyaloscyphaceae
based on morphology of the associated sexual state (Abdullah et al. 2005) and DNA sequence data
(Johnston et al. 2019, Vu et al. 2019) (H. Madrid).

Pseudoanungitea Crous
Crous et al. (2018) introduced this new genus in Venturiaceae (S. Fryar).
Pseudoastrosphaeriellopsis Devadatha et al.
Phookamsak et al. (2019) introduced Pseudoastrosphaeriellopsis as a monotypic genus in
Pseudoastrosphaeriellaceae to accommodate trematosphaeria-like taxon. The genus is typified by
Pseudoastrosphaeriellopsis kaveriana Devadatha et al. collected from decaying wood of
Avicennia marina (Forssk.) Vierh. and Suaeda monoica Forssk. ex J.F.Gmel. in India. Based on
multi-gene phylogenetic analyses, Pseudoastrosphaeriellopsis formed a distinct lineage basal to
Pseudoastrosphaeriella (R. Phookamsak).

Pseudobambusicola Hern.-Restr. & Crous


This new genus was placed in Sulcatisporaceae using multi-gene phylogenetics and
morphological characters by Rupcic et al. (2018). It is closely related to Neobambusicola but
differs in having cylindrical-necked conidiomata surrounded by dark brown, smooth to slightly
verruculose hyphae (Rupcic et al. 2018) (S. Fryar & S. Somrithipol).

Pseudofusicoccum Mohali
Yang et al. (2017) considered that this genus warrants a separate family and thus introduced
Pseudofusicoccumaceae to accommodate it. Phillips et al. (2019) took into account phylogeny

1415
(ITS, LSU), morphology and evolutionary divergence times and concluded that Pseudofusicoccum
resides within Phyllostictaceae. Hence, Phillips et al. (2019) synonymised
Pseudofusicoccumaceae under Phyllostictaceae (A.J.L. Phillips).

Pseudographis Nyl.
Based on the phylogenetic analysis of a three-gene dataset (ITS, LSU, mtSSU), Karakehian
et al. (2019) placed Pseudographis in Rhytismataceae, not Triblidiaceae. Because of the inclusion
of Pseudographis in the family, the authors expanded the morphological description of
Rhytismataceae to include “ascospore cell walls that produce a strong blue/purple reaction in
iodine-based reagents” (D. Haelewaters).

Pseudogymnoascus Raillo
In Johnston et al.’s (2019) 15-gene tree, Pseudogymnoascus was strongly supported as sister
genus to Leuconeurospora within Pseudeurotiaceae, Thelebolales. In a genomic-scale tree based
on 3156 single-copy genes, Pseudogymnoascus destructans was sister to Thelebolus microsporus,
confirming its position in the order Thelebolales (Johnston et al. 2019) (D. Haelewaters).

Pseudolanzia Baral & G. Marson


Introduced to accommodate a species that bears morphological similarities to Lanzia but is
phylogenetically distinct. It is placed on a long branch within Rutstroemiaceae based on an
ITS+LSU phylogeny (Baral 2019) (D. Haelewaters).

Pseudomelanconidaceae C.M. Tian & X.L. Fan


The asexual morph of the family Pseudomelanconidaceae is somewhat similar to members
of Melanconiellaceae, and Juglanconidaceae. However, phylogenetic inferences resolved this
family as an individual group with well-supported group from other families of Diaporthales (Fan
et al. 2018a) (S.S.N. Maharachchikumbura & S. Fryar).

Pseudomelanconis C.M. Tian & X.L. Fan


Pseudomelanconis caryae is the type species of new genus Pseudomelanconis, and only
occurs on Carya cathayensis in China (Fan et al. 2018a) (S.S.N. Maharachchikumbura).

Pseudoneoconiothyrium Wanas. et al.


Wanasinghe et al. (2018) introduced a monotypic genus to accommodate neoconiothyrium-
like species in Thyridariaceae. However, Phookamsak et al. (2019) accommodated the genus in
Roussoellaceae based on multi-gene analyses and this concurred with Jiang et al. (2019a) and
Karunarathna et al. (2019) (R. Phookamsak).

Pseudoophiobolus Phook. et al.


Phookamsak et al. (2017) introduced Pseudoophiobolus to accommodate ophiobolus-like
species in Phaeosphaeriaceae based on multi-gene phylogenetic analyses including P. achilleae,
P. erythrosporus, P. galii, P. italicus, P. mathieui, P. rosae, P. subhyalinisporus and P. urticicola
(R. Phookamsak).

Pseudopaucispora A. Hashim.
Hashimoto et al. (2018) introduced this monotypic genus in Lophiostomataceae based on
molecular and morphological characters (S. Fryar).

Ramalinaceae C. Agardh
The taxonomy of family Ramalinaceae was recently revised by Kistenich et al. (2018).
According to phylogenetic analysis genera Adelolecia, Catinaria, Compsocladium,
Crustospathula, Frutidella, Japewia, Schadonia, Tasmidella do not belong this family (A. Suija).

1416
Ramasricellites Kalgutkar & Janson. (fossil)
Kalgutkar & Jansonius (2000) opined that Multicellaesporites differentialis Ramanujam &
Srisailam is a misfit in Multicellaesporites and therefore they proposed Ramasricellites to
accommodate it. The sharp differentiation between the dark, broad central cells and the narrower,
elongate hyaline terminal cells, as well as the lack of constriction at the median septum,
differentiate this form from species in Multicellites (R.K. Saxena).

Ramomarthamyces P.R. Johnst.


Marthamyces was found to be polyphyletic and therefore Johnston & Park (2019)
described Ramomarthamyces within Marthamycetaceae (Marthamycetales) for species separated
from Marthamyces sensu stricto. The four species in Ramomarthamyces have distinctly branched
rather than propoloid paraphyses (D. Haelewaters).

Ramophialophora M. Calduch et al.


This genus is polyphyletic within Sordariales (Zhang et al. 2017), but its type species, R.
vesiculosa is clearly phylogenetically placed in Lasiosphaeriaceae (Madrid et al. 2010) (H.
Madrid)

Ranadivia Zmitr.
Zmitrovich (2018) introduced this new genus in Fomitopsidaceae to accommodate
Daedalea allantoidea M.L. Han, B.K. Cui & Y.C. Dai, D. africana I. Johans. & Ryvarden, D.
stereoides Fr., and Polyporus modestus Kunze ex Fr. Based on a multi-gene phylogeny, Han et al.
(2016) accepted these species in the genus Daedalea Pers. (V. Papp).

Ratnagiriathyrites R.K. Saxena & N.K. Misra (fossil)


This monotypic genus is characterized by its non-radiating, hexagonal porate cells (R.K.
Saxena).

Remersonia Samson & Seifert


Wang et al. (2018) showed that the genus belongs in the Chaetomiaceae (K. Bensch).

Requienellaceae Boise
This family was introduced by Boise (1986) in the class Dothideomycetes (as
Loculoascomycetes) and she kept the family in Melanommatales (sensu Barr 1983) or Pyrenulales
(sensu Eriksson 1984). However, Requienellaceae was not treated as a distinguished family by
Hawksworth & Eriksson (1986), who maintained it under Pyrenulaceae. Barr (1990) and Aptroot
(1991) accepted Requienellaceae as a family of Melanommatales and again Kirk et al. (2008) as a
family of the Pyrenulales. Based on the sequence data, Jaklitsch et al. (2016b) reinstated
Requienellaceae as a family of Xylariales (S.S.N. Maharachchikumbura).

Resiniporus Zmitr.
Zmitrovich (2018) introduced this new genus in Irpicaceae to accommodate Ceriporiopsis
resinascens (Romell) Domański and C. pseudogilvescens (Pilát) Niemelä & Kinnunen (V. Papp).

Resinogalea Rikkinen & A.R. Schmidt


Rikkinen et al. (2016) proposed Resinogalea for Resinogalea humboldtensis collected from
resin of Araucaria humboldtensis in New Caledonia (S. Somrithipol).

Retihelicosporonites Ramanujam & K.P. Rao (fossil)

1417
Helical spores (conidia) are found in various hyphomycetes, viz. Helicoma Corda,
Helicomina L.S. Olive, Helicoon Morgan, Helicodendron Peyronel, Xenosporella Höhn, Hiospira
R.T. Moore, etc. (Barnett 1956, Ellis 1971, Ainsworth et al. 1973) (R.K. Saxena).

Retroconis de Hoog & Bat. Vegte


This genus belongs in Chaetomiaceae, Sordariales according to Crous et al. (2007) (H.
Madrid).

Rhamphoria Niessl
Ramphoria is the type genus of the newly erected family Rhamphoriaceae (Réblová &
Štěpánek (2018) (K. Bensch).

Rhamphoriaceae Réblová
Rhamphoriaceae is a novel family introduced by Réblová & Štěpánek (2018) to represent
genera Rhamphoria, Rhamphoriopsis, Linkosia and Xylolentia (S.S.N. Maharachchikumbura).

Rhamphoriopsis Réblová & Gardiennet


Genus in the Rhamphoriaceae with Rhamphoriopsis muriformis as the type species
(Réblová & Štěpánek 2018) (S.S.N. Maharachchikumbura).

Rhexoacrodictys W.A. Baker & Morgan-Jones


Xia et al. (2017) treated this genus as a member in Savoryellales, Savoryellaceae. (J. Ma)

Rhodoveronaea Arzanlou, W. Gams & Crous


Réblová & Štěpánek (2108) referred this genus to the newly erected family
Rhamphoriaceae (K. Bensch).

Rhizoglomus Sieverd. et al.


The genus Rhizophagus was not accepted in the Fungal Kingdom, as Rhizophagus
populinus is not an arbuscular mycorrhizal fungi but a plant root pathogen originally attributed to
the Peronosporales (Sieverding et al. 2014) which at time is attributed to the kingdom Chromista
(Cavalier-Smith 2018). Glomus intraradices became the type species of the new genus
Rhizoglomus with several new species described using Rhizoglomus as generic name (Sudová et al.
2015, Błaszkowski et al. 2018a, b, 2019a. b, Turrini et al. 2018).

Rimularia Nyl.
Four species were included in the phylogeny (Resl. et al. 2015), but 25 species still need
molecular data for the correct genus placement (M. Kukwa).

Roesleria Thüm. & Pass.


Baral (2016) maintained the family Roesleriaceae within his “Lineage B” (Helotiaceae
sensu lato) to accommodate the genus Roesleria with its peculiar morphological characteristics.
However, Johnston et al. (2019) found high support for the placement of this genus deep within
the family Helotiaceae (D. Haelewaters).

Roselymyces Fiuza et al.


The monotypic genus Roselymyces was erected in the Xylariales based on morphological
chacracters and a molecular phylogeny based on ITS and LSU data by Fiuza et al. (2018) with
Roselymyces brasiliensis as the type species. The genus was not yet associated to one of the
families of Xylariales, but shows morphological affinities to Cylindrium, Polyscytalum,
Pseudoidriella and Tristratiperidium (M. Stadler).

1418
Rostania Trevis.
According to Košuthová et al. (2019) the genus is not monophyletic and 2 species were
transferred to Leptogium and Scytinum. At present 3 species are known to belong to the genus in
its strict sense, but the species delimitation within Rostania needs further studies (M. Kukwa).

Rutstroemiaceae Holst-Jensen et al.


The Rutstroemiaceae + Sclerotiniaceae clade was retrieved with high statistical support in
the sclerotinioid clade of Helotiales in the 15-gene tree of Johnston et al. (2019). If
Sclerotiniaceae is retained, Rutstroemiaceae as currently recognized is not monophyletic and
would need to be split in four families. More multigene and genomic-scale work is needed to
resolve this sclerotinioid clade. Ekanayaka et al. (2019) proposed that this family belongs to an
informal clade named “Sclerotiniales”. However, this placement was without support (D.
Haelewaters & N. Wijayawardene).

Saccharomycetes O.E. Erikss. & Winka.


In the case of Saccharomycetes yeasts, the status of several families and the status and
familial placement of several genera has not been unequivocally cleared (Daniel et al. 2014,
Hittinger et al. 2015, Shen et al. 2016, 2018, Kurtzman & Boekhout 2017) since the publication of
the 5th edition of The Yeasts: A Taxonomic Study (Kurtzman 2011). Based on accumulating
phylogenomic data, the status of some families are expected to change substantially (e.g. Shen et
al. 2018). Thus, a comprehensive list of currently accepted sexual genera and non-synonymized
asexual genera (that are expected to be retained following the Melbourne code, notably Candida
Berkhout) are listed among the notes with additional notes on expected changes and protected
names, following Kurtzman & Boekhout (2017) (W.P. Pfliegler & E. Horváth).

Saccisporonites Kalgutkar & Janson. (fossil)


Kalgutkar & Jansonius (2000) opined that Lacrimasporonites stoughiae Elsik is a misfit in
Lacrimasporonites Elsik and therefore they proposed Saccisporonites to accommodate it (R.K.
Saxena).

Sakireeta Subram. & K. Ramakr.


Based on ITS and LSU sequence phylogeny, Crous et al. (2015a) transferred Sakireeta to
Botryosphaeriaceae (A.J.L. Phillips).

Saprochaete Coker & Shanor ex D.T.S. Wagner & Dawes


Expected transfer of species to Magnusiomyces Zender (Kurtzman & Boekhout 2017) (W.P.
Pfliegler and E. Horváth).

Sarcoleotia S. Ito & S. Imai


Phylogeny in Hustad et al. (2011) demonstrates that Sarcoleotia is sister species to
Nothomitra, both in a separate clade than the rest of Geoglossomycetes (V.P. Hustad).

Sarcopeziza Loizides et al.


This genus was recently introduced by Agnello et al. (2018) (P. Alvarado).

Sardiniella Linaldeddu et al.


This genus was introduced by Linaldeddu et al. (2016) to accommodate a diplodia-like
species from diseased Celtis africana trees in Sardinia. Morphologically similar to Diplodia and
Dothiorella, but phylogenetically distinct (A.J.L. Phillips).

Saxophila Selbmann & de Hoog

1419
This is a monotypic genus, typified by S. tyrrhenica, a dematiaceous microcolonial fungus
obtained from marble and limestone in Europe. Its placement in Extremaceae is supported by
multilocus DNA sequence data (Isola et al. 2016) (H. Madrid).

Schadonia Körb.
Kistenich et al. (2018) transferred this genus from Ramalinaceae to Pilocarpaceae (E.
Timdal).

Sclerencoelia Pärtel & Baral


This genus was introduced by Pärtel et al. (2017) to accommodate two species of Encoelia
(E. fascicularis and E. pruinosa) that belonged to Sclerotiniaceae, whereas the type species E.
furfuraceae was placed in Cenangiaceae. A third species of Sclerencoelia was also described by
Pärtel et al. (2017) mostly based molecular data (D. Haelewaters).

Sclerococcum Fr.
Réblová et al. (2016) transferred it to Sclerococcaceae and this was supported by Yu et al.
(2018). However, Sclerococcum has been transferred to Dactylosporaceae Bellem. & Hafellner [=
Sclerococcaceae Réblová, Unter. & W. Gams] by Diederich et al. (2018) (H. Zhang & J. Etayo).

Scolecolachnum Guatimosim et al.


Scolecolachnum was introduced in Hyaloscyphaceae (Guatimosim et al. 2016). Based on
both ITS and LSU phylogenetic analyses, it is retrieved as sister to Hyphodiscus in
Hyaloscyphaceae Han Clade 4 (sensu Han et al. 2014, Johnston et al. 2019) (D. Haelewaters).

Scutula Tul.
Kistenich et al. (2018) transferred this genus from Pilocarpaceae to Ramalinaceae (E.
Timdal).

Seltsamia Jaklitsch & Voglmayr


Jaklitsch et al. (2018) proposed this new genus in Cucurbitariaceae based on morphological
and molecular characters (S. Fryar).

Septomelanconiella Samarak. & K.D. Hyde


Phookamsak et al. (2019) introduced a monotypic genus Septomelanconiella to
accommodate a single species S. thailandica Samarak. & K.D. Hyde. Septomelanconiella can be
distinguished from Melanconiella Sacc. in having finely verrucose brown mature conidia.
Phylogenetic analysis revealed that the genus formed a distinct lineage with other genera in
Melanconiellaceae (R. Phookamsak).

Septoriella Oudem.
Crous et al. (2015c) treated Wojnowicia as a synonym of Septoriella based on a neotypic
study of Wojnowicia hirta Sacc. (R. Phookamsak).

Septorioides Quaedvl. et al.


This genus was introduced for species morphologically similar to Septoria but
distinguishable on account of conidiomata that open by an irregular split, and paraphyses
intermingled with the conidiogenous cells. Furthermore, they constitute a phylogenetic lineage in
Botryosphaeriaceae and thus separate from Septoria and allied genera (Quaedvlieg et al. 2013).
Wyka & Broders (2016) introduced the family Septorioideaceae in Botryosphaeriales to
accommodate Septorioides species. Phillips et al. (2019) took into account phylogeny (ITS, LSU),
morphology and evolutionary divergence times and concluded that Septorioides resides within

1420
Saccharataceae. For this reason, Phillips et al. (2019) regarded Septorioideaceae as a synonym of
Saccharataceae (A.J.L. Phillips).

Sepultariella Van Vooren et al.


This genus was erected to accommodate two species previously ascribed to Leucoscypha. It
represents a separate phylogenetic lineage within Pyrenomataceae (Van Vooren et al. 2017) (I.
Kušan & N. Matočec).

Sheathospora X.L. Fan


Fan et al. (2018b) proposed this new genus based on unique pycnidial conidiomata and
conidia with distinct hyaline sheath in Melanconiellaceae. Sheathospora cornuta is the type to
accommodate Melanconiella cornuta and currently so far known from Cornus controversa and
Juglans regia in China (X.L. Fan).

Snippocia Ertz et al.


Ertz et al. (2018) introduced this genus and placed it in Arthoniaceae based on phylogenetic
analyses (N. Wijayawardene).

Solanella Vaňha
Ekanayaka et al. (2018) proposed that this genus should be transferred to Pezizomycotina
incertae sedis (I. Kušan & N. Matočec).

Solicorynespora R.F. Castañeda & W.B. Kendr.


Hernández-Restrepo et al. (2014) showed that Solicorynespora insolita has a high affinity
with members of Dothideomycetes, and more specifically with Astrosphaeriella livistonicola (J.
Ma)

Spadicoides S. Hughes
Réblová et al. (2018) accommodated this genus in Xenospadicoidaceae based on
phylogenetic analyses (S. Fryar).

Spegazzinia Sacc.
This genus was shown to belong in Didymosphaeriaceae by Tanaka et al. (2015) (P.
Alvarado).

Sphaerosporium Schwein.
According to a multilocus phylogenetic study in Song et al. (2019), the generic type, S.
lignatile Schwein., belongs to Pyronemataceae (Pezizomycetes, Pezizales), whereas S. equinum
(Desm.) J.L. Crane & Schokn. was placed among Onygenales based on LSU sequence data
(Rokas et al. 2012) (D. Haelewaters).

Sphaerosporium Schwein.
According to a multilocus phylogenetic study by Haelewaters et al. in Song et al. (2019), the
generic type, S. lignatile Schwein., belongs to Pyronemataceae (Pezizomycetes, Pezizales),
whereas S. equinum (Desm.) J.L. Crane & Schokn. was placed among Onygenales based on LSU
sequence data (Rokas et al. 2012) (D. Haelewaters).

Spinosporonites R.K. Saxena & S. Khare (fossil)


This monotypic genus includes circular to subcircular, inaperturate, multicellular spores,
each cell giving rise to a robustly built spine. They readily resemble the setose pycnidia found in
some Coelomycetes (R.K. Saxena).

1421
Spiromastigaceae Hirooka et al.
The family was first invalidly published in Rizzo et al (2014) (Arts 38.11 and 42.1) and later
validated in Hirooka et al. (2015) (K. Bensch).

Spirotremesporites Dueñas (fossil)


This genus includes ellipsoidal to elongate, aseptate, psilate fungal spores having aperture in
the form of a single furrow at an angle to the axis of the spore, straight or curved to S-shaped or
sigmoidal in outline, or spiral around the spore axis. Varisulcosporites Rouse & Mustard is a
junior taxonomic synonym of Spirotremesporites (R.K. Saxena).

Sporacestra A. Massal.
Kistenich et al. (2018) resurrected this genus in Ramalinaceae (E. Timdal).

Sporastatiales Lumbsch & Leavitt


Kraichak et al. (2018a) raised Sporastatiaceae to ordinal level as Sporastatiales (N.
Wijayawardene).

Sporocadaceae Corda
Liu et al. (2019) provided a revision of the Sporocadaceae based on multi-locus
phylogenetic analyses, using LSU, ITS, tef-1α, tub2 and rpb2 loci, in combination with
morphological data. A total of 30 well-supported monophyletic clades were recognized,
representing 23 known and seven new genera. Typifications for the type species Diploceras,
Discosia, Monochaetia, Sporocadus and Truncatella) and emendations of various genera and
species were also provided (M. Stadler).

Sporocarpon Will. (fossil)


Dubiocarpon S.A. Hutch. and Oidospora Will. are later taxonomic synonyms of
Sporocarpon Will. (R.K. Saxena).

Sporormiella Ellis & Everh.


The ostiolate Sporormiella has been recognized as a probable synonym of the earlier non-
ostiolate Preussia for several decades and, based on morphology and phylogeny, Zhang et al.
(2012) and Hyde et al. (2013) adopted Preussia. However, the type species S. nigropurpurea has
not been sequenced, and Sporormiella is widely used in the literature of coprophilous fungi (e.g.
Doveri 2004, Bell 2005) and palaeoecology (Raper & Bush 2009, Raczka et al. 2016) and
contains many more species. If the genera are eventually proved to be congeneric molecularly, we
consider that Sporormiella should be proposed for conservation over Preussia so both names are
currently retained here (D.L. Hawksworth & N.N. Wijayawardene).

Staphlosporonites Sheffy & Dilcher (fossil)


Transeptaesporites V.S. Ediger is a later taxonomic synonym of Staphlosporonites (R.K.
Saxena).

Steinera Zahlbr.
The genus, previously placed in Koerberiaceae, has been recently moved into Arctomiaceae
(Ertz et al. 2017a). A new genus, Henssenia, was established for Steinera species remaining in
Koerberiaceae (Ertz et al. 2017a) (M. Kukwa).

Stemphylium Wallr.
The major part of the family Pleosporaceae is represented by species of Pleospora, a genus
that is considered paraphyletic (Kodsueb et al. 2006, Inderbitzin et al. 2009). The type species of
Pleospora, P. herbarum, was synonymized with Stemphylium herbarum. At this time, however,

1422
several hundreds of Pleospora epithets still have not been assigned to Stemphylium or other
genera and are not included in this outline (P.B. Gannibal).

Stereophlebia Zmitr.
Zmitrovich (2018) segregated this new monotypic genus from Lilaceophlebia (Parmasto)
Spirin & Zmitr. to accommodate Phlebia tuberculata (Berk. & M.A. Curtis) Ţura, Zmitr., Wasser
& Spirinski (V. Papp).

Stictographaceae D.Q. Dai & K.D. Hyde


Dai et al. (2018) introduced this family in Asterinales to accommodate Stictographa Mudd,
Karschia Körb., Labrocarpon Etayo & Pérez-Ortega and Melaspileopsis (Müll. Arg.) Ertz &
Diederich (N. Wijayawardene).

Stigmatodiscus Voglmayr & Jaklitsch


Voglmayr & Pintos (2018) synonymised Asterodiscus Voglmayr et al. with Stigmatodiscus
(P. Alvarado).

Striadiporites C.P. Varma & Rawat (fossil)


This genus includes unicellular, diporate fungal spores with striated spore wall.
Stridiporosporites Ke & Shi is a junior taxonomic synonym of Striadiporites (R.K. Saxena).

Symbiotaphrina Kühlw. & Jurzitza ex W. Gams & Arx


Baral et al. (2017) validated the order Symbiotaphrinales and introduced the new family
Symbiotaphrinaceae (K. Bensch).

Symmetrospora Q.M. Wang et al.


Symmetrospora was recently introduced for species previously placed in the asexual genera
Sporobolomyces and Rhodotorula in the “gracilis/marina clade” of Cystobasidiomycetes (Wang et
al. 2015b). Haelewaters et al. (2020) recently proposed three new species and a new combination,
making 10 recognized species (D. Haelewaters).

Synnemasporella X.L. Fan & J.D.P. Bezerra


Synnemasporella was introduced by Fan et al. (2018a) to accommodate fungi with
synnematous conidiomata. This genus is typified by Synnemasporella toxicodendri (S.S.N.
Maharachchikumbura).

Synnemasporellaceae X.L. Fan & J.D.P. Bezerra


Fan et al. (2018a) proposed this new family to accommodate one new genus,
Synnemasporella (Type species: Synnemasporella toxicodendri). The new genus and species have
been collected from Toxicodendron sylvestre in China, and S. aculeans was transferred from
Cryptodiaporthe aculeans (basionym: Sphaeria aculeans) (S. Fryar & S. Somrithipol).

Szczepkamyces Zmitr.
Zmitrovich (2018) introduced this new monotypic genus in Polyporaceae to accommodate
Dichomitus campestris (Quél.) Domański & Orlicz (V. Papp).

Taeniolella S. Hughes
Heuchert et al. (2018) showed that the genus is polyphyletic with type species belonging
Kirschsteiniotheliaceae (Dothideomycetes) while saprobic species cluster within Sordariomycetes
in different families. Lichenicolous species form a monophyletic clade within Asterotexiales,
Dothideomycetes but many species are still not sequenced (A. Suija).

1423
Taitaia Suija et al.
Suija et al. (2018) introduced this lichenicolous genus and confirmed its placement in
Gomphillaceae, Graphidales (N. Wijayawardene).

Tamsiniella S.W. Wong et al.


Phookamsak et al. (2019) treated Tamsiniella in Phyllachoraceae based on phylogenetic
analysis (R. Phookamsak).

Tapesia (Pers.) Fuckel


Tapesia is considered a synonym of Mollisia (Hawksworth & David 1989) but many
species are still classified under Tapesia (Species Fungorum 2020, Tanney & Seifert 2020). As a
result, Tapesia is included in this outline (D. Haelewaters).

Tasmidella Kantvilas et al.


Kistenich et al. (2018) transferred this genus from Ramalinaceae to the Lecanorales
incertae sedis (E. Timdal).

Thalloidima A. Massal.
Kistenich et al. (2018) resurrected this genus in the Ramalinaceae (E. Timdal).

Thecotheus Boud.
Placed in Ascobolaceae, Pezizales. This genus was treated by Kušan et al. (2015) who listed
23 known species, including the newly described T. platyapiculatus (I. Kušan & N. Matočec).

Tenuitholiascaceae S.H. Jiang et al.


Jiang et al. (2020) introduced this family based on the new genus Tenuitholiascus, which
resides in Strigulales (N. Wijayawardene).

Thysanorea Arzanlou et al.


According to phylogenetic studies by Arzanlou et al. (2007) and Dong et al. (2018), this
genus is a member of Herpotrichiellaceae. Dong et al. (2018) introduced the second species
Thysanorea aquatica W. Dong, H. Zhang & K.D. Hyde. However, this species has been reported
as a synonym with the type species Thysanorea papuana (Aptroot) Arzanlou, W. Gams & Crous
(Wang et al. 2018) (H. Madrid & H. Zhang).

Toninia A. Massal.
Kistenich et al. (2018) placed Arthrosporum A. Massal. as a synonym of Toninia (E.
Timdal)

Torrentispora K.D. Hyde et al.


Réblová et al. (2018) transferred Torrentispora from the Annulatascaceae to
Xenospadicoidaceae based on multi-gene phylogenetic analyses (S. Fryar).

Tremellochaete Raitv.
Tremellochaete was reinstated in Auriculariaceae by Malysheva & Spirin (2017) based on
morphological characteristics and phylogenetic analyses. Phookamsak et al. (2019) updated a
species number in this genus. Based on morphological characteristics and phylogenetic analysis,
three species are accommodated in this genus (Malysheva & Spirin 2017, Index Fungorum 2019,
Phookamsak et al. 2019) (R. Phookamsak).

Triadelphia Shearer & J.L. Crane

1424
Recent studies suggested that Triadelphia is polyphyletic, but no DNA sequence data is
available for many of its members. The type species, T. heterospora, belongs in Microascales
(Crous et al. 2015b) (H. Madrid).

Triblidiaceae Rehm
Triblidiaceae is considered a monophyletic family within Rhytismatales, including two
genera, Huangshania and Triblidium. The previous order Triblidiales is synonymized under
Rhytismatales (Karakehian et al. 2019) (D. Haelewaters).

Tribolites W.H. Bradley (fossil)


The fossil conidia resemble conidia of extant genera Tetrachaetum Ingold and Lemonniera
D. Wild. Trihyphaecites Peppers is a junior taxonomic synonym of Tribolites (R.K. Saxena).

Trichomonascaceae Kurtzman & Robnett


Family status expected to change upon resolving Blastobotrys/Trichomonascus (W.P.
Pfliegler and E. Horváth).

Trichomonascus H.S. Jackson emend. Kurtzman & Robnett


Expected transfer of species to Blastobotrys Klopotek to comply with the Melbourne Code
(Kurtzman & Boekhout 2017) (W.P. Pfliegler & E. Horváth).

Trichothyrites Rosend. (fossil)


Notothyrites Cookson and Sphaerialites Venkatach. & R.K. Kar are later taxonomic
synonyms of Trichothyrites (Kalgutkar & Jansonius 2000) (R.K. Saxena).

Trihyphites Kalgutkar & Janson. (fossil)


Trihyphaecites fractus Z.C. Song & Liu Cao, in Song et al. (1989) belongs to Trihyphites.
(R.K. Saxena).

Triporicellaesporites Ke & Shi (fossil)


The spores of two species of Triporicellaesporites, viz. T. elongatus P. Ke & Z.Y. Shi and T.
simplex (Elsik & Janson.) Kalgutkar & Janson., are very similar to spores of the extant
Ceratosporella bicornis (Morgan) Höhnel. (R.K. Saxena).

Trullella Zmitr.
Zmitrovich (2018) treated that the name Trulla Miettinen & Ryvarden is illegitimate (non
Trulla T.M. Harris), thus the new genus Trullella was proposed to accommodate Trulla dentipora
(Ryvarden & Iturr.) Miettinen & Ryvarden and five other Trulla species (V. Papp).

Tubakiaceae U. Braun et al.


Braun et al. (2018) introduced Tubakiaceae (in Diaporthales) to accommodate Tubakia B.
Sutton. and six other genera (viz. Apiognomonioides U. Braun, J.Z. Groenew. & Crous,
Involutscutellula U. Braun & C. Nakash., Paratubakia U. Braun & C. Nakash., Racheliella Crous
& U. Braun, Saprothyrium U. Braun, Crous & J.Z. Groenew., Sphaerosporithyrium U. Braun,
Crous, O. Moreno-Rico & Marm.) (N. Wijayawardene).

Tylothallia P. James & H. Kilias


Kistenich et al. (2018) transferred this genus from Lecanoraceae to Ramalinaceae (E.
Timdal).

Umbilicaria Hoffm.

1425
Davydov et al. (2017) revised the Umbilicariaceae sensu stricto and accepted eight
subgenera in the genus Umbilicaria: viz. Actinogyra (type: U. muehlenbergii), subg. Agyrophora
(type: A. atropruinosa), subg. Floccularia subg. nov. (type: U. deusta), subg. Gyrophora (type: U.
vellea), subg. Iwatakia subg. nov. (type: U. esculenta), subg. Lasallia (type: L. pustulata), subg.
Umbilicaria (type: U. hyperborea), and subg. Umbilicariopsis subg. nov. (type: Umbilicaria
polyrhiza) (G. Rambold).

Umthunziomyces Crous & M.J. Wingf.


Umthunziomyces was introduced for a septoria-like species that resides in
Planistromellaceae (Crous et al. 2016) (A.J.L. Phillips).

Uncinulites Pampal. (fossil)


Graamspora Sal.-Cheb. & Locq. is a later taxonomic synonym of Uncinulites (R.K.
Saxena).

Unguicularia Hohn.
Previously considered as a member of Hyaloscyphaceae (Baral 2016), but currently placed
in Helotiales genera incertae sedis based on the ITS placement by Johnston et al. (2019).
Additional sequence data are needed to resolve the placement of this genus (D. Haelewaters).

Varicellaria Nyl.
In Kraichak et al. (2018b), the monotypic family Varicellariaceae was validated (K.
Bensch).

Varicosporellopsis Lechat & J. Fourn.


Lechat & Fournier (2016) described Varicosporellopsis and placed it in Nectriaceae (I.
Kušan & N. Matočec).

Varmasporites Kalgutkar & Janson. (fossil)


Kalgutkar & Jansonius (2000) opined that Fusiformisporites tonakkalensis Y.N.R. Varma &
R.S. Patil is a misfit in Fusiformisporites Rouse and therefore they proposed Varmasporites to
accommodate it (R.K. Saxena).

Velebitea I. Kušan et al.


Velebitea was introduced in Phookamsak et al. (2019) to accommodate a single species,
Velebitea chrysotexta I. Kušan, Matočec & Jadan in Lachnaceae based on molecular data. The
genus was collected from decorticated branches and stump base of Fagus sylvatica L. (Fagaceae)
in Croatia and is characterized by having apothecial ascomata, elongated cylindrical-deltoid asci,
protruding above paraphyses at maturity, with in Lugol’s solution apical apparatus moderately
euamyloid, of Calycina-type and hyaline, elongated fusoid ascospores (R. Phookamsak).

Vermiconidia Egidi & Onofri


A multilocus phylogenetic study by Isola et al. (2016) placed all members of this genus in
Extremaceae (H. Madrid).

Vitreoporus Zmitr.
Zmitrovich (2018) introduced this new genus to accommodate Gloeoporus dichrous (Fr.)
Bres., G. africanus P.E. Jung & Y.W. Lim, G. citrinoalbus Yuan Yuan & Jia J. Chen, and G.
orientalis P.E. Jung & Y.W. Lim. However, phylogenetic analyses by Jung et al. (2018) revealed
that these species belong to a monophyletic clade in Gloeoporus sensu stricto (V. Papp).

Vittaliana Devadatha et al.

1426
Devadatha et al. (2019) introduced this genus and showed that it is a member of
Phaeosphaeriaceae. Its ascospores are similar to Acericola and Vagicola (S. Tibpromma).

Wheelerophlyctis P.M Letcher et al.


Letcher et al. (2018) introduced this genus, which comprises two species. Phylogenetic
analyses confirmed its placement in Asterophlyctaceae (P. Letcher).

Xanthonectria Lechat et al.


Lechat et al. (2016b) described the monotypic genus Xanthonectria to accommodate Nectria
pseudopeziza within Bionectriaceae (I. Kušan & N. Matočec).

Xenodactylaria Crous
Crous et al. (2018) introduced this genus and showed that it has a distinct lineage in
Myrmecridiales. As a result, the family Myrmecridiaceae was introduced (N. Wijayawardene).

Xenospadicoidaceae Hern.-Restr et al.


Réblová et al. (2018) accepted Calyptosphaeria, Lentomitella, Spadicoides and
Torrentispora as members of the Xenospadicoidaceae, Xenospadicoidales. Furthermore, Réblová
et al. (2018) reduced Xenospadicoides and Pseudodiplococcium under Spadicoides and
synonymised Lentomitellaceae with Xenospadicoidaceae based on a multi-gene phylogeny (J. Ma
& S. Fryar).

Xyladictyochaetaceae Crous & Hern.-Restr


Crous et al. (2018) introduced this family to accommodate the genus Xyladictyochaeta
within the Xylariales (S. Fryar & K. Bensch).

Xylolentia Réblová
Xylolentia is a newly introduced genus in the family Rhamphoriaceae, with the type species
Xylolentia brunneola (Réblová & Štěpánek 2018) (S.S.N. Maharachchikumbura).

Xylomyces Goos et al.


Nine species are currently reported within Xylomyces, but Xylomyces chlamydosporus Goos,
R.D. Brooks & Lamore is the only species phylogenetically related to the Jahnulales (H. Raja).

Zopfochytrium M.J. Powell et al.


Powell et al. (2018) introduced this genus and confirmed its placement in Chytridiaceae (P.
Letcher).

Zymochalara Guatimosim et al.


Zymochalara was introduced in Helotiales genera incertae sedis (Guatimosim et al. 2016)
but in the ITS tree of Johnston et al. (2019) it was retrieved within Pezizellaceae (D.
Haelewaters).

Acknowledgements
Nalin N. Wijayawardene thanks Mushroom Research Foundation and National Science
Foundation of China (No. NSFC 31950410558) for financially supporting this project. Kevin D.
Hyde acknowledges the Foreign Experts Bureau of Yunnan Province, Foreign Talents Program
(2018; grant no. YNZ2018002), Thailand Research grants entitled Biodiversity, phylogeny and
role of fungal endophytes on above parts of Rhizophora apiculata and Nypa fruticans (grant no:
RSA5980068), the future of specialist fungi in a changing climate: baseline data for generalist and
specialist fungi associated with ants, Rhododendron species and Dracaena species (grant no:
DBG6080013), Impact of climate change on fungal diversity and biogeography in the Greater

1427
Mekong Subregion (grant no: RDG6130001). H.T. Lumbsch thanks support by the Grainger
Bioinformatics Center. E. Malosso is grateful to CAPES for financial support (grant no.
88881.062172/2014-01). B.T. Goto, G.A. Silva and K. Jobim, L.C. Maia acknowledges CNPq
(Brazilian Scientific Council, grants no. 465.420/2014-1, 307.129/2015-2 and 408011/2016-5) and
CAPES for support. The study was partially supported by the National Science Centre, Poland,
under Grants No. 2015/17/D/NZ8/00778 and 2017/25/B/NZ8/00473 to Julia Pawłowska. The
research of Martin Kukwa received support from the National Science Centre (NCN) in Poland
(project no 2015/17/B/NZ8/02441). Alan J.L. Phillips acknowledges the support from
UID/MULTI/04046/2019 Research Unit grant from FCT, Portugal to BioISI. H. Zhang is
financially supported by the National Natural Science Foundation of China (Project ID: NSF
31500017). S. Boonmee would like to thank the Thailand Research Fund (Project No.
TRG6180001). Dong-Qin Dai and Li-Zhou Tang would like to thank the National Natural Science
Foundation of China (No. NSFC 31760013, NSFC 31260087, NSFC 31460561), the Scientific
Research Foundation of Yunnan Provincial Department of Education (2017ZZX186) and the
Thousand Talents Plan, Youth Project of Yunnan Provinces for support. R. Phookamsak, M.
Doilom, D. N. Wanasinghe, S.C. Karunarathna and J.C. Xu express sincere appreciations to Key
Research Program of Frontier Sciences of the Chinese Academy of Sciences (grant no. QYZDY-
SSW-SMC014) for research financial support. R. Phookamsak thanks the Yunnan Provincial
Department of Human Resources and Social Security (grant no. Y836181261), Chiang Mai
University and National Science Foundation of China (NSFC) project code 31850410489 for
research financial support. S.C. Kaunarathna thanks CAS President’s International Fellowship
Initiative (PIFI) for funding his postdoctoral research (No. 2018PC0006) and the National Science
Foundation of China (NSFC) for funding this work under the project code 31851110759. S.
Tibpromma would like to thank the International Postdoctoral Exchange Fellowship Program
(number Y9180822S1), CAS President’s International Fellowship Initiative (PIFI) (number
2020PC0009), China Postdoctoral Science Foundation and the Yunnan Human Resources, and
Social Security Department Foundation for funding her postdoctoral research. Yuri S. Tokarev,
Elena S. Nassonova and Irma V. Issi are indebtful to Yuliya Y. Sokolova (Institute of Cytology
RAS, St. Petersbug, Russia) and Anastasia V. Simakova (Tomsk State University, Tomsk, Russia)
for kind permission of reproduction of electron microscopy images of Metchnikovella incurvata
and Crepidulospora beklemishevi, respectively. Yuri S. Tokarev and Irma V. Issi thank Russian
Foundation of Basic Research, grant number 17-04-00871 (taxonomy of Rozellomycota). Elena S.
Nassonova thank Russian Foundation of Basic Research, grant number 18-04-01359 (early
evolution of Microsporidia, phylogeny of Metchnikovellida). Adam Flakus and Pamela
Rodriguez-Flakus are greatly indebted to all staff of the Herbario Nacional de Bolivia, Instituto de
Ecología, Universidad Mayor de San Andrés, La Paz and the SERNAP (http://sernap.gob.bo), for
their generous cooperation providing permits, assistance and facilities support for scientific
studies. The research of AF and PRF were financially supported by the National Science Centre
(NCN) in Poland (DEC-2013/11/D/NZ8/03274). Adam Flakus and Pamela Rodriguez-Flakus
received additional support under statutory funds from the W. Szafer Institute of Botany, Polish
Academy of Sciences, Krakow, Poland. The authors would like to thank Yunnan Innovation
Platform for Development and Utilization of Symbiotic Fungi Resources for finance support. Li-
Fang Zhang would like to thank grant-in-aid from Science and Technology Department of
Yunnan Province (2018FD080) for finance support. Chun-Ying Deng thanks the Biodiversity
Survey and Assessment Project of the Ministry of Ecology and Environment, China
(2019HJ2096001006). Yingqian Kang would like to thank Guizhou Scientific Plan Project [(2019)
2873]; Excellent Youth Talent Training Project of Guizhou Province [(2017) 5639]; Guiyang
Science and Technology Project [(2017) No. 5-19]; Talent Base Project of Guizhou Province,
China [FCJD2018-22]; Research Fund of Education Bureau of Guizhou Province, China [(2018)
481]. D. N. Wanasinghe would like to thank the CAS President’s International Fellowship
Initiative (PIFI) for funding his postdoctoral research (number 2019PC0008), the National Science
Foundation of China and the Chinese Academy of Sciences for financial support under the

1428
following grants: 41761144055, 41771063 and Y4ZK111B01. Yuri K. Novozhilov and Oleg N.
Shchepin acknowledge support from the Russian Foundation of Basic Research, project 18-04-
01232 А. Ivana Kušan, Neven Matočec, Armin Mešić and Zdenko Tkalčec are grateful to
Croatian Science Foundation for their financial support under the project grant HRZZ-IP-2018-
01-1736 (ForFungiDNA). K. Tanaka would like to thank the Japan Society for the Promotion of
Science (JSPS 19K06802)

References

Abdel-Wahab MA, El-Samawaty AE, El Gorban AM, Yassin MA et al. 2018 – Khaleijomyces
marinus gen. et sp. nov. (Juncigenaceae, Torpedosporales) a new lignicolous marine fungus
from Saudi Arabia. Phytotaxa 340, 277–285.
Abdullah SK, Gené J, Guarro J. 2005 – A synopsis of the aero–aquatic genus Pseudaegerita and
description of two new species. Mycological Research 109, 590–594.
Accioly T, Sousa JO, Moreau P-A, Lécuru C et al. 2019 – Hidden fungal diversity from the
Neotropics: Geastrum hirsutum, G. schweinitzii (Basidiomycota, Geastrales) and their
allies. Plos One 14, e0211388.
Adl SM, Bass D, Lane CE, Lukeš J et al. 2019 – Revisions to the classification nomenclature and
diversity of Eukaryotes. Journal of Eukaryotic Microbiology 66, 4–119.
Agnello C, Alvarado P, Loizides M. 2018 – Sarcopeziza (Pezizceae, Ascomycota), a new
monotypic genus for Inzenga’s old taxon Peziza sicula. Ascomycete.org 10, 177–186.
Ainsworth GC, Sparrow FK, Sussman AS. (Eds.) 1973 – The fungi, an advanced treatise. I–IVB;
Academic Press, New York, 3416 p.
Alvarado P, Healy RA, Moreno G, Cabero J et al. 2018 – Phylogenetic studies in Genabea,
Myrmecocystis, and related genera. Mycologia 110, 401–418.
Álvarez E, Cano J, Stchigel AM, Sutton DA et al. 2011 – Two new species of Mucor from clinical
samples. Medical Mycology 49, 62–72.
Alves MH, Trufem SFB, Milanez AI. 2002 – Táxons de Mucor Fresen. (Zygomycota) em fezes de
herbívoros, Recife, PE, Brasil. Revista Brasileira de Biologia 25, 147–160.
Aptroot A, Lücking R. 2016 – A revisionary synopsis of the Trypetheliaceae (Ascomycota:
Trypetheliales). Lichenologist 48, 763–982.
Aptroot A, Maphangwa KW, Zedda L, Tekere M et al. 2019 – The phylogenetic position of
Culbersonia is in the Caliciaceae (lichenized ascomycetes). Lichenologist 51, 187–191.
Aptroot A, Sparrius LB, Alvarado P. 2018 – Aquacidia, a new genus to accommodate a group of
skiophilous temperate Bacidia species that belong in the Pilocarpaceae (lichenized
ascomycetes). Gorteria 40, 11–14.
Aptroot A. 1991 – Monograph of the Pyrenulaceae (excluding Anthracothecium and Pyrenula)
and the Requienellaceae. J. Cramer.
Aptroot A. 1995 – A monograph of Didymosphaeria. Studies in Mycology 37, 1–16.
Ariyawansa HA, Phookamsak R, Tibpromma S, Kang JC et al. 2014 – A molecular and
morphological reassessment of Diademaceae. The Scientific World Journal. Article ID
675348, http://dx.doi.org/10.1155/2014/675348
Arzanlou M, Groenewald JZ, Gams W, Braun U et al. 2007 – Phylogenetic and morphotaxonomic
revision of Ramichloridium and allied genera. Studies in Mycology 58, 57–93.
Bader JA, Shotts EB Jr, Steffens WL, Lom J. 1998 – Occurrence of Loma cf. salmonae inbrook,
brown and rainbow trout from Buford trout hatchery, Georgia, USA. DisAquat Organ 34,
211–216.
Balbiani G. 1882 – Sur les microsporidies ou psorospermies des articules. Comptes Rendus de
l’Académie des Sciences 95, 1168–1171.
Baral HO, Polhorský A. 2019 – Chrysodisca peziculoides gen. et sp. nov. from xeric coniferous
bark across Europe. Mycologia Montenegrina 20, 79–98.

1429
Baral HO, Weber E, Marson G, Quijada L. 2017 – A new connection between wood saprobism
and beetle endosymbiosis: the rarely reported saprobic discomycete Tromeropsis is
congeneric with the symbiotic yeast Symbiotaphrina (Symbiotaphrinales, Xylonomycetes)
and two asexual morphs misplaced in Hyphozyma. Mycological Progress 17, 215–254.
Baral HO. 2016 – Inoperculate discomycetes. In: Jaklitsch W, Baral HO, Lücking R, Lumbsch HT,
Frey W (eds) Syllabus of plant families: A. Engler’s syllabus der Pflanzenfamilien part 1/2.
Borntraeger, Stuttgart, 157–205.
Baral HO. 2019 – Pseudolanzia piceetorum gen. et sp. nov. (Rutstroemiaceae, Helotiales) from
fallen Picea abies needles in Mecklenburg-Vorpommern (Germany). Mycologia
Montenegrina 20, 151–166.
Barbosa FR, Gusmão LFP, Castañeda-Ruiz RF. 2017 – Craspedodidymella matogrossensis gen.
& sp. nov. from the Brazilian Amazon rainforest. Mycotaxon 132, 643–647.
Barnett HL. 1956 – Illustrated genera of imperfect fungi. Burgess Publishing Company,
Minneapolis, 218 p.
Barnett HL. 1960 – Illustrated genera of imperfect fungi. Burgess Publishing Company,
Minneapolis, 225 p.
Barr ME. 1983 – The ascomycete connection. Mycologia 75, 1–13.
Barr ME. 1990 – Melanommatales (Loculoascomycetes). North American Flora, Series 2, part 13.
New York Botanical Garden, New York.
Bass D, Czech L, Williams BAP, Berney C et al. 2018 – Clarifying the relationships between
Microsporidia and Cryptomycota. Journal of Eukaryotic Microbiology 65, 773–782.
Beakes GW, Thines M 2017 – Hyphochytriomycota and Oomycota. In: Archibald J, Simpson A,
Slamovits C (eds) Handbook of the Protists, Second Edition. Springer International
Publishing, Cham, 435–505.
Becnel JJ, Andreadis TG. 1999 – Microsporidia in insects. In: Wittner M (ed) The microsporidia
and microsporidiosis. ASM Press, Washington DC, 447–501.
Becnel JJ, Takvorian PM, Cali A. 2014 – Checklist of available generic names for Microsporidia
with type species and type hosts. In: Weiss LM, Becnel JJ (eds) Microsporidia: Pathogens of
Opportunity. Wiley–Blackwell, pp 671–686.
Begerow D, McTaggart A, Agerer R, Frey W. 2018 – Syllabus of Plant Families-A. Engler's
Syllabus der Pflanzenfamilien Part 1/3.
Bell A. 2005 – An illustrated guide to the coprophilous ascomycetes of Australia. CBS
Biodiversity Series 3, 172 pp. Utrecht: Centraalbureau voor Schimmelcultures.
Bennett RM, Thines M 2019 – Revisiting Salisapiliaceae. Fungal Systematics and Evolution, 3,
353–366.
Benny GL, Smith ME, Kirk PM, Tretter ED et al. 2016 – Challenges and future perspectives in
the systematics of Kickxellomycotina, Mortierellomycotina, Mucoromycotina, and
Zoopagomycotina. In: Li D.–W (ed) Biology of Microfungi, Fungal Biology. Switzerland,
Springer International Publishing, pp 65–126.
Bessey EA. 1950 – Morphology and taxonomy of fungi. Blakiston Company, Philadelphia, 791 p.
Beznoussenko GV, Dolgikh VV, Morzhina EV, Semenov PB et al. 2007 – Analogs of the Golgi
complex in microsporidia: structure and avesicular mechanisms of function. Journal of Cell
Science 120, 1288–1298.
Bigliardi E, Sacchi L. 2001 – Cell biology and invasion of the microsporidia. Microbes &
Infection 3, 373 –379.
Blackwell M, Haelewaters D, Pfister DH. 2020 – Laboulbeniomycetes: Evolution, natural history,
and Thaxter’s final word. Mycologia in press. doi:10.1080/00275514.2020.1718442.
Błaszkowski J, Chwat G, Góralska A, Ryszka P et al. 2015 − Two new genera, Dominikia and
Kamienskia, and D. disticha sp. nov. in Glomeromycota. Nova Hedwigia 100, 225–238.
Błaszkowski J, Kozłowska A, Niezgoda P, Goto BT et al 2018a – A new genus, Oehlia with
Oehlia diaphana comb. nov. and an emended description of Rhizoglomus vesiculiferum
comb. nov. in the Glomeromycotina. Nova Hedwigia 107, 501–518.

1430
Błaszkowski J, Niezgoda P, Goto BT, Kozłowska A. 2018b – Halonatospora gen. nov. with H.
pansihalos comb. nov. and Glomus bareae sp. nov. (Glomeromycota; Glomeraceae).
Botany 96, 737–748.
Błaszkowski J, Niezgoda P, Paiva JN, Silva KJG et al. 2019a – Sieverdingia gen. nov., S. tortuosa
comb. nov., and Diversispora peloponnesiaca sp. nov. in the Diversisporaceae
(Glomeromycota). Mycological Progress 18, 1363–1382.
Błaszkowski J, Niezgoda P, Piatek M, Marguno F et al. 2019b – Rhizoglomus dalpae, R. maiae,
and R. silesianum new species. Mycologia 1, 1–15.
Bohne W, Böttcher K, Gross U. 2011 – Theparasitophorous vacuole of Encephalitozooncuniculi:
biogenesis and characteristics of the host cell–pathogen interface. International Journal of
Medical Microbiology 301, 395 –399.
Boise J. 1986 – Requienellaceae, a new family of Loculoascomycetes. Mycologia 78, 37–41.
Bon M. 1986 – Novitates. Validations et taxons nouveaux. Documents Mycologiques 17, 51–56.
Boonmee S, Phookamsak R, Hongsanan S, Doilom M et al. 2017 – Mycosphere notes 51 – 101.
Revision of genera in Perisporiopsidaceae and Pseudoperisporiaceae and other Ascomycota
genera incertae sedis. Mycosphere 8, 1695–1801.
Braun U, Nakashima C, Crous PW, Groenewald JZ et al. 2018 –Phylogeny and taxonomy of the
genus Tubakia s. lat. Fungal Systematics & Evolution 1, 41–99.
Brefeld O. 1869 – Dictyosteliummucoroides. Einneuer Organismusaus der Verwandtschaft der
Myxomyceten. Abhandl Senckenb Naturforsch Ges, 85–107.
Brown M, Silberman M. 2013 – The non–dictyostelidsorocarpic amoebae. Pages 219–242 in
Dictyostelids: Evolution, Genomics and Cell Biology. Romeralo M, Baldauf S, Escalante R
(Eds). Springer–Verlag, Berlin–Heidelberg.
Brown MW, Spiegel FW, Silberman JD. 2009 – Phylogeny of the “Forgotten” cellular slime mold
Fonticula alba reveals a key evolutionary branch within Opisthokonta. Molecualr Biology
& Evolution 26, 2699–2709.
Buaya AT, Thines M 2020 – Diatomophthoraceae–a new family of olpidiopsis-like diatom
parasitoids largely unrelated to Ectrogella. Fungal Systematics & Evolution 5, 113–118.
Buaya AT, Ploch S, Hanic L, Nam B, Nigrelli L, Kraberg A, Thines M 2017 – Phylogeny of
Miracula helgolandica gen. et sp. nov. and Olpidiopsis drebesii sp. nov., two basal
oomycete parasitoids of marine diatoms, with notes on the taxonomy of Ectrogella-like
species. Mycological Progress 16, 1041–1050.
Buaya AT, Ploch S, Inaba S, Thines M 2019 – Holocarpic oomycete parasitoids of red algae are
not Olpidiopsis. Fungal Systematics & Evolution 4, 21–31.
Buyck B, Duhem B, Das K, Jayawardena RS et al. 2017 – Fungal biodiversity profiles 21–30.
Cryptogamie Mycologie 38, 101–146.
Čadež N, Bellora N, Ulloa R, Hittinger CT et al. 2019 – Genomic content of a novel yeast species
Hanseniaspora gamundiae sp. nov. from fungal stromata (Cyttaria) associated with a
unique fermented beverage in Andean Patagonia, Argentina. PLoS One 14, e0210792.
Cai L, Jeewon R, Hyde K. 2006 – Molecular systematics of Zopfiella and allied genera: evidence
from multigene sequence analyses. Mycological Research 110, 359–368.
Cali A, Becnel JJ, Takvorian PM. 2017 – Microsporidia. In: Archibald JM, Simpson AGB,
Slamovits CH (eds) Handbook of the Protists. Springer.pp 1569–1618.
Cali A, Takvorian PM. 2014 – Developmental morphology and life cycles of the Microsporidia.
In: Weiss LM, Becnel JJ (eds) Microsporidia: pathogens of opportunity. Wiley–Blackwell
Press, Ames, 71–133.
Cannon PF, Alcorn JL. 1994 − Omnidemptus affinis gen. et sp. nov., teleomorph of
Mycoleptodiscus affinis sp. nov. Mycotaxon 51, 483–487
Carmarán CC, Berretta M, Martínez S, Barrera B et al. 2015 – Species diversity of Cladorrhinum
in Argentina and description of a new species, Cladorrhinum australe. Mycological
Progress 14, 94.

1431
Castellano MA, Elliott TF, Truong C, Séné O et al. 2016 – Kombocles bakaiana gen. sp. nov.
(Boletaceae), a new sequestrate fungus from Cameroon. IMA fungus 7, 239–245.
Cavalier-Smith T. 1981 – Eukaryote kingdoms: seven or nine? Bio Systems. 14, 461–481.
Cavalier-Smith T. 1983 – A 6–kingdom classification and a unified phylogeny. In: Schenk HEA
and Schwemmler WS (eds) Endocytobiology, W. S. Vol II. Walter de Gruyter & Co, Berlin,
pp. 1027–1034.
Cavalier-Smith T. 2018 – Kingdom Chromista and its eight phyla: a new synthesis emphasising
periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences.
Protoplasma 255, 297–357.
Cavender JC, Raper KB. 1965a – The Acrasieae in nature. I. Isolation. American Journal of
Botany 52, 294–296.
Cavender JC, Raper KB. 1965b – The Acrasieae in nature. II. Forest soil as a primary habitat.
American Journal of Botany 52, 297–302.
Chen CC, Wu SH, Chen CY. 2018 – Hydnophanerochaete and Odontoefibula, two new genera of
phanerochaetoid fungi (Polyporales, Basidiomycota) from East Asia. MycoKeys 39, 75–96.
Chen GQ, Zheng RY 1986 – A new species of Mucor with giant spores. Acta Mycologica Sinica,
Supplement I 1, 56–60.
Choi Y-J, Thines M 2015 – u Host jumps and radiation, not co‐divergence drives diversification
of obligate pathogens. A case study in downy mildews and Asteraceae. PloS ONE 10,
e0133655.
Choudhary MM, Metcalfe MG, Arrambide K, Bern C et al. 2011 – Tubulinosema sp.
microsporidian myositis in immunosuppressed patient. Emerging Infectious Diseases 17,
1727–1730.
Clarke RT 1965 – Fungal spores from Vermejo Formation coal beds (Upper Cretaceous) of
central Colorado. Mountain Geologist 2, 85–93.
Cookson IC 1947 – Fossil fungi from Tertiary deposits in the southern hemisphere. Part I.
Proceedings of the Linnean Society of New South Wales 72, 207–214.
Corazon-Guivin MA, Cerna-Mendoza A, Guerrero-Abad JC, Vallejos-Tapullima A et al. 2019
Microkamienskia gen. nov. and Microkamienskia peruviana, a new arbuscular mycorrhizal
fungus from Western Amazonia. Nova Hedwigia 109, 355–368
Corazon-Guivin MA, Mendoza AC, Guerrero-Abad JC, Vallejos-Tapullima A et al. 2019a –
Funneliglomus, gen. nov., and Funneliglomus sanmartinensis, a new arbuscular mycorrhizal
fungus from the Amazonia region in Peru. Sydowia 71 E-Book/S 17–24.
Corazon-Guivin MA, Mendoza AC, Guerrero-Abad JC, Vallejos-Tapullima A et al. 2019b –
Nanoglomus plukenetiae, a new fungus from Peru, and a key to small-spored Glomeraceae
species, including three new genera in the “Dominikia complex/clades”. Mycological
Progress 18, 1395–1409.
Corsaro D, Michel R, Walochnik J, Venditti D et al. 2016 – Molecular identification of
Nucleophaga terricolae sp. nov. (Rozellomycota), and new insights on the origin of the
Microsporidia. Parasitology Research 115, 3003–3011.
Corsaro D, Walochnik J, Venditti D, Hauröder B et al. 2020 – Solving an old enigma:
Morellospora saccamoebae gen. nov., sp. nov. (Rozellomycota), a Sphaerita-like parasite of
free-living amoebae. Parasitology Research 119, 925–934.
Corsaro D, Walochnik J, Venditti D, Müller KD et al 2014a – Rediscovery of Nucleophaga
amoebae, a novel member of the Rozellomycota. Parasitology Research 113, 4491–4498.
Corsaro D, Walochnik J, Venditti D, Steinmann J et al. 2014b – Microsporidia–like parasites of
amoebae belong to the early fungal lineage Rozellomycota. Parasitology Research 113,
1909–1918.
Corsaro D, Wylezich C, Venditti D, Michel R et al. 2019 – Filling gaps in the microsporidian tree:
rDNA phylogeny of Chytridiopsis typographi (Microsporidia: Chytridiopsida). Parasitology
Research 118, 169–180.
Crivelli PG. 1983 – Über die heterogene Ascomycetengattung Pleospora Rabh. 1–213.

1432
Crivelli PG. 1983 – Ueber die heterogene Ascomycetengattung Pleospora Rabh. Vorschlag für
eine Aufteilung, Zürich. Diss. ETH Nr. 7318. pp 1–215.
Crous PW, Braun U, Schubert K, Groenewald JZ. 2007 – Delimiting Cladosporium from
morphologically similar genera. Studies in Mycology 58, 33–56.
Crous PW, Carris LM, Giraldo A, Groenewald JZ, Hawksworth DL et al. 2015c – The Genera of
Fungi – fixing the application of the type species of generic names – G 2: Allantophomopsis,
Latorua, Macrodiplodiopsis, Macrohilum, Milospium, Protostegia, Pyricularia, Robillarda,
Rotula, Septoriella, Torula, and Wojnowicia. IMA fungus 1, 163–198.
Crous PW, Groenewald JZ 2016 – They seldom occur alone. Fungal Biology 120, 1392–1415.
Crous PW, Müller MM, Sánchez RM, Giordano L et al. 2015a – Resolving Tiarosporella spp.
allied to Botryosphaeriaceae and Phacidiaceae. Phytotaxa. 202, 73–93.
Crous PW, Wingfield MJ, Burgess TI, Carnegie AJ et al. 2017 – Fungal Planet description sheets:
625 –715. Persoonia 39, 270–467.
Crous PW, Wingfield MJ, Burgess TI, Hardy GESt et al. 2016 – Fungal Planet description sheets:
469 –557. Persoonia 37, 218–403.
Crous PW, Wingfield MJ, Burgess TI, Hardy GEStJ, Gené, J, Guarro J et al. 2018 – Fungal Planet
description sheets: 716 – 784. Persoonia 40, 240–393.
Crous PW, Wingfield MJ, Guarro J, Hernández–Restrepo M et al. 2015b – Fungal Planet
description sheets: 320–370. Persoonia 34, 167–266.
Cui BK, Li HJ, Dai YC. 2011b – Wood–rotting fungi in eastern China 6. Two new species of
Antrodia (Basidiomycota) from Mt. Huangshan, Anhui Province. Mycotaxon 116, 13–20.
Cuomo CA, Desjardins CA, Bakowski MA, Goldberg J et al. 2012 – Microsporidian genome
analysis reveals evolutionary strategies for obligate intracellular growth. Genome Research
22, 2478–2488.
da Silva M, Pinho DB, Pereira OL, Fernandes FM, Barreto RW. 2016 –Naming potentially
endangered parasites: foliicolous mycobiota of Dimorphandra wilsonii, a highly threatened
Brazilian tree species. PloS one 11, e0147895.
Dai DQ, Tang LZ, Liu C, Wang HB, Hyde KD. 2018 – Studies on Parmulariaceae I. A
phylogeny based on available sequence data; introducing Parmulariales ord. nov., and
Hemigraphaceae, Melaspileellaceae and Stictographaceae fam. nov. Phytotaxa 369, 63–79.
Daniel HM, Lachance MA, Kurtzman CP. 2014 – On the reclassification of species assigned to
Candida and other anamorphic ascomycetous yeast genera based on phylogenetic
circumscription. Antonie Leeuwenhoek 106, 67–84.
Daranagama DA, Hyde KD, Sir EB, Thambugala KM et al. 2018 – Towards a natural
classification and backbone tree for Graphostromataceae, Hypoxylaceae,
Lopadostomataceae and Xylariaceae. Fungal Diversity 88, 1–65.
Daru BH, Bowman EA, Pfister DH, Arnold AE. 2018 – A novel proof of concept for capturing
the diversity of endophytic fungi preserved in herbarium specimens. Philosophical
Transactions of the Royal Society of London Series B – Biological Sciences 374, 20170395.
Datlof EM, Amend AS, Earl K, Hayward J et al. 2017 – Uncovering unseen fungal diversity from
plant DNA banks. PeerJ 5, e3730.
Davydov EA, Peršoh D, Rambold G. 2017 – Umbilicariaceae (lichenized Ascomycota)–Trait
evolution and a new generic concept. Taxon 66, 1282–1303.
Dayarathne MC, Maharachchikumbura SSN, Jones EBG, Dong W et al. 2019 – Phylogenetic
revision of Savoryellaceae and evidence for its ranking as a subclass. Frontiers in
Microbiology 10, 840.
de Bary A. 1887 – Comparative morphology and biology of the fungi mycetozoa and bacteria
Clarendon Press, Oxford.
de Hoog GS, Guarro J, Gené J, Figueras MJ. 2015 – Atlas of clinical fungi, 3rd web edition.
Utrecht, CBS–Fungal Biodiversity Centre.
De Kesel A, Haelewaters D. 2019 – Laboulbeniales (Fungi, Ascomycota) of cholevine beetles
(Coleoptera, Leiodidae) in Belgium and the Netherlands. Sterbeeckia 35, 60–66.

1433
De Lima CLF, Lima DX, De Souza CAF, De Oliveira RJV et al. 2018 – Description of Mucor
pernambucoensis (Mucorales, Mucoromycota), a new species isolated from the Brazilian
Upland Rainforest. Phytotaxa 350, 274–282.
De Long Q, Liu LL, Zhang X, Wen TC et al. 2019 – Contributions to species of Xylariales in
China-1. Durotheca species. Mycological Progress 18, 495–510.
De Souza CAF, Voig K, Gurgel LS, Cordeiro TRL et al 2018 – A new species of Mucor
(Mucoromycotina, Mucorales) isolated from an enclave of Upland Atlantic Forest in the
semi–arid region of Brazil. Phytotaxa 351, 53–62.
Delbac F, Polonais V. 2008 – The microsporidian polar tube and its role in invasion. Subcellular
Biochemistry 47, 208–220.
Delgado G, Miller AN, Piepenbring M. 2018 – South Florida microfungi: Castanedospora, a new
genus to accommodate Sporidesmium pachyanthicola (Capnodiales, Ascomycota).
Cryptogamie, Mycologie 39, 109–128.
Desirò A, Rimington WR, Jacob A, Vande Pol N et al. 2017 – Multigene phylogeny of
Endogonales, an early diverging lineage of fungi associated with plants. IMA fungus 8,
245–257.
Desjardin D, Binder M, Roekring S, Flegel T. 2009 – Spongiforma, a new genus of gastroid
boletes from Thailand. Fungal Diversity 37, 1–8.
Devadatha B, Mehta N, Wanasinghe DN, Baghela A et al. 2019 – Vittaliana mangrovei
Devadatha, Nikita, A. Baghela & VV Sarma, gen. nov, sp. nov. (Phaeosphaeriaceae), from
mangroves near Pondicherry (India), based on morphology and multigene phylogeny.
Cryptogamie Mycologie 40, 117–132
Dick MW 2001 – Straminipilous Fungi: Systematics of the Peronosporomycetes Including
Accounts of the Marine Straminipilous Protists, the Plasmodiophorids and Similar
Organisms. Dordrecht / Boston / London: Kluwer Academic Publishers.
Didier ES, Stovall ME, Green LC, Brindley PJ et al. 2004 – Epidemiology of microsporidiosis:
sources and modes of transmission. Veterinary Parasitology 126, 145–166.
Diederich P, Lawrey JD, Ertz D. 2018 – The 2018 classification and checklist of lichenicolous
fungi, with 2000 non–lichenized, obligately lichenicolous taxa. Bryologist 121, 340–425.
Dilcher DL. 1965 – Epiphyllous fungi from Eocene deposits in western Tennessee, U.S.A.
Palaeontographica Abt B 116, 1–54.
Dong W, Hyde KD, Bhat DJ, Zhang H. 2018 – Introducing Aculeata aquatica, gen. et sp. nov.,
Minimelanolocus thailandensis sp. nov. and Thysanorea aquatica sp. nov.
(Herpotrichiellaceae, Chaetothyriales) from freshwater in northern Thailand. Mycological
Progress 17, 617–629.
Dougoud R. 2012 – Aeruginoscyphus, un nouveau genre pour Peziza sericea (Helotiales,
Hyaloscyphaceae). Ascomycete.org 4, 1–4.
Doveri F. 2005 – Fungi Fimicoli Italici.1104 pp. Trento: Assozciazione Micoligica Bresadola.
Dukik K, Muñoz JF, Jiang Y, Feng P et al. 2017 – Novel taxa of thermally dimorphic systemic
pathogens in the Ajellomycetaceae (Onygenales). Mycoses 60, 296–309.
Dumont KP. 1975 – Sclerotiniaceae VIII. The generic name Pachydisca, Mycologia 67, 161–167.
Dunn AM, Smith JE. 2001 – Microsporidian life cycles and diversity: the relationship between
virulence and transmission. Microbes and Infection 3, 381–388.
Edwards WN. 1922 – An Eocene microthyriaceous fungus from Mull, Scotland. Transactions of
the British Mycological Society 8, 66–72.
Ekanayaka AH, Hyde KD, Gentekaki E, McKenzie EHC et al. 2019 – Preliminary classification
of Leotiomycetes. Mycosphere 10, 310–489.
Ekanayaka AH, Hyde KD, Jones EBG, Zhao QI. 2018 – Taxonomy and phylogeny of operculate
discomycetes: Pezizomycetes. Fungal Diversity 90, 161–243.
Ellis MB. 1971 – Dematiaceous Hyphomycetes. Commonwealth Mycological Institute, Kew,
England, 608 p.

1434
Elsik WC. 1968 – Palynology of a paleocene Rockdale lignite, Milam County, Texas. I.
Morphology and taxonomy. Pollen Spores 10, 263–314.
Elsik WC. 1978 – Classification and geologic history of the microthyriaceous fungi. Proceedings
of the Fourth International Palynological Conference, Lucknow 1976–77 volume 1, 331–
342.
Elsik WC. 1990 – Hypoxylonites and Spirotremesporites, form genera for Eocene to Pleistocene
fungal spores bearing a single furrow. Palaeontographica Abt. B 216, 137–169.
Eriksson O. 1984 – Outline of the Ascomycetes 1983 – Systema Ascomycetum 3, 1–72.
Ertz D, Poulsen RS, Charrier M, Søchting U. 2017a – Taxonomy and phylogeny of the genus
Steinera (Arctomiales, Arctomiaceae) in the subantarctic islands of Crozet and Kerguelen.
Phytotaxa 324, 201–238.
Ertz D, Sanderson N, Łubek A, Kukwa M. 2018 – Two new species of Arthoniaceae from old–
growth European forests, Arthonia thoriana and Inoderma sorediatum, and a new genus for
Schismatomma niveum. Lichenologist 50, 161–72.
Ertz D, Søchting U, Gadea A, Charrier M et al. 2017b – Ducatina umbilicata gen. et sp. nov., a
remarkable Trapeliaceae from the subantarctic islands in the Indian Ocean. Lichenologist
49, 127–40.
Ethridge Glass DL, Brown DD, Elsik WC. 1986 – Fungal spores from the Upper Eocene Manning
Formation, Jackson Group, east and south–central Texas, U.S.A. Pollen Spores 28, 403–
420.
Fan XD, Bezerra JDP, Tian CM, Crous PW. 2018a – Families and genera of diaporthalean fungi
associated with canker and dieback of tree hosts. Persoonia 40, 119–134.
Fan XD, Du Z, Bezerra JDP, Tian CM. 2018b – Taxonomic circumscription of melanconis-like
fungi causing canker disease in China. Mycokeys 42, 89–124.
Fiore-Donno AM, Clissmann F, Meyer M, Schnittler M et al. 2013 – Two-gene phylogeny of
bright–spored Myxomycetes (slime moulds superorder Lucisporidia). PLoS One 8, e62586.
Fiore-Donno AM, Kamono A, Meyer M, Schnittler M et al. 2012 – 18S rDNA phylogeny of
Lamproderma and allied genera (Stemonitales Myxomycetes Amoebozoa). PLoS One 7,
e35359.
Fiuza PO, Silva CR, Borges Santos TA, Raja H et al. 2018 – Roselymyces, a new asexual genus of
the Xylariales (Ascomycota) from Brazil. Sydowia 70, 59–65.
Fokin SI, Giuseppe GDi, Erra F, Dini F. 2008 – Euplotespora binucleata n. gen., n. sp. (Protozoa:
Microsporidia), a parasite infecting the hypotrichous ciliate Euplotes woodruffi, with
observations on microsporidian infections in Ciliophora. Journal of Eukaryotic
Microbiology 55, 214–228.
Franzen C, Nassonova ES, Scholmerich J, Issi IV. 2006 – Transfer of the members of the genus
Brachiola (Microsporidia) to the genus Anncaliia based on ultrastructural and molecular
data. Journal of Eukaryotic Microbiology 53, 26–35.
Franzen C. 2004 – Microsporidia: how can they invade other cells? Trends in Parasitology 20,
275–279.
Frisch A, Thor G, Moon KH, Ohmura Y. 2018 – Galbinothrix, a new monotypic genus of
Chrysotrichaceae (Arthoniomycetes) lacking pulvinic acid derivatives. Plant & Fungal
Systematics 63, 31–37.
Fryar SC, Haelewaters D, Catcheside DEA. 2019 – Annabella australiensis gen. & sp. nov.
(Helotiales, Cordieritidaceae) from South Australian mangroves. Mycological Progress 18,
973–981.
Fryday AM, Printzen C, Ekman S. 2014 – Bryobilimbia, a new generic name for Lecidea
hypnorum and closely related species. Lichenologist 46, 25–37.
Galindo LJ, Torruella G, Moreira D, Timpano H et al. 2018 – Evolutionary genomics of
Metchnikovella incurvata (Metchnikovellidae): An Early Branching Microsporidium.
Genome Biology & Evolution 10, 2736–2748.

1435
Gams W. 2000 – Phialophora and some similar morphologically little-differentiated anamorphs
of divergent ascomycetes. Studies in Mycology 45, 187–199.
Gannibal PhB, Lawrence DP. 2018 – Distribution of Alternaria species among sections. 6.
Species formerly assigned to genus Ulocladium. Mycotaxon 133, 293-299.
Gannibal PhB. 2018 – Distribution of Alternaria species among sections. 4. Species formerly
assigned to genus Nimbya. Mycotaxon 133, 37–43.
Gerdemann JW, Trappe JM. 1974 – The Endogonaceae of the Pacific Northwest. Micologia
Memoir 5, 1–76
Ghani IA, Dieng H, Abu Hassan ZA, Ramli N et al. 2013 – Pathogenicity of a microsporidium
isolate from the Diamondback moth against Noctuid moths: characterization and
implications formicrobiological pest management. PLoS One 8, e81642.
Gherbawy Y, Kesselboth C, Elhariry H, Hoffmann K. 2010 – Molecular barcoding of microscopic
fungi with emphasis on the mucoralean genera Mucor and Rhizopus. In: Gherbawy Y,
Voight K (eds) Molecular identification of fungi. Springer–Verlag, Berlin Heidelberg, pp
225–265.
Gill EE, Fast NM. 2006 – Assessing the microsporidia–fungi relationship: Combined
phylogenetic analysis of eight genes. Gene 375, 103–109.
Gilman JC. 1959 – A manual of soil fungi. Constable and Company Ltd, London.
Goldmann L, Weir A. 2018 – Molecular phylogeny of the Laboulbeniomycetes (Ascomycota).
Fungal Biology 122, 87–100.
Gonçalves MF, Vicente TF, Esteves AC, Alves A. 2019 – Neptunomyces aureus gen. et sp. nov.
(Didymosphaeriaceae, Pleosporales) isolated from algae in Ria de Aveiro, Portugal.
MycoKeys 60, 31.
Gordillo A, Decock C. 2018 – Myrothecium–like (Ascomycota, Hypocreales) species from
tropical areas: Digitiseta gen. nov. and additions to Inaequalispora and Parvothecium.
Mycological Progress 17, 179–190.
Goto BT, Silva GA, Assis DMA, Silva DK et al. 2012 – Intraornatosporaceae (Gigasporales), a
new family with two new genera and two new species. Mycotaxon 119, 117–132.
Gräfenhan T, Schroers H–J, Nirenberg HI, Seifert KA. 2011 – An overview of the taxonomy,
phylogeny, and typification of nectriaceous fungi in Cosmospora, Acremonium, Fusarium,
Stilbella, and Volutella. Studies in Mycology 68, 79–113.
Grube M, Matzer M, Hafellner J. 1995 – A preliminary account of the lichenicolous Arthonia
species with reddish, K+ reactive pigments. Lichenologist 27, 25–42.
Grünig CR, Duo A, Sieber TN, Holdenrieder O. 2008 – Assignment of species rank to six
reproductively isolated cryptic species of the Phialocephala fortinii s.l.-Acephala applanata
species complex. Mycologia 100, 47–67.
Guatimosim E, Schwartsburd PB, Crous PW, Barreto RW. 2016 – Novel fungi from an ancient
niche: lachnoid and chalara-like fungi on ferns. Mycological Progress 15, 1239–1267.
Gulden G, Stensrud Ø, Shalchian-Tabrizi K, Kauserud H. 2005 – Galerina Earle: A polyphyletic
genus in the consortium of dark-spored agarics. Mycologia 97, 823–837.
Guswenrivo I, Tseng SP, Yang CCS, Yoshimura T. 2018 – Development of multiplex nested PCR
for simultaneous detection of ectoparasitic fungi Laboulbeniopsis termitarius and
Antennopsis gallica on Reticulitermes speratus (Blattodea: Rhinotermitidae). Journal of
Economic Entomology 111, 1330–1336.
Haag KL, James TY, Pombert JF, Larsson R et al. 2014 – Evolution of a morphological novelty
occurred before genome compaction in a lineage of extreme parasites. Proceedings of the
National Academy of Sciences 111, 15480–15485.
Haelewaters D, De Kesel A, Gorczak M, Bao K et al. 2019a – Laboulbeniales (Ascomycota) of
the Boston Harbor Islands II: species parasitizing Carabidae, and the Laboulbenia flagellata
species complex. Northeastern Naturalist 25 (Special Issue 9), 110–149.
Haelewaters D, De Kesel A, Pfister DH. 2018a – Integrative taxonomy reveals hidden species
within a common fungal parasite of ladybirds. Scientific Reports 8, 15966.

1436
Haelewaters D, Page RA, Pfister DH. 2018b – Laboulbeniales hyperparasites (Fungi,
Ascomycota) of bat flies: Independent origins and host associations. Ecology and Evolution
8, 8396–8418.
Haelewaters D, Pfliegler WP, Gorczak M, Pfister DH. 2019b – Birth of an order: comprehensive
molecular phylogenetic study reveals that Herpomyces (Fungi, Laboulbeniomycetes) is not
part of Laboulbeniales. Molecular Phylogenetics & Evolution 133, 286–301.
Haelewaters D, Rossi W. 2017 – Laboulbeniales parasitic on American small carrion beetles: new
species of Corethromyces, Diphymyces, and Rodaucea. Mycologia 109, 655–666.
Haelewaters D, Toome-Heller M, Albu S, Aime MC. 2020 – Red yeasts from leaf surfaces and
other habitats: three new species and a new combination of Symmetrospora
(Pucciniomycotina, Cystobasidiomycetes). Fungal Systematics and Evolution 5, 187–196.
Haelewaters D, Zhao SY, Clusella-Trullas S, Cottrell TE et al. 2017 – Parasites of Harmonia
axyridis: current research and perspectives. Biological Control 62, 355–371.
Haelewaters D, Zhao SY, De Kesel A, Royer IR et al. 2015 – Laboulbeniales (Ascomycota) of the
Boston Harbor Islands I: species parasitizing Coccinellidae and Staphylinidae. Northeastern
Naturalist 22, 459–477.
Hagiwara H. 1989 – The taxonomic study of Japanese dictyostelid cellular slime molds. National
Science Museum, Tokyo.
Han JG, Hosoya T, Sung GH, Shin HD. 2014 – Phylogenetic reassessment of Hyaloscyphaceae
sensu lato (Helotiales, Leotiomycetes) based on multigene analyses. Fungal Biology 118,
150–167.
Han ML, Chen YY, Shen LL, Song J et al. 2016 – Taxonomy and phylogeny of the brown–rot
fungi: Fomitopsis and its related genera. Fungal Diversity 80, 343–373.
Hansen K, Perry BA, Dranginis AW, Pfister DH. 2013 – A phylogeny of the highly diverse cup-
fungus family Pyronemataceae (Pezizomycetes, Ascomycota) clarifies relationships and
evolution of selected life history traits. Molecular Phylogenetics & Evolution 67, 311–335.
Hao YJ, Qin J, Yang ZL. 2014 – Cibaomyces, a new genus of Physalacriaceae from East Asia.
Phytotaxa 162, 198–210.
Hashimoto A, Hirayama K, Takahashi H, Matsumura M et al. 2018 – Resolving the Lophiostoma
bipolare complex: Generic delimitations within Lophiostomataceae. Studies in Mycology
90, 161–189.
Hawksworth DL & Lücking R. 2017 – Fungal diversity revisited: 2.2 to 3.8 million species.
Microbiology Spectrum 5, doi: 10.1128/microbiolspec.FUNK-0052-2016.
Hawksworth DL, Cole MS. 2002 – Intralichen, a new genus for lichenicolous ‘Bispora’ and
‘Trimmatostroma’ species. Fungal Diversity11, 87–97.
Hawksworth DL, Eriksson OE. 1986 – The names of accepted orders of ascomycetes. Systema
Ascomycetum 5, 175–184.
Hawksworth DL, Kirk PM, Sutton BC, Pegler DN, eds. 1983 – Ainsworth & Bisby's Dictionary
of the Fungi. 7th ed. Commonwealth Mycol Institute, Kew.
He M Q, Zhao RL, Hyde KD, Begerow D et al. 2019 – Notes, outline and divergence times of
Basidiomycota. Fungal Diversity 1–263.
Healy R, Bonito G, Trappe JM. 2009 – Calongea, a new genus of truffles in the Pezizaceae
(Pezizales). Anales del Jardín Botánico de Madrid. Consejo Superior de Investigaciones
Científicas 66, 25–32.
Helaly SE, Thongbai B, Stadler M. 2018 – Diversity of biologically active secondary metabolites
from endophytic and saprotrophic fungi of the ascomycete order Xylariales. Natural Product
Reports 35, 992–1014.
Henkel TW, Aime MC, Chin MM L, Miller SL et al. 2012 – Ectomycorrhizal fugal sporocarp
diversity and discovery of new taxa in Dicymbe monodominant forests of the Guiana
Shield. Biodiversity & Conservation 21, 2195–2220.
Henkel TW, Obase K, Husbands D, Uehling JK et al. 2016 – New Boletaceae taxa from Guyana:
Binderoboletus segoi gen. and sp. nov., Guyanaporus albipodus gen. and sp. nov.,

1437
Singerocomus rubriflavus gen. and sp. nov., and a new combination for Xerocomus
inundabilis. Mycologia 108, 157–173.
Hermet A, Méheust D, Monunier J, Barbier G et al. 2012 – Molecular systematics in the genus
Mucor with special regards to species encountered in cheese. Fungal Biology 116, 692–705.
Hernández-Restrepo M, Castañeda-Ruíz RF, Gené J, Silvera-Simón C et al. 2014 – Two new
species of Solicorynespora from Spain. Mycological Progress 13, 157–164.
Heuchert B, Braun U, Diederich P, Ertz D. 2018 – Taxonomic monograph of the genus
Taeniolella s. lat. (Ascomycota). Fungal Systematics & Evolution 2, 69–261.
Hibbett D, Abarenkov K, Kõljalg U, Öpik M et al. 2016 – Sequence–based classification and
identification of Fungi. Mycologia 108, 1049–1068.
Hibbett DS, Binder M, Bischoff JF, Blackwell M et al. 2007 – A higher-level phylogenetic
classification of the Fungi. Mycological Research 111, 509–547.
Hinney B, Sak B, Joachim A, Kváč M. 2016 – More than a rabbit’s tale – Encephalitozoon spp. in
wild mammals and birds. International Journal for Parasitology: Parasites & Wildlife 5, 76–
87.
Hirooka Y, Tanney JB, Nguyen HDT, Seifert KA. 2015 – Xerotolerant fungi in house dust:
Taxonomy of Spiromastix, Pseudospiromastix and Sigleria gen. nov. in Spiromastigaceae
(Onygenales, Eurotiomycetes). Mycologia 108, 135–156.
Hittinger CT, Rokas A, Bai FY, Boekhout T et al. 2015 – Genomics and the making of yeast
biodiversity. Current Opinion in Genetics & Development 35, 100–109.
Hjortstam K, Ryvarden L. 2010 – Phaerodontia and Phaneroites two corticioid taxa
(Basidiomycotina) proposed from tropical areas. Syn Fung 27, 26–33.
Hoek C van den DG, Mann HM, Jahns. 1995 – Algae: An Introduction to Phycology. Cambridge:
Cambridge University Press. pp. 104, 124, 134, 166.
Hofstetter V, Redhead SA, Kauff F, Moncalvo JM et al. 2014 – Taxonomic revision and
examination of ecological transitions of the Lyophyllaceae (Basidiomycota, Agaricales)
based on a multigene phylogeny. Cryptogamie Mycologie 35, 399–425.
Hongsanan S, Maharachchikumbura SS, Hyde KD, Samarakoon MC et al. 2017 – An updated
phylogeny of Sordariomycetes based on phylogenetic and molecular clock evidence. Fungal
Diversity 84, 25–41.
Huang SK, Tangthirasunun N, Phillips AJL, Dai DQ et al. 2016 – Morphology and phylogeny of
Neoscytalidium orchidacearum sp. nov. (Botryosphaeriaceae). Mycobiology 44, 79–84.
Huhndorf SM, Miller AN, Fernández FA. 2004 – Molecular systematics of the Sordariales: the
order and the family Lasiosphaeriaceae redefined. Mycologia 96, 368–387.
Huhndorf SM. 1994 – Neotropical Ascomycetes 5. Hypsostromataceae, a new family of
Loculoascomycetes and Manglicola samuelsii, a new species from Guyana. Mycologia 86,
266–269.
Humber RA. 2012 – Entomophthoromycota: a new phylum and reclassification for
entomophthoroid fungi. Mycotaxon 120, 477–492.
Humber RA. 2016 – Entomophthoromycota: a new overview of some of the oldest terrestrial
fungi. In: Li DW (ed) Biology of microfungi. Springer, Cham, pp 127–145.
Hussein JM, Tibuhwa DD, Tibell S. 2018 – Phylogenetic position and taxonomy of Kusaghiporia
usambarensis gen. et sp. nov. (Polyporales). Mycology 9, 136–144.
Hustad VP, Miller AN, Dentinger BTM, Cannon PF. 2013 – Generic circumscriptions in
Geoglossomycetes. Persoonia 31, 101–111.
Hustad VP, Miller AN, Moingeon JM, Priou JP. 2011 – Inclusion of Nothomitra in
Geoglossomycetes. Mycosphere 2, 646–654.
Hustad VP, Miller AN. 2015 – Maasoglossum, a basal genus in Geoglossomycetes. Mycoscience
56, 572–579.
Hutchinson SA. 1955 – A review of the genus Sporocarpon Williamson. Annals of Botany 19,
425–435.

1438
Hyde KD, Chaiwan N, Norphanphoun C, Boonmee S et al. 2018a – Mycosphere notes 169–224.
Mycosphere 9, 271–430.
Hyde KD, Jones EBG, Liu JK, Ariyawansa H et al. 2013 – Families of Dothideomycetes. Fungal
Diversity 63, 1–313.
Hyde KD, Norphanphoun C, Bazzicalupo A, Karunarathna A et al. 2017 – Fungal diversity notes
603–708: Taxonomic and phylogenetic notes on genera and species. Fungal Diversity 87, 1–
235.
Hyde KD, Norphanphoun C, Chen J, Dissanayake AJ et al. 2018b – Thailand’s amazing diversity:
up to 96% of fungi in northern Thailand may be novel. Fungal Diversity 93, 215–239.
Hyde KD, Norphanphoun C, Maharachchikumbura SSN, Bhat DJ et al. 2020 – Refined families
of Sordariomycetes. Mycosphere 11, 305–1059.
Ibañez CG & Zamuner AB. 1996 – Hyphomycetes (Deuteromycetes) in cones of Araucaria
mirabilis (Spegazzini) Windhausen, Middle Jurassic of Patagonia, Argentina; Mycotaxon
59, 137–143.
Inderbitzin P, Mehta YR, Berbee ML. 2009 – Pleospora species with Stemphylium anamorphs: a
four locus phylogeny resolves new lineages yet does not distinguish among species in the
Pleospora herbarum clade. Mycologia 101, 329–339.
Index Fungorum (2019) http://www.indexfungorum.org/Names/Names.asp
Ironside JE, Wilkinson TJ, Rock J. 2008 – Distribution and host range of the microsporidian
Pleistophora mulleri. Journal of Eukaryotic Microbiology 55, 355–362.
Isola D, Zucconi L, Onofri S, Caneva G et al. 2016 – Extremotolerant rock–inhabiting black fungi
from Italian monumental sites. Fungal Diversity 76, 75–96.
Issi IV 1980 – Modern situation in systematics of Microsporidia. Proceedings of Zoological
Institute 94, 75–84 (In Russian)
Issi IV, Tokarev YS, Seliverstova EV, Voronin VN. 2012a – Taxonomy of Neopereziachironomi
and Neoperezia semenovaiae comb. nov. (Microsporidia, Aquasporidia): lessons from
ultrastructure and ribosomal DNA sequence data. European Journal of Protistology 48, 17–
29.
Issi IV, Tokarev YS, Seliverstova EV, Voronin VN. 2012b – The parasite-host interface between
Crispospora chironomi (Microsporidia, Terresporidia) and Chironomus plumosus (Diptera,
Chironomidae) enterocytes. Euroasian Entomological Journal 11, 395–400.
Issi IV, Tokarev YS, Voronin VN, Seliverstova EV et al. 2010 – Ultrastructure and molecular
phylogeny of Mrazekia macrocyclopis sp. n. (Microsporidia, Mrazekiidae), a microsporidian
parasite of Macrocyclops albidus (Jur.) (Crustacea, Copepoda). Acta Protozoologica 49, 75–
84.
Issi IV. 1986 – Microsporidia as a phylum of parasitic protozoa. In: Beyer TV and Issi IV (eds.),
Protozoology 10. Leningrad: Nauka, pp 1–136.
Iturrieta–González I, Gené J, Guarro J, Castañeda–Ruiz RF, García D. 2018 – Neodendryphiella,
a novel genus of the Dictyosporiaceae (Pleosporales). MycoKeys 37, 19–38.
Jacobs K, Botha A. 2008 – Mucor renisporus sp. nov., a new coprophilous species from Southern
Africa. Fungal Diversity 29, 27–35.
Jain KP, Gupta RC. 1970 – Some fungal remains from the Tertiaries of Kerala Coast. The
Palaeobotanist 18, 177–182.
Jaklitsch WM, Baral HO, Lücking R, Lumbsch HT. 2016a – Ascomycota. In: Frey W (ed)
Syllabus of Plant Families – Adolf Engler’s Syllabus der Pflanzenfamilien, 13th ed.
Borntraeger, Stuttgart
Jaklitsch WM, Checa J, Blanco MN, Olariaga I et al. 2018 – A preliminary account of the
Cucurbitariaceae. Studies in Mycology 90, 71–118.
Jaklitsch WM, Gardiennet A, Voglmayr H. 2016b – Resolution of morphology–based taxonomic
delusions: Acrocordiella, Basiseptospora, Blogiascospora, Clypeosphaeria, Hymenopleella,
Lepteutypa, Pseudapiospora, Requienella, Seiridium and Strickeria. Persoonia 37, 82–105.

1439
Jaklitsch WM, Voglmayr H. 2019 –European species of Dendrostoma (Diaporthales). MycoKeys
59, 1–26.
James TY, Kauff F, Schoch CL, Matheny P et al. 2006 – Reconstructing the early evolution of
Fungi using a six–gene phylogeny. Nature 443, 818–822.
James TY, Pelin A, Bonen L, Ahrendt S et al. 2013 – Shared signatures of parasitism and
phylogenomics unite Cryptomycota and microsporidia. Current Biology 23, 1548–1553.
Jansonius J, Hills LV. 1976 – Genera file of fossil spores. Special Publication, Department of
Geology, University of Calgary.
Jiang HB, Hyde KD, Jayawardena RS, Doilom M et al. 2019a – Taxonomic and phylogenetic
characterizations reveal two new species and two new records of Roussoella
(Roussoellaceae, Pleosporales) from Yunnan, China. Mycol Prog 18, 577–591.
Jiang N, Fan XL, Crous PW, Tian CM. 2019b – Species of Dendrostoma (Erythrogloeaceae,
Diaporthales) associated with chestnut and oak canker diseases in China. MycoKeys 48,
67–96.
Jiang N, Yang Q, Liang YM, Tian CM. 2019c – Taxonomy of two synnematal fungal species
from Rhus chinensis, with Flavignomonia gen. nov. described. MycoKeys 60, 17–29.
Jiang SH, Hawksworth DL, Lücking R, Wei JC 2020 – A new genus and species of foliicolous
lichen in a new family of Strigulales (Ascomycota: Dothideomycetes) reveals remarkable
class-level homoplasy. IMA fungus 11, 1–13.
Jobim K, Błaszkowski J, Niezgoda P, Kowsloska A et al. 2019 – New sporocarpic taxa in the
phylum Glomeromycota: Sclerocarpum amazonicum gen. et sp. nov. in the family
Glomeraceae (Glomerales) and Diversispora sporocarpia sp. nov. in Diversisporaceae
(Diversisporales). Mycological Progress 18, 369–384.
Johnston PR, Park D. 2019 – New species of Marthamyces and Ramomarthamyces gen. nov. from
New Zealand and the Cook Islands. Mycotaxon 134, 489–516.
Johnston PR, Quijada L, Smith CA, Baral HO, Hosoya T et al. 2019 – A multigene phylogeny
toward a new phylogenetic classification for the Leotiomycetes. IMA Fungus 10, 1.
Jones EBG, Sakayaroj J, Suetrong S, Somrithipol S et al. 2009 – Classification of marine
Ascomycota, anamorphic taxa and Basidiomycota. Fungal Diversity 35, 1–187.
Jones MDM, Forn L, Gadelha C, Egan MJ et al. 2011 – Discovery of novel intermediate forms
redefines the fungal tree of life. Nature 474, 200–203.
Joseph S, Sinha GP, Ramachandran VS. 2018 – Taxonomic revision of the lichen genus
Opegrapha sense lato (Roccellaceae) in India. Indian Journal of Forestry Additional Series
6, 1–172.
Jung PE, Lee H, Wu SH, Hattori T et al. 2018 – Revision of the taxonomic status of the genus
Gloeoporus (Polyporales, Basidiomycota) reveals two new species. Mycological Progress
17, 855–863.
Justo A, Miettinen O, Floudas D, Ortiz-Santana B et al. 2017 – A revised family–level
classification of the Polyporales (Basidiomycota). Fungal Biology 121, 798–824.
Kalgutkar RM, Jansonius J. 2000 – Synopsis of fungal spores, mycelia and fructifications. AASP
Contribution Series 39, 1–423.
Kalgutkar RM, Nambudiri EMV, Tidwell WD. 1993 – Diplodites sweetii sp. nov. from the Late
Cretaceous (Maastrichtian) Deccan Intertrappean beds of India. Review of Palaeobotany
and Palynology 77, 107–118.
Kalgutkar RM. 1997 – Fossil fungi from the lower Tertiary Iceberg Bay Formation, Eureka Sound
Group, Axel Heiberg Island, Northwest Territories, Canada. Review of Palaeobotany and
Palynology 97, 197–226.
Kang S, Tice AK, Spiegel FW, Silberman JD et al. 2017 – Between a pod and a hard test: The
deep evolution of Amoebae. Molecular Biology and Evolution 34, 2258–2270.
Karakehian JM, Quijada L, Friebes G, Tanney JB et al. 2019 – Placement of Triblidiaceae in
Rhytismatales and comments on unique ascospore morphologies in Leotiomycetes (Fungi,
Ascomycota). Mycokeys 54, 99–133.

1440
Karpov SA, Mamkaeva MA, Aleoshin VV, Nassonova E et al. 2014 – Morphology, phylogeny,
and ecology of the aphelids (Aphelidea, Opisthokonta) and proposal for the new
superphylum Opisthosporidia. Frontiers in Microbiology 5, 112.
Karunarathna A, Phookamsak R, Jayawardena RS, Cheewangkoon R et al. 2019 – Neoroussoella
alishanense sp. nov on Pennisetum purpureum (Poaceae) with an asexual/sexual morph
connection. Phytotaxa 406, 218–236.
Keane R, Berlemann J. 2016 – The predatory life cycle of Myxococcusxanthus. Microbiol 162, 1–
11.
Keeling PJ, Fast NM, Corradi N. 2014 – Microsporidian genome structure and function. In: Weiss
LM, Becnel JJ (eds) Microsporidia: pathogens of opportunity. Wiley–Blackwell Press,
Ames, pp 221–229
Keeling PJ, Luker MA, Palmer JD. 2000 – Evidence from beta–tubulin phylogeny that
microsporidia evolved from within the fungi. Mol Biol Evol 17, 23–31.
Kendrick WB, Carmichael JW. 1973 – Hyphomycetes. in Ainsworth GC, Sparrow FK & Sussman
AA (eds.) The Fungi. An Advanced Treatise, volume 4A, Academic Press, New York: 323–
509.
Kendrick WB. 1958 – Helicoma monospora sp. nov. from pine litter. Transactions of the British
Mycological Society 41, 446–448.
Kiran M, Sattar A, Zamir K, Haelewaters D et al. 2020 – Additions to the genus Chroogomphus
(Boletales, Gomphidiaceae) from Pakistan. MycoKeys 66, 23–38.
Kirk PM, Cannon PF, Minter DW, Stalpers JA. 2008 – Dictionary of the Fungi. (10th edn).
Wallingford, UK.
Kirk PM, Stalpers JA, Braun U, Crous PW et al. 2013 – A without prejudice list of generic names
of fungi for protection under the International Code of Nomenclature for algae, fungi, and
plants. IMA Fungus 4, 381–443.
Kiss L, Kovács GM, Bóka K, Bohár G et al. 2018 – Deciphering the biology of
Cryptophyllachora eurasiatica gen. et sp. nov., an often cryptic pathogen of an allergenic
weed, Ambrosia artemisiifolia. Scientific Reports 8, 10806.
Kistenich S, Timdal E, Bendiksby M, Ekman S. 2018 – Molecular systematics and character
evolution in the lichen family Ramalinaceae (Ascomycota: Lecanorales). Taxon 67, 871–
904.
Kodsueb R, Dhanasekaran V, Aptroot A, Lumyong S et al. 2006 – The family Pleosporaceae:
intergeneric relationships and phylogenetic perspectives based on sequence analyses of
partial 28S rDNA. Mycologia 98, 571–583.
Kohout P, Sudová R, Janoušková M, Čtvrtlíková M et al. 2014 – Comparison of commonly used
primer sets for evaluating arbuscular mycorrhizal fungal communities: Is there a universal
solution? Soil Biology & Biochemistry 68, 482–493.
Košuthová A, Westberg M, Otálora MAG, Wedin M. 2019 – Rostania revised: testing generic
delimitations in Collemataceae (Peltigerales, Lecanoromycetes). MycoKeys 47, 17–33.
Kraichak E, Huang JP, Nelsen M, Leavitt SD et al. 2018a – A revised classification of orders and
families in the two major subclasses of Lecanoromycetes (Ascomycota) based on a temporal
approach. Botanical Journal of the Linnean Society 188, 233–249.
Kraichak E, Huang JP, Nelsen M, Leavitt SD et al. 2018b – Nomenclatural novelties. Index
Fungorum 375, 1–1.
Kumar LM, Smith ME, Nouhra ER, Orihara T et al. 2017 – A molecular and morphological re–
examination of the generic limits of truffles in the tarzetta–geopyxis lineage – Densocarpa,
Hydnocystis, and Paurocotylis. Fungal Biology 121, 264–284.
Kumar P. 1990 – Fungal remains from the Miocene Quilon beds of Kerala State, South India.
Review of Palaeobotany and Palynology 63, 13–28.
Kurtzman CP, Boekhout T. 2017 – Yeasts as distinct life forms of fungi. In: Buzzini P, Lachance
MA, Yurkov A (eds) Yeasts in natural ecosystems: Ecology. Springer International, Cham,
pp 1–37.

1441
Kurtzman CP. 2011 – Discussion of teleomorphic and anamorphic ascomycetous yeasts and
yeast–like Taxa. In: Kurtzman CP, Fell JW, Boekhout T (eds) The yeasts – A taxonomic
study, 5th edn. Elsevier Science, Burlington, pp 293–307.
Kušan I, Matočec N, Jadan M, Tkalčec Z et al. 2018 – An overview of the genus Coprotus
(Pezizales, Ascomycota) with notes on the type species and description of C. epithecioides
sp, nov. Mycokeys 29, 15–47.
Kušan I, Matočec N, Mešić A, Tkalčec Z. 2015 – A new species of Thecotheus from Croatia with
a key to the known species with apiculate spores. Sydowia 67, 51–63.
Labbé A. 1899 – Microsporidiida. In: Bütschli O, Ed Das Tierreich Vol 5 Sporozoa, Berlin,
Germany: Friedländer und Sohn, pp. 104–112.
Lakhanpal RN, Maheshwari HK, Awasthi N. 1976 – A Catalogue of Indian Fossil Plants. Birbal
Sahni Institute of Palaeobotany, Lucknow, India. pp. 1–318.
Lambert C, Wendt L, Hladki AI, Stadler M et al. 2019 – Hypomontagnella (Hypoxylaceae): a new
genus segregated from Hypoxylon by a polyphasic taxonomic approach. Mycological
Progress 18, 187–201.
Lang PL, Willems FM, Scheepens JF, Burbano HA et al. 2019 – Using herbaria to study global
environmental change. New Phytologist 221, 110–122.
Larsson JIR 2014 – The primitive microsporidia. In: Weiss LM and Becnel JJ (eds.),
Microsporidia pathogens of opportunity. John Wiley & Sons, Inc., Ames, Iowa, pp 605–634.
Lechat C, Fournier J, Moreau PA. 2016b – Xanthonectria, a new genus for the nectrioid fungus
Nectria pseudopeziza. Ascomycete.org 8, 172–178.
Lechat C, Fournier J, Priou JP. 2018a – Chrysonectria, a new genus in the Nectriaceae with the
new species C. finisterrensis from France. Ascomycete.org 10, 121–125.
Lechat C, Fournier J, Vega M, Priou J–P 2018b – Geonectria, a new genus in the Bionectriaceae
from France. Ascomycete.org 10, 81–85.
Lechat C, Fournier J. 2016 – Varicosporellopsis, a new aquatic genus from southern France.
Ascomycete.org 8, 96–100.
Lechat C, Fournier J. 2016a – Lasionectriella, a new genus in the Bionectriaceae, with two new
species from France and Spain. Ascomycete.org 8, 59–65.
Leedale GF 1974 – How many are the kingdoms of organisms? Taxon 23, 261–270.
Lee SC, Heitman J, Ironside JE. 2014 – Sex and the Microsporidia In: Weiss LM, Becnel JJ (eds)
Microsporidia: pathogens of opportunity. Wiley–Blackwell Press, Ames, 231–243.
Leontyev DV, Schnittler M, Stephenson S, NovozhilovYK et al. 2019 – Towards a phylogenetic
classification of Myxomycetes. Phytotaxa 399, 209–238.
Leontyev DV, Schnittler M, Stephenson SL. 2014 – Pseudocapillitium or true capillitium? A
study of capillitial structures in Alwisia bombarda (Myxomycetes). Nova Hegwigia 99, 441–
451.
Leontyev DV, Schnittler M. 2017 – The phylogeny of myxomycetes. Pages 83–105 in:
Stephenson SL, Rojas C (Eds.) Myxomycetes. Biology, Systematics, Biogeography and
Ecology. Elsevier Academic Press.
Letcher PM, Powell MJ, Davis WJ. 2018 – Morphology, zoospore ultrastructure, and molecular
position of taxa in the Asterophlyctis lineage (Chytridiales, Chytridiomycota). Fungal
Biology 122, 1109–1123.
Letcher PM, Powell MJ. 2018 – A taxonomic summary and revision of Rozella (Cryptomycota).
IMA fungus 9, 383–399.
Li GJ, Hyde KD, Zhao RL, Hongsanan S et al. 2016 – Fungal diversity notes 253–366: taxonomic
and phylogenetic contributions to fungal taxa. Fungal Diversity77, 1–237.
Li Y, Xiao J, de Hoog GS, Wang X et al. 2017 – Biodiversity and human pathogenicity of
Phialophora verrucosa and relatives in Chaetothyriales. Persoonia 38, 1–19.
Li YM, Shivas RG, Cai L. 2017 – Cryptic diversity in Tranzscheliella spp. (Ustilaginales) is
driven by host switches. Scientific Reports 7, 43549.

1442
Lima DX, Souza–Motta CM, Wagner L, Voigt K et al. 2017 – Circinella simplex –– a misapplied
name of Mucor circinatus sp. nov. Phytotaxa 329, 269–276.
Lin CC, Aronson JM 1970 – Chitin and cellulose in the cell walls of the oomycete, Apodachlya
sp. Archiv für Mikrobiologie, 72, 111–114.
Linaldeddu BT, Alves A, Phillips AJL. 2016 – Sardiniella urbana gen. et sp. nov., a new member
of the Botryosphaeriaceae isolated from declining Celtis australis trees in Sardinian
streetscapes. Mycosphere 7, 893–905.
Lindemann U, Vega M, Alvarado P. 2015 – Revision der Gattung Kotlabaea: K. deformis, K.
delectans und K. Benkertii. Zeitschrift für Mykologie 81, 373–402.
Lister A 1894 – A monograph of the Mycetozoa. British Museum London.
Liu F, Bonthond G, Groenewald JZ, Cai L et al. 2019 – Sporocadaceae, a family of
coelomycetous fungi with appendage-bearing conidia. Studies in Mycology 92, 287–415.
Liu JK, Hyde KD, Jeewon R, Phillips AJL et al. 2017 – Ranking higher taxa using divergence
times: a case study in Dothideomycetes. Fungal Diversity 84, 75–99.
Liu NG, Hyde KD, Bhat DJ, Jumpathong J, Liu JK. 2019 – Morphological and phylogenetic
studies of Pleopunctum gen. nov. (Phaeoseptaceae, Pleosporales) from China. Mycosphere
10, 757–775.
Liu NG, Lin CG, Liu JK, Samarakoon MC et al. 2018a – Lentimurisporaceae, a new
Pleosporalean family with divergence times estimates. Cryptogamie, Mycologie 39, 259–
283.
Liu SL, Nakasone KK, Wu SH, He SH, Dai YC. 2018b – Taxonomy and phylogeny of Lopharia
s.s., Dendrodontia, Dentocorticium and Fuscocerrena (Basidiomycota, Polyporales).
MycoKeys 32, 25–48.
Liu XZ, Wang Q M, Göker M et al. 2015 – Towards an integrated phylogenetic classification of
the Tremellomycetes. Studies in Mycology 81, 85–147.
Lücking R, Dal-Forn, M, Sikaroodi M, Gillevet PM et al. 2014 – A single macrolichen constitutes
hundreds of unrecognized species. Proceedings of the National Academy of Sciences of the
USA 111, 11091–11096.
Lücking R, Hodkinson BP, Leavitt SD. 2017 – The 2016 classification of lichenized fungi in the
Ascomycota and Basidiomycota–Approaching one thousand genera. Bryologist 119, 361–
417.
Lücking R, Kalb K. 2018 – Formal instatement of Allographa (Graphidaceae): How to deal with
a hyperdiverse genus complex with cryptic differentiation and paucity of molecular data. –
Herzogia 31, 535–561.
Lücking R, Moncada B, Hawksworth DL. 2019 – Gone with the wind: sequencing its type species
supports inclusion of Cryptolechia in Gyalecta (Ostropales: Gyalectaceae). Lichenologist
51, 287–299.
Lücking R, Tehler A, Bungartz F, Rivas Plata E et al. 2013 – Journey from the West: did tropical
Graphidaceae (lichenized Ascomycota: Ostropales) evolve from a saxicolous ancestor along
the American Pacific coast?. American Journal of Botany 100, 844–56.
Lumbsch HT, Huhndorf SM 2010 – Outline of Ascomycota – 2009. Myconet 14, 1–64.
Ma X, Zhao CL. 2019 – Crepatura ellipsospora gen. et sp. nov. in Phanerochaetaceae
(Polyporales, Basidiomycota) bearing a tuberculate hymenial surface. Mycological Progress
18, 785–793
Ma YR, Xia JW, Gao JM, Li Z et al. 2016 – Dictyoceratosporella gen. nov. with the description
of two new species collected from Hainan, China. Sydowia 68, 57–61.
Madden AA, Stchigel AM, Guarro J, Sutton D et al. 2012 – Mucor nidicola sp. nov., a novel
fungal species isolated from an invasive paper wasp nest. International Journal of
Systematic & Evolutionary Microbiology 62, 1710–1714.
Madrid H, Cano J, Gené J, Guarro J. 2011 – Two new species of Cladorrhinum. Mycologia 103,
795–805.

1443
Madrid H, Gené J, Cano J, Stchigel A et al. 2010 – Ramophialophora humicola and
Fibulochlamys chilensis, two new microfungi from soil. Mycologia 102, 605–612.
Madrid H, Hernández-Restrepo M, Gené J, Cano J et al. 2016 – New and interesting
chaetothyrialean fungi from Spain. Mycological Progress 15, 1179–1201.
Magyar D, Merényi Z, Udvardy O, Kajtor-Apatini D et al. 2018 – Mycoceros antennatissimus
gen. et sp. nov.: a mitosporic fungus capturing pollen grains. Mycological Progress 17, 33–
43.
Malysh JM, Ignatieva AN, Artokhin KS, Frolov AN et al. 2018b – Natural infection of the beet
webworm Loxostegesticticalis L. (Lepidoptera: Crambidae) with three Microsporidia and
host switching in Nosema ceranae. Parasitology Research 117, 3039–3044.
Malysh JM, Kononchuk AG, Frolov AN 2019 – Detection of microsporidia infecting beet
webworm Loxostege sticticalis (Pyraloidea: Crambidae) in European part of Russia in
2006–2008. Plant Protection News [Vestnik zashchity rasteniy] 2, 45–51.
Malysh JM, Vorontsova YL, Glupov VV, Tsarev AA, Tokarev YS. 2018a – Vairimorpha
ephestiae is a synonym of Vairimorpha necatrix (Opisthosporidia: Microsporidia) based on
multilocus sequence analysis. European Journal of Protistology 66, 63–67.
Malysheva V, Spirin V. 2017 – Taxonomy and phylogeny of the Auriculariales (Agaricomycetes,
Basidiomycota) with stereoid basidiocarps. Fungal Biology 121, 689–715.
Mapook A, Hyde KD, McKenzie EHC, Gareth Jones EBG et al. 2020 – Taxonomic and
phylogenetic contributions to fungi associated with the invasive weed Chromolaena odorata
(Siam weed). Fungal Diversity https://doi.org/10.1007/s13225-020-00444-8
Marano AV, Jesus AL, de Souza JI, Jeronimo GH, Gonçalves DR, Boro MC, Rocha SCO, Pires-
Zottarelli CLA 2016 – Ecological roles of saprotrophic Peronosporales (Oomycetes,
Straminipila) in natural environments. Fungal Ecology 19, 77–88.
Marchetta A, Gerrits van den Ende B, Al-Hatmi AMS, Hagen F et al. 2018 – Global molecular
diversity of the halotolerant fungus Hortaea werneckii. Life 8, 31.
Marinho F, Silva GA, Ferreira ACA, Veras JSN et al. 2014 – Bulbospora minima, new genus and
new species in the Glomeromycetes from semi–arid Northeast Brazil. Sydowia 66, 313–323.
Marmolejo J, Siahaan SAS, Takamatsu S, Braun U. 2018 – Three new records of powdery
mildews found in Mexico with one genus and one new species proposed. Mycoscience 59,
1–7.
Martin GW, Alexopoulos CJ. 1969 – The Myxomycetes. Iowa Univ Press, Iowa City.
Massee G. 1892 – A monograph of the Myxogasteres. Methuen Co, London.
Matočec N, Kušan I, Ozimec R. 2014 – The genus Polycephalomyces (Hypocreales) in the frame
of monitoring Veternica cave (Croatia) with the new segregate genus Perennicordyceps.
Ascomycete.org 6, 125–133.
Mehrabi M, Asgari B, Hemmati R. 2018 – Knufia perfecta, a new black yeast from Iran, and a key
to Knufia species. Nova Hedwigia 106, 519–534.
Mehrotra BS, Mehrotra BM. 1979 [1978] – Another azygosporic species of Mucor from India.
Sydowia 31, 94–96.
Meissner EG, Bennett JE, Qvarnstrom Y, da Silva A et al. 2012 – Disseminated microsporidiosis
in an immunosupplpressed patient. Emerging Infectious Diseases 18, 1155–1158.
Mikhailov KV, Simdyanov TG, Aleoshin VV. 2016 – Genomic survey of a hyperparasitic
microsporidian Amphiamblys sp. (Metchnikovellidae). Genome Biology and Evolution 9,
454–467.
Minnis AM, Kennedy AH, Grenier DB, Palm ME, Rossman AY. 2012 – Phylogeny and
taxonomic revision of the Planistromellaceae including its coelomycetous anamorphs:
contributions towards a monograph of the genus Kellermania. Persoonia 29, 11–28.
Minnis AM, Lindner DL 2013 – Phylogenetic evaluation of Geomyces and allies reveals no close
relatives of Pseudogymnoascus destructans, comb. nov., in bat hibernacula of eastern North
America. Fungal Biology 117, 638–649.

1444
Mirza JH, Khan SM, Begum S, Shagufta S. 1979 – Mucorales of Pakistan. University of
Agriculture, Faisalabad, Pakistan.
Mishra B, Choi Y-J, Thines M 2018 – Phylogenomics of Bartheletia paradoxa reveals its basal
position in Agaricomycotina and that the early evolutionary history of basidiomycetes was
rapid and probably not strictly bifurcating. Mycological Progress 17, 333–341.
Moncalvo JM, Vilgalys R, Redhead SA, Johnson JE et al. 2002 – One hundred and seventeen
clades of euagarics. Molecular Phylogenetics and Evolution 23, 357–400.
Morton JB, Benny GL. 1990 – Revised classification of arbuscular mycorrhizal fungi
(Zygomycetes). A new order, Glomales, two new suborders, Glomineae and Gigasporineae,
and two new families, Acaulosporaceae and Gigasporaceae, with an emendation of
Glomaceae. Mycotaxon 37, 471–491.
Morton JB, Msiska Z. 2010 – Phylogenies from genetic and morphological characters do not
support a revision of Gigasporaceae (Glomeromycota) into four families and five genera.
Mycorrhiza 20, 483–496.
Motato-Vásquez V, Grassi E, Gugliotta AM, Robledo GL. 2018 – Evolutionary relationships of
Bresadolia (Basidiomycota, Polyporales) based on molecular and morphological evidence.
Mycological Progress 17, 1031–1048.
Moussa TA, Gerrits van den Ende BH, Al Zahrani HS, Kadasa NM et al. 2017 – The genus
Anthopsis and its phylogenetic position in Chaetothyriales. Mycoses 60, 254–259.
Muggia L, Kopun T, Ertz D. 2015 – Phylogenetic placement of the lichenicolous, anamorphic
genus Lichenodiplis and its connection to Muellerella–like teleomorphs. Fungal Biology
119, 1115–1128.
Müller E. 1950 – Die schweizerischen Arten der Gattung Leptosphaeria und ihrer Verwandten.
Sydowia 4, 185–319
Nägeli C. 1857 – Über die neue Krankheit der Seidenraupe und verwandte Organismen. Bot
Zeitung 15, 760–761.
Nascimento MM, Selbmann L, Sharifynia S, Al-Hatmi AM et al. 2016 – Arthrocladium, an
unexpected human opportunist in Trichomeriaceae (Chaetothyriales). Fungal Biology 120,
207–218.
Nasr S, Soudi MR, Fazeli SAS, Nguyen HD et al. 2014 – Expanding evolutionary diversity in the
Ustilaginomycotina: Fereydouniaceae fam. nov. and Fereydounia gen. nov., the first
urocystidalean yeast lineage. Mycological Progress 13, 1012.
Nylund S, Nylund A, Watanabe K, Arnesen CE et al. 2010 – Paranucleospora theridion n. gen., n.
sp. (Microsporidia, Enterocytozoonidae) with a Life Cycle in the Salmon Louse
(Lepeophtheirussalmonis, Copepoda) and Atlantic Salmon (Salmosalar). Journal of
Eukaryotic Microbiology 57, 95–114.
Oehl F, da Silva DKA, Maia LC, de Sousa NMF, da Silva GA et al. 2011d – Orbispora gen. nov.,
ancestral in the Scutellosporaceae (Glomeromycetes). Mycotaxon 116, 161–169.
Oehl F, da Silva GA, Goto BT, Sieverding E. 2011e – New recombinations in Glomeromycota.
Mycotaxon 117, 429–434.
Oehl F, da Silva GA, Sánchez-Castro I, Goto BT et al. 2011f – Revision of Glomeromycetes with
entrophosporoid and glomoid spore formation with three new genera. Mycotaxon 117, 297 –
316.
Oehl F, de Souza FA, Sieverding E. 2008 – Revision of Scutellospora and description of five new
genera and three new families in the arbuscular mycorrhiza–forming Glomeromycetes.
Mycotaxon 106, 311–360.
Oehl F, Sánchez–Castro I, Palenzuela J, da Silva GA. 2015 – Palaeospora spainii, a new
arbuscular mycorrhizal fungus from Swiss agricultural soils. Nova Hedwig 101, 89–102.
Oehl F, Sieverding E, Palenzuela J, Ineichen K et al. 2011c – Advances in Glomeromycota
taxonomy and classification. IMA fungus 2, 191–199.
Oehl F, Silva GA, Goto BT, Maia LC et al. 2011b – Glomeromycota: two new classes and a new
order. Mycotaxon 116, 75–120.

1445
Oehl F, Silva GA, Goto BT, Sieverding E. 2011a – Glomeromycetes: three new genera and
glomoid species reorganized. Mycotaxon 116, 75–120.
Olive LS, Stoianovitch С. 1975 – The Mycetozoans. Academic Press, New York.
Orihara T, Smith ME. 2017 – Unique phylogenetic position of the African truffle–like fungus,
Octaviania ivoryana (Boletaceae, Boletales), and the proposal of a new genus,
Afrocastellanoa. Mycologia 109, 323–332.
Ovcharenko M, Wróblewski P, Kvach Y, Drobiniak O. 2017 – Study of Loma acerinae
(Microsporidia) detected from three Ponto-Caspian gobies (Gobiidae) in Ukraine.
Parasitology Research 116, 1453–1462.
Papp V, Dima B. 2018 – New systematic position of Aurantiporus alborubescens (Meruliaceae,
Basidiomycota), a threatened old–growth forest polypore. Mycological Progress 17, 319–
332.
Pärtel K, Baral HO, Tamm H, Põldmaa K. 2017 – Evidence for the polyphyly of Encoelia and
Encoelioideae with reconsideration of respective families in Leotiomycetes. Fungal
Diversity 82, 183–219.
Pei KQ. 2000 – A new variety of Mucor variosporus and the validation of M. luteus Linnemann
and M. variosporus Schipper. Mycosystema 19, 10–12.
Pérez-Ortega S, Suija A, de los Rios A. 2011 – The connection between Abrothallus and its
anamorph state Vouauxiomyces established by Denaturing Gradient Gel Electrophoresis
(DGGE). Lichenologist 43, 277–279.
Perry BA, Hansen K, Pfister DH. 2007 – A phylogenetic overview of the family Pyronemataceae
(Ascomycota, Pezizales). Mycological Research 111, 549–571.
Petersen HE 1905 – Contributions a la connaissance des Phycomycetes marins (Chytridineae
Fischer). Oversigt over det Kongelige Danske videnskabernes selskabs forhandlinger 5,
439–188.
Petersen RH, Hughes KW. 2010 – The Xerula/ Oudemansiella complex (Agaricales). Nova
Hedwigia 137, 1–165.
Peyretaillade E, Boucher D, Parisot N, Gasc C et al. 2015 – Exploiting the architecture and the
features of the microsporidian genomes to investigate diversity and impact of these parasites
on ecosystems. Heredity 114, 441–449.
Phillips AJL, Hyde KD, Alves A, Liu JK. 2018 – Families in Botryosphaeriales: a phylogenetic,
morphological and evolutionary perspective. Fungal Diversity 94, 1–22.
Phookamsak R, Boonmee S, Norphanphoun C, Wanasinghe DN et al. 2016 – Schizothyriaceae.
Mycosphere 7, 154–189.
Phookamsak R, Hyde KD, Jeewon R, Bhat DJ et al. 2019 – Fungal Diversity notes 929–1035:
taxonomic and phylogenetic contributions on genera and species of fungal taxa. Fungal
Diversity 95, 1–273.
Phookamsak R, Liu JK, McKenzie EH, Manamgoda DS et al. 2014 – Revision of
Phaeosphaeriaceae. Fungal Diversity 68, 159–238.
Phookamsak R, Wanasinghe DN, Hongsanan S, Phukhamsakda C et al. 2017 – Towards a natural
classification of Ophiobolus and ophiobolus-like taxa; introducing three novel genera
Ophiobolopsis, Paraophiobolus and Pseudoophiobolus in Phaeosphaeriaceae
(Pleosporales). Fungal Diversity 87, 299–339.
Pino-Bodas R, Zhurbenko MP, Stenroos S. 2017 – Phylogenetic placement within
Lecanoromycetes of lichenicolous fungi associated with Cladonia and some other genera. –
Persoonia 39, 91–117.
Pirozynski KA, Weresub LK. 1979 – The classification and nomenclature of fossil fungi. in
Kendrick, B. (ed.), The whole fungus, the sexual–asexual synthesis. Proceedings of the 2nd
International Mycological Conference, University of Calgary, Kananaskis, Alberta
(published by National Museum of Natural Sciences, National Museums of Canada and the
Kananaskis Foundation) volume 2, 653–688.

1446
Pirozynski KA. 1978 – Fungal spores through the ages – a mycologist’s view. Proceedings of the
Fourth International Palynological Conference, Lucknow 1976–77, volume 1, 327–330.
Poulain M, Meyer M, Bozonnet J. 2011 – Les Myxomycètes. FédMycol Bot Dauphiné–Savoie,
Delémont.
Powell MJ, Letcher PM, Longcore JE, Blackwell WH. 2018 – Zopfochytrium is a new genus in
the Chytridiales with distinct zoospore ultrastructure. Fungal Biology 122, 1041–1049.
Prasher IB, Sharma R, Singh G 2016 – Gelatinoamylaria gen. nov. (Dermateaceae, Helotiales)
from Bhutan. Kavaka 46, 35–36.
Prieto M, Schultz M, Olariaga I, Wedin M. 2018 – Lichinodium is a new lichenized lineage in the
Leotiomycetes. Fungal Diversity 94, 23–39.
Pringle A, Baker DM, Platt JL, Wares JP et al. 2005 – Cryptic speciation in the cosmopolitan and
clonal human pathogenic fungus Aspergillus fumigatus. Evolution 59, 1886–1899.
Printzen C, Spribille T, Tønsberg T. 2008 – Myochroidea, a new genus of corticolous, crustose
lichens to accommodate the Lecidea leprosula group. Lichenologist 40, 195–207.
Purrini K, Weiser J. 1985 – Ultrastructural study of the microsporidian Chytridiopsis typographi
(Chytridiopsida: Microspora) infecting the bark beetle, Ips typographus (Scolytidae:
Coleoptera), with new data on spore dimorphism. Journal of Invertebrate Pathology 45, 66–
74.
Puytorac P de, Grain J, Mignot JP. 1987 – Précis de protistologie. Lubrecht & Cramer Ltd, 1987,
581 p.
Quaedvlieg W, Verkley GJM, Shin HD, Barreto RW et al. 2013 – Sizing up Septoria. Studies in
Mycology 75, 307–390.
Quandt CA, Beaudet D, Corsaro D, Walochnik J et al. 2017 – The genome of an intranuclear
parasite, Paramicrosporidium saccamoebae, reveals alternative adaptations to obligate
intracellular parasitism. Elife 24; 6, e29594.
Quijada L, Johnston PR, Cooper JA & Pfister D. 2018a – Overview of Phacidiales, including
Aotearoamyces gen. nov. on Nothofagus. IMA fungus 9, 371–38.
Quijada L, Matočec N, Kušan I, Baral HO et al. 2018b – Insights into Claussenomyces Kirschst.:
Past, Present and Future. 11th International mycological congress. Mycological discoveries
for a better world. Abstract Book 273, San Juan, Puerto Rico
Quijada L, Tanney JB, Popov E, Johnston PR et al. 2020 – Cones, needles and wood: Micraspis
(Micraspidaceae, Micraspidales fam. et ord. nov.) speciation segregates by host plant
tissues. Fungal Systematics and Evolution 5, 99–112.
Raczka MF, Bush MB, Folcik AM, McMichael CH. 2016 – Sporormiella as a tool for detecting
the presence of large herbivores in the Neotropics. Biota Neotropica 16(1).
Rao AR. 1959 – Fungal remains from some Tertiary deposits of India. Palaeobotanist 7, 43–46.
Raper D, Bush M. 2009 – A test of Sporormiella representation as a predictor of megaherbivore
presence and abundance. Quaternary Research 71, 490-496.
Raper KB, Smith NR. 1939 – The growth of Dictyostelium discoideum on pathogentic bacteria.
Journal of Microbiology 38, 431–445.
Raper KB. 1984 – The Dictyostelids. Princeton University Press, United States.
Réblová M, Hubka V, Thureborn O, Lundberg J et al. 2016 – From the tunnels into de treetops:
new lineages of black yeasts from biofilm in the Stockholm Metro system and their relatives
among ant – associated fungi in the Chaetothyriales. PloS One 11, e0163396.
Réblová M, Miller AN, Réblová K, Štěpánek V 2018 – Phylogenetic classification and generic
delineation of Calyptosphaeria gen. nov., Lentomitella, Spadicoides and Torrentispora
(Sordariomycetes). Studies in Mycology 89, 1–62.
Réblová M, Štěpánek V. 2018 – Introducing the Rhamphoriaceae fam. nov. (Sordariomycetes),
two new genera and new life histories for taxa with Phaeoisaria–and Idriella–like
anamorphs. Mycologia 110, 750–770.

1447
Réblová M, Untereiner WA, Réblová K. 2013 – Novel evolutionary lineages revealed in the
Chaetothyriales (Fungi) based on multigene phylogenetic analyses and comparison of ITS
secondary structure. PloS ONE 8, e63547, 1–28.
Réblová M, Untereiner WA, Štěpánek V, Gams W. 2016 – Disentangling Phialophora section
Catenulatae: disposition of taxa with pigmented conidiophores and recognition of a new
subclass, Sclerococcomycetidae (Eurotiomycetes). Mycological Progress 16, 27–46.
Resl P, Schneider K, Westberg M, Printzen C et al. 2015 – Diagnostics for a troubled backbone:
testing topological hypotheses of trapelioid lichenized fungi in a large–scale phylogeny of
Ostropomycetidae (Lecanoromycetes). Fungal Diversity 73, 239–258.
Ride WDL, Cogger HG, Dupuis C, Kraus O et al. 1999 – International code of zoological
nomenclature. Fourth Edition. The Natural History Museum, London.
Riess K, Schön ME, Lutz M, Butin H et al. 2016 – On the evolutionary history of Uleiella
chilensis, a smut fungus parasite of Araucaria araucana in South America: Uleiellales ord.
nov. in Ustilaginomycetes. PloS one 11, e0147107.
Rikkinen J, Beimforde C, Seyfullah LJ, Perrichot V et al. 2016 – Resinogalea humboldtensis gen.
et sp. nov., a new resinicolous fungus from New Caledonia, placed in Bruceomycetaceae
fam. nova (Ascomycota). Annales Botanici Fennici 53, 205–215.
Rizzo L, Sutton DA, Wiederhold NP, Thompson EH et al. 2014 – Isolation and characterisation of
the fungus Spiromastix asexualis sp. nov. from discospondylitis in a German Shepherd dog,
and review of Spiromastix with the proposal of the new order Spiromastixales (Ascomycota).
Mycoses 57, 419–428.
Rollins AW, Landolt JC, Stephenson SL. 2010 – Dictyostelid cellular slime molds associated with
grasslands of the central and western United States. Mycologia 102, 996–1003.
Romeralo M, Baldauf S, Escalante R 2013 – Dictyostelids Evolution, Genomics and Cell Biology.
Springer, Berlin Heidelberg.
Romeralo M, Cavender JC, Landolt JC, Stephenson SL, Baldauf SL. 2011 – An expanded
phylogeny of social amoebas (Dictyostelia) shows increasing diversity and new
morphological patterns. BMC Evolutionary Biology 11, 84.
Romeralo M, Escalante R, Baldauf SL. 2012 – Evolution and diversity of dictyostelid social
Amoebae. Protist 63, 327–343.
Ronikier A, Halamski AT. 2018 – Is Myxomycetes (Amoebozoa) a truly ambiregnal group? A
major issue in protist nomenclature. Protist 169, 484–493.
Ropars J, Cruaud C, Lacoste S, Dupont J. 2012 – A taxonomic and ecological overview of cheese
fungi. International Journal of Food Microbiology 155, 199–210.
Rosendahl CO 1943 – Some fossil fungi from Minnesota. Bulletin of the Torrey Botanical Club
70, 126–138.
Rosling A, Cox F, Cruz-Martinez K, Ihrmark K et al. 2011 – Archaeorhizomycetes: unearthing an
ancient class of ubiquitous soil fungi. Science 333, 876–879.
Rossman AY, Cavan AW, Braun U, Castlebury LA et al. 2016 – Overlooked competing asexual
and sexually typifed generic names of Ascomycota with recommendations for their use or
protection. IMA fungus 7, 289–308.
Rossman AY, Crous PW, Hyde KD, Hawksworth DL et al. 2015 – Recommended names for
pleomorphic genera in Dothideomycetes. IMA fungus 6, 507–523.
Rupcic Z, Chepkirui C, Hernández-Restrepo M, Crous PW et al. 2018 – New nematicidal and
antimicrobial secondary metabolites from a new species in the new genus,
Pseudobambusicola thailandica. MycoKeys 33, 1–23.
Salgado-Salazar C, Beirn LA, Ismaiel A, Boehm MJ et al. 2018 – Clarireedia: A new fungal
genus comprising four pathogenic species responsible for dollar spot disease of turfgrass.
Fungal Biology 122, 761–773.
Samarakoon MC, Thongbai B, Hyde KD, Brönstrup M et al. 2020 – Elucidation of the life cycle
of the endophytic genus Muscodor and its transfer into the genus Induratia in Induratiaceae

1448
fam. nov., based on a polyphasic taxonomic approach. Fungal Diversity DOI:
10.1007/s13225-020-00443-9.
Sanders D, Borys KD, Kisa F, Rakowski SA et al. 2017 – Multiple dictyostelid species destroy
biofilms of Klebsiellaoxytoca and other gram negative species. Protist 168, 311–325.
Santamaria S, Enghoff H, Gruber J, Reboleira ASPS 2017 – First Laboulbeniales from
harvestmen: the new genus Opilionomyces. Phytotaxa 305, 285–292.
Santamaria S, Enghoff H, Reboleira ASPS. 2016 – Hidden biodiversity revealed by collections-
based research – Laboulbeniales in millipedes: genus Rickia. Phytotaxa 243, 101–127.
Saxena RK, Sarkar S. 1986 – Morphological study of Frasnacritetrus taugourdeau emend. from
the tertiary sediments of Himachal Pradesh, India. Review of Palaeobotany & Palynology
46, 209–225.
Saxena RK, Tripathi SKM. 2011 – Indian Fossil Fungi. Palaeobotanist 60, 1–208.
Saxena RK. 1991 – A catalogue of fossil plants from India – Part 5B. Tertiary fungi. Special
Publication, Birbal Sahni Institute of Palaeobotany, Lucknow. pp. 1–19.
Saxena RK. 2006 – A Catalogue of Tertiary Fungi from India (1989–2005). Special Publication,
Birbal Sahni Institute of Palaeobotany, Lucknow. pp. 1–37.
Scambler R, Niskanen T, Assyov B, Ainsworth AM et al. 2018 – Diversity of Chroogomphus
(Gomphidiaceae, Boletales) in Europe, and typification of C. rutilus. IMA fungus 9, 271–
290.
Schaap P, Winckler T, Nelson M, Alvarez-Curto E et al. 2006 – Molecular phylogeny and
evolution of morphology in the social amoebas. Science 14(5799), 661–3.
Schenck NC, Pérez Y. 1990 – Manual for identification of VA mycorrhizal fungi. Synergistic
Publications, Gainesville.
Schipper MAA, Samson RA. 1994 – Miscellaneous notes on Mucoraceae. Mycotaxon 50, 475–
491.
Schipper MAA. 1973 – A study on variability in Mucor hiemalis and related species. Studies in
Mycology 4, 1–40.
Schipper MAA. 1975 – Mucor mucedo, Mucor flavus and related species. Studies in Mycology 10,
1–33.
Schipper MAA. 1976 – On Mucor circinelloides, Mucor racemosus and related species. Studies in
Mycology 12, 1–40.
Schipper MAA. 1978 – On certain species of Mucor with a key to all accepted species. Studies in
Mycology 17, 1–69.
Schnepf E, Deichgräber G, Drebes G 1977 – Development and ultrastructure of the marine,
parasitic oomcete, Lagenisma coscinodisci (Lagenidiales): sexual reproduction. Canadian
Journal of Botany, 56, 1315–1325.
Schoch CL, Sung GH, López-Giráldez F, Townsend JP et al. 2009 – The Ascomycota Tree of Life:
A phylum-wide phylogeny clarifies the origin and evolution of fundamental reproductive
and ecological traits. Systematic Biology 58, 224–239
Schuld M, Madel G, Schmuck R. 1999 – Impact of Vairimorpha sp. (Microsporidia: Burenellidae)
on Trichogramma chilonis (Hymenoptera, Trichogrammatidae), a hymenopteran parasitoid
of the cabbage moth, Plutella xylostella (Lepidoptera, Yponomeutidae). Journal of
Invertebrate Pathology 74, 120–126.
Schüßler A, Schwarzott D, Walker C. 2001 – A new fungal phylum, the Glomeromycota:
phylogeny and evolution. Mycological Research 105, 1413–1421
Seifert K, Morgan-Jones G, Gams W, Kendrick B 2011 – The genera of hyphomycetes. CBS
Biodiversity Series 9, 1–997.
Senderskiy IV, Timofeev SA, Seliverstova EV, Pavlova OA et al. 2014 – Secretion of
Antonospora (Paranosema) locustae proteins into infected cells suggests an active role of
Microsporidia in the control of host programs and metabolic processes. PLoS ONE 9,
e93585.

1449
Sert HB, Sumbul H, Sterflinger K. 2007 – Microcolonial fungi from antique marbles in
Perge/Side/Termessos (Antalya/Turkey). Antonie van Leewenhoek 91, 217–227.
Shadwick LL, Spiegel FW, Shadwick JD, Brown MW et al. 2009 – Eumycetozoa = Amoebozoa?
SSU rDNA phylogeny of protosteloid slime molds and its significance for the amoebozoan
supergroup. PLoS One 4, e6754.
Sharma R, Kulkarni G, Sonawane MS. 2017 – Alanomyces, a new genus of Aplosporellaceae
based on four loci phylogeny. Phytotaxa 297, 168–175.
Sharma R, Xia X, Cano LM, Evangelisti E, Kemen E et al (2015) Genome analyses of the
sunflower pathogen Plasmopara halstedii provide insights into effector evolution in downy
mildews and Phytophthora. BMC Genomics 16, 741.
Shchepin O, Schnittler M, Dagamac N, Leontyev D et al. 2019 – Unexplored diversity of
microscopic myxomycetes: evidence from environmental DNA. Plant Ecology & Evolution,
152, 499–506.
Sheikh S, Thulin M, Cavender JC, Escalante R et al. 2018 – A new classification of the
Dictyostelids. Protist 169, 1–28.
Shen XX, Opulente DA, Kominek J, Zhou X et al. 2018 – Tempo and mode of genome evolution
in the budding yeast subphylum. Cell 75, 1533-1545.e20
Shen XX, Zhou X, Kominek J, Kurtzman CP et al. 2016 – Reconstructing the backbone of the
Saccharomycotina yeast phylogeny using genome–scale data. G3 (Bethesda) 6, 3927–3939.
Shi WP, Zheng X, Jia WT, Li AM et al. 2018 – Horizontal transmission of Paranosema locustae
(Microsporidia) in grass hopper populations via predatory natural enemies. Pest
Management Science 74, 2589–2593.
Shoemaker RA, Babcock CE. 1992 – Applanodictyosporous Pleosporales: Clathrospora,
Comoclathris, Graphyllium, Macrospora, and Platysporoides. Canadian Journal of Botany
70, 1617–1658.
Sieverding E, Silva GA, Berndt R, Oehl F. 2014 – Rhizoglomus, a new genus in the Glomeraceae.
Mycotaxon 129, 373–386.
Silva GA, Maia LC, Oehl F. 2012 – Phylogenetic systematics of the Gigasporales. Mycotaxon
122, 207–220.
Silva RM, Oliveira RJ, Bezerra JD, Bezerra JL et al. 2019 – Bifusisporella sorghi gen. et sp. nov.
(Magnaporthaceae) to accommodate an endophytic fungus from Brazil. Mycological
Progress 18, 847–854.
Simakova AV, Pankova TF, Tokarev YS, Issi IV. 2005 – New genus of microsporidia Senoma
gen. n. with type species Senoma globulifera comb. n. (syn. Issia globulifera Issi, Pankova,
1983 from malaria mosquito Anopheles messae Fall. Protistology 4, 134–145.
Simmons EG. 2007 – Alternaria: an identification manual. CBS Biodiversity Series 6. 775 pp.
Singh BN. 1947 – Studies on soil Acrasieae. 1. Distribution of species of Dictyostelium in soils of
Great Britain and the effects of bacteria on their development. Journal of General
Microbiology 1.
Singh G, Dal Grande F, Divakar PK, Otte J et al. 2015 – Coalescent-based species delimitation
approach uncovers high cryptic diversity in the cosmopolitan lichen-forming fungal genus
Protoparmelia (Lecanorales, Ascomycota). Plos One 10, e0124625.
Slippers B, Boissin E, Phillips AJL, Groenewald JZ et al. 2013 – Phylogenetic lineages in the
Botryosphaeriales: A systematic and evolutionary framework. Studies in Mycology 76, 31–
49.
Smith ME, Amses KR, Elliott TF, Obase K et al. 2015 – New sequestrate fungi from guyana:
Jimtrappea guyanensis gen. sp. nov., Castellanea pakaraimophila gen. sp. nov, and
Costatisporus cyanescens gen. sp. nov. (Boletaceae, Boletales). IMA fungus 6, 297–317.
Sokolova Y, Pelin A, Hawke J, Corradi N. 2015 – Morphology and phylogeny of
Agmasomapenaei (Microsporidia) from the type host, Litopenaeus setiferus, and the
typelocality, Louisiana, USA. International Journal for Parasitology 45, 1–16.

1450
Sokolova YK, Issi IV, Voronin VN. 2018 – Annotated list of species of the Microsporidia
described in the Former Soviet Union and Russia in 20th century 1967–2000. Protistology
12, 12–37.
Sokolova YY, Bowers LC, Alvarez X, Didier ES. 2019 – Encephalitozoon cuniculi and
Vittaforma corneae (Phylum Microsporidia) inhibit staurosporine–induced apoptosisin
human THP–1 macrophages in vitro. Parasitology 146, 569–579.
Sokolova YY, Paskerova GG, Rotari YM, Nassonova ES et al. 2013 – Fine structure of
Metchnikovella incurvata Caullery & Mesnil 1914 (Microsporidia), a hyperparasite of
gregarines Polyrhabdina sp. from the polychaete Pygospioelegans. Parasitology 140, 855–
867.
Song J, Liang JF, Mehrabi-Koushki M, Krisai-Greilhuber I, Ali B et al. 2019 – Fungal
Systematics and Evolution 5. Sydowia 71, 141–245.
Song Z, Li G, Cao L, Luo H et al. 1989 – Early Tertiary sporo–pollen assemblages from the
Dongpu region. Edited by Research Institute of Exploration and Development, Zhongyuan
Petroleum Exploration Bureau, Nanjing Institute of Geology and Palaeontology, Academia
Sinica, 192 p.
Spatafora JW, Chang Y, Benny GL, Lazarus K et al. 2016 – A phylum–level phylogenetic
classification of zygomycete fungi based on genome–scale data. Mycologia 108, 1028–1046.
Species Fungorum 2019 http://www.speciesfungorum.org/Names/SynSpecies.asp
Stefani FOP, Jones RH, May TW. 2014 – Concordance of seven gene genealogies compared to
phenotypic data reveals multiple cryptic species in Australian dermocyboid Cortinarius
(Agaricales). Molecular Phylogenetics & Evolution 71, 249–260.
Stentiford GD, Feist SW, Stone DM, Bateman KS et al. 2013 – Microsporidia: diverse, dynamic,
and emergent pathogens in aquatic systems. Trends in Parasitology 29, 567–578.
Stentiford GD, Ramilo A, Abollo E, Kerr R et al. 2017 – Hypersporaaquatica n. gn., n. sp.
(microsporidia), hyperparasitic in Marteiliacochillia (Paramyxida), is closely related to
crustacean–infectingmicrospordian taxa. Parasitology 144, 186–199.
Stentiford GD, Ross S, Minardi D, Feist SW et al. 2018 – Evidence for trophic transfer of
Inodosporus octospora and Ovipleistophora arlo n. sp. (microsporidia) between crustacean
and fish hosts. Parasitology 145, 1105–1117.
Stephenson SL, Landolt JC. 1992 – Vertebrates as vectors of cellular slime moulds in temperate
forests. Mycological Research 96, 670–672.
Stephenson SL, Landolt JC. 1998 – Dictyostelid cellular slime molds in canopy soils of tropical
forests. Biotropica 30, 657–661.
Stephenson SL, Landolt JC. 2011 – Dictyostelids from aerial “canopy soil” microhabitats. Fungal
Ecology 4, 191–195.
Stephenson SL, Rajguru SN. 2010 – Dictyostelid cellular slime moulds in agricultural soils.
Mycosphere 1, 333–336.
Subrahamanyam A. 1983 – Studies on themomycology. Mucor thermo–hyalospora sp. nov.
Bibliotheca Mycologica 91, 421–423.
Subramanian CV. 1971 – Hyphomycetes, an account of Indian species, except Cercosporae.
Indian Council of Agricultural Research, New Delhi, 930 p.
Sudová R, Sýkorová Z, Rydlová J, Čtvrtlíková M, Oehl F. 2015 – Rhizoglomus melanum, a new
arbuscular mycorrhizal fungal species associated with submerged plants in freshwater lake
Avsjøen in Norway. Mycologial Progress 14, 9
Suija A, de los Ríos A, Pérez-Ortega S. 2015 – A molecular reappraisal of Abrothallus species
growing on lichens of the order Peltigerales. Phytotaxa, 195, 201−226.
Suija A, Kaasalainen U, Kirika PM, Rikkinen J. 2018 – Taitaia, a novel lichenicolous fungus in
tropical montane forests in Kenya (East Africa). Lichenologist 50, 173–184.
Suija A, van den Boom PPG, Zimmermann E, Zhurbenko MP et al. 2017 – Lichenicolous species
of Hainesia belong to Phacidiales (Leotiomycetes) and are included in an extended concept
of Epithamnolia. Mycologia 109, 882–889.

1451
Sumbali G. (ed. Johri BM) 2005 – The fungi. ISBN 978–1–84265–153–7
Suthers HB. 1985 – Ground–feeding migratory songbirds as cellular slime mold distribution
vectors. Oecologia 65, 526–530.
Tanaka K, Hirayama K, Yonezawa H, Sato G et al. 2015 – Revision of the Massarineae
(Pleosporales, Dothideomycetes). Studies in Mycology 82, 75–136.
Tanney JB, Douglas B, Seifert KA. 2016 – Sexual and asexual states of some endophytic
Phialocephala species of Picea. Mycologia 108, 255–280.
Tanney JB, Seifert KA. 2020 – Mollisiaceae: An overlooked lineage of diverse endophytes,
Studies in Mycology, https://doi.org/10.1016/j.simyco.2020.02.005.
Taugourdeau P. 1968 – Sur un curieux microfossile incertae sedis du Frasnien du Boulonnais,
Frasnacritetrus nov. gen. (Acritarche). Cahiers de Micropaléontologie, série 1, no. 10
(Archives originales du Centre de Documentation du C.N.R.S. no. 452, 1–4.
Tavares II. 1985 – Laboulbeniales (Fungi, Ascomycetes). Mycologia Memoir 9, 1–627.
Tedersoo L, Sánchez–Ramírez S, Koljalg U et al. 2018 – High–level classification of the Fungi
and a tool for evolutionary ecological analyses. Fungal Diversity 90, 135–159.
Teterevnikova-Babaian DN, Taslakhchian MG. 1973 – New data on fossil fungal spores in
Armenia. Mycology and Phytology 4, 159–164.
Thambugala KM, Wanasinghe DN, Phillips AJ, Camporesi E et al. 2017 – Mycosphere notes 1-
50: grass (Poaceae) inhabiting Dothideomycetes. Mycosphere 8, 697–796.
Thines M, Nam B, Nigrelli L, Beakes G, Kraberg A. 2015 – The diatom parasite Lagenisma
coscinodisci (Lagenismatales, Oomycota) is an early diverging lineage of the
Saprolegniomycetes. Mycological Progress 14, 75.
Thines M, Crous PW, Aime MC, Aoki T et al. 2018 – Ten reasons why a sequence-based
nomenclature is not useful for fungi anytime soon. IMA fungus 9, 177–183.
Thines M. 2019 – An evolutionary framework for host shifts-jumping ships for survival. New
Phytologist 224, 605–617.
Tice AK, Shadwick LL, Fiore-Donno AM, Geisen S et al. 2016 – Expansion of the molecular and
morphological diversity of Acanthamoebidae (Centramoebida, Amoebozoa) and
identification of a novel life cycle type within the group. Biology Direct 11, 69–90.
Tokarev YS, Voronin VN, Seliverstova EV, Dolgikh VV et al. 2010 – Ultrastructure and
molecular phylogeny of Anisofilariata chironomi sp. n. g. n. (Microsporidia: Terresporidia),
a microsporidian parasite of Chironomus plumosus L. (Diptera: Chironomidae).
Parasitology Research 106, 39–46.
Tokarev YS, Voronin VN, Seliverstova EV, Grushetskaya TA et al. 2012 – Ultrastructure and
molecular phylogenetics of Helmichialacustris, a microsporidium with an uncoiled isofilar
polar filament. Parasitology Research 110, 1201–1208.
Tonka T, Weiser JJr, Weiser J. 2010 – Budding: a new stage in the development of Chytridiopsis
typographi (Zygomycetes: Microsporidia). Journal of Invertebrate Pathology 104, 17–22.
Torruella G, Grau–Bové X, Moreira D, Karpov SA et al. 2018 – Global transcriptome analysis of
the aphelid Paraphelidium tribonemae supports the phagotrophic origin of fungi.
Communications Biology 1, 231.
Trakunyingcharoen T, Lombard L, Groenewald JZ, Cheewangkoon R et al. 2014 – Mycoparasitic
species of Sphaerellopsis, and allied lichenicolous and other genera. IMA fungus 5, 391–
414.
Truong C, Mujic AB, Healy R, Kuhar F et al. 2017 – How to know the fungi: combining field
inventories and DNA–barcoding to document fungal diversity. New Phytologist 214, 913–
919.
Tsaousis AD, Kunji ER, Goldberg AV et al. 2008 – A novel route for ATP acquisition by the
remnant mitochondria of Encephalitozoon cuniculi. Nature 453(7194), 553–556.
Tsuneda A, Davey ML, Tsuneda I, Hudgins A et al. 2011a – Endophoma, a new didymellaceous
endoconidial genus from bat – cave soil. Mycologia 103, 1146–1155.

1452
Tsuneda A, Hambleton S, Currah RS. 2011b – The anamorph genus Knufia and its
phylogenetically allied species in Coniosporium, Sarcinomyces and Phaeococcomyces.
Botany 89, 523–536.
Turland NJ, Wiersema JH, Barrie FR et al. 2018 – International Code of Nomenclature for algae
fungi and plants (Shenzhen Code) adopted by the Nineteenth International Botanical
Congress Shenzhen, China, July 2017. Regnum Vegetabile, 159 Koeltz Botanical Books
Glashütten.
Turrini A, Saran M, Giovannetti M, Oehl F. 2018 – Rhizoglomus venetianum, a new arbuscular
mycorrhizal fungal species from a heavy metal contaminated site, downtown Venice in
Italy. Mycological Progress 17, 1213–1224.
Van der Hammen T. 1954 – El desarrollo de la flora Colombiana en los periodos geológicos. I.
Maestrichtiano hasta Terciario más inferior. Boletín Geológico 2, 49–106.
Van Vooren N, Lindemann, Healy R. 2017 – Emendation of the genus Tricharina (Pezizales)
based on phylogenetic, morphological and ecological data. Ascomycete.org 9, 101–123.
Vavra J, Larsson JIR. 2014 – Structure of Microsporidia. In: Weiss LM, Becnel JJ (eds)
Microsporidia: Pathogens of Opportunity. Wiley-Blackwell, pp 1–70
Vavra J, Lukes J. 2013 – Microsporidia and ‘the art of living together’. Advances in Parasitology
82, 253–319.
Vellinga EC, Sysouphanthong P, Hyde KD. 2011 – The family Agaricaceae: phylogenies and two
new white–spored genera. Mycologia 103, 494–509.
Venkatachala BS, Kar RK 1969 – Palynology of the Tertiary sediments in Kutch—2. Epiphyllous
fungal remains from the borehole no. 14. Palaeobotanist 17, 179–183.
Vivier E 1975 – The microsporidia of the protozoa. Protistologica 9, 345–361.
Vizzini A, Contu M, Ercole E. 2011 – Musumecia gen. nov. in the Tricholomatoid clade
(Basidiomycota, Agaricales) related to Pseudoclitocybe. Nordic Journal of Botany 29, 734–
740.
Vlk W 1939 – Über die Geißelstruktur der Saprolegniaceenschwärmer. Archiv für Protistenkunde
92, 157–160.
Voglmayr H, Aguirre-Hudson MB, Wagner HG, Tello S et al. 2019 – Lichens or endophytes? The
enigmatic genus Leptosillia in the Leptosilliaceae fam. nov. (Xylariales), and Furfurella gen.
nov. (Delonicicolaceae). Persoonia 42, 228–260.
Voglmayr H, Clémençon H. 2016 – Identification and taxonomic position of two mucoralean
endoparasites of Hysterangium (Basidiomycota) based on molecular and morphological
data. Mycological Progress 15, 1–17.
Voglmayr H, Friebes G, Gardiennet A, Jaklitsch WM. 2018 – Barrmaelia and Entosordaria in
Barrmaeliaceae (fam. nov., Xylariales) and critical notes on Anthostomella-like genera
based on multigene phylogenies. Mycological Progress 17, 155–177.
Voglmayr H, Pintos A. 2018 – Three new species of Stigmatodiscus from Mallorca (Spain).
Mycological Progress 17, 1189–1201.
Vossbrinck CR, Debrunner-Vossbrinck BA, Weiss LM. 2014 – Molecular phylogeny of the
Microsporidia. In: Weiss LM, Becnel JJ (eds) Microsporidia: Pathogens of Opportunity.
Wiley–Blackwell, pp 203–220.
Vossbrinck CR, Debrunner-Vossbrinck BA. 2005 – Molecular phylogeny of the Microsporidia:
ecological, ultrastructural and taxonomic considerations. Folia Parasitologica 52, 131–142.
Vu D, Groenewald M, de Vries M, Gehrmann T et al. 2019 – Large–scale generation and analysis
of filamentous fungal DNA barcodes boosts coverage for kingdom Fungi and reveals
thresholds for fungal species and higher taxon delimitation. Studies in Mycology 92, 135–
154.
Wanasinghe DN, Phukhamsakda C, Hyde KD, Jeewon R et al. 2018 – Fungal diversity notes 709–
839: taxonomic and phylogenetic contributions to fungal taxa with an emphasis on fungi on
Rosaceae. Fungal Diversity 89, 1–236.

1453
Wang HK, Aptroot A, Crous PW, Hyde KD et al. 2007 –. The polyphyletic nature of
Pleosporales: an example from Massariosphaeria based on rDNA and RBP2 gene
phylogenies. Mycological Research 111, 1268–1276.
Wang QM, Groenewald M, Takashima M, Theelen B et al. 2015a – Phylogeny of yeasts and
related filamentous fungi within Pucciniomycotina determined from multigene sequence
analyses. Studies in Mycology 8127–53.
Wang QM, Theelen B, Groenewald M, Bai FY et al. 2014a – Moniliellomycetes and
Malasseziomycetes, two new classes in Ustilaginomycotina. Persoonia 33, 41–47.
Wang QM, Yurkov AM, Göker M, Lumbsch HT et al. 2015b – Phylogenetic classification of
yeasts and related taxa within Pucciniomycotina. Studies in Mycology 81, 149–189.
Wang XW, Yang FY, Meijer M, Kraak B et al. 2018 – Redefining Humicola sensu stricto and
related genera in the Chaetomiaceae. Studies in Mycology 93, 65–153.
Watanabe T. 1994 – Two new species of homothallic Mucor in Japan. Mycologia 86, 691–695.
Wei X, Schmitt I, Hodkinson BP, Flakus A et al. 2017 – Circumscription of the genus Lepra, a
recently resurrected genus to accommodate the “Variolaria” – group of Pertusaria sensu
lato (Pertusariales, Ascomycota). PLoS ONE 12, e0180284.
Weir A, Blackwell M. 2001 – Molecular data support the Laboulbeniales as a separate class of
Ascomycota, Laboulbeniomycetes. Mycological Research 105, 715–722.
Weir A, Hammond PM. 1997 – Laboulbeniales on beetles: host utilization patterns and species
richness of the parasites. Biodiversity & Conservation 6, 701–719.
Wendt L, Sir EB, Kuhnert E, Heitkämper S et al. 2018 – Resurrection and emendation of the
Hypoxylaceae, recognised from a multi-gene phylogeny of the Xylariales. Mycological
Progress 17, 115–154.
Westphalen MC, Rajchenberg M, Tomšovský M, Gugliotta AM. 2018 – A re–evaluation of
Neotropical Junghuhnia s. lat. (Polyporales, Basidiomycota) based on morphological and
multigene analyses. Persoonia 41, 130–141.
Wijayawardene DNN, McKenzie EHC, Hyde KD. 2012 – Towards incorporating anamorphic
fungi in a natural classification checklist and notes for 2011. Mycosphere 3, 157–228.
Wijayawardene NN, Hyde KD, Lumbsch HT, Liu JK et al. 2018a – Outline of Ascomycota: 2017
Fungal Diversity 88, 167–263.
Wijayawardene NN, Hyde KD, Rajeshkumar KC, Hawksworth DL et al. 2017a – Notes for genera:
Ascomycota. Fungal Diversity 86, 1–594.
Wijayawardene NN, Hyde KD, Tibpromma S, Wanasinghe DN et al. 2017b – Towards
incorporating asexual fungi in a natural classification: checklist and notes 2012–2016.
Mycosphere 8, 1457–1554.
Wijayawardene NN, Hyde KD, Wanasinghe DN, Papizadeh M et al. 2016 – Taxonomy and
phylogeny of dematiaceous coelomycetes. Fungal Diversity 77, 1–316.
Wijayawardene NN, Pawłowska J, Letcher PM, Kirk PM et al. 2018b – Notes for genera: basal
clades of Fungi (including Aphelidiomycota, Basidiobolomycota, Blastocladiomycota,
Calcarisporiellomycota, Caulochytriomycota, Chytridiomycota, Entomophthoromycota,
Glomeromycota, Kickxellomycota, Monoblepharomycota, Mortierellomycota,
Mucoromycota, Neocallimastigomycota, Olpidiomycota, Rozellomycota and
Zoopagomycota). Fungal Diversity 92, 43–129.
Williams BA, Hirt RP, Lucocq JM, Embley TM. 2002 – A mitochondrial remnant in the
microsporidian Trachipleistophor hominis. Nature 418(6900), 865–869.
Wittstein K, Cordsmeier A, Lambert C, Wendt L et al. 2020 – Identification of Rosellinia species
as producers of cyclodepsipeptide PF1022 A and resurrection of the genus Dematophora as
inferred from polythetic taxonomy. Studies in Mycology 96, 1–16.
Woudenberg JHC, Seidl MF, Groenewald JZ, De Vries M et al. 2015 – Alternaria section
Alternaria: Species, formae speciales or pathotypes? Studies in Mycology 82, 1–21.
Woudenberg JHC, Truter M, Groenewald JZ, Crous PW. 2014 – Large–spored Alternaria
pathogens in section Porri disentangled. Studies in Mycology 79, 1–47.

1454
Wu G, Zhao K, Li YC, Zeng NK et al. 2016 – Four new genera of the fungal family Boletaceae.
Fungal Diversity 81, 1–24
Wu ZQ, Xu TM, Shen S, Liu XF et al. 2018 – Elaphroporia ailaoshanensis gen. et sp. nov. in
Polyporales (Basidiomycota). MycoKeys 29, 81–95.
Wyka SA, Broders KD. 2016 – The new family Septorioideaceae, within the Botryosphaeriales
and Septorioides strobi as a new species associated with needle defoliation of Pinus strobus
in the United States. Fungal Biology 120, 1030–1040.
Xia JW, Ma YR, Li Z, Zhang XG. 2017 – Acrodictys–like wood decay fungi from southern
China, with two new families Acrodictyaceae and Junewangiaceae. Scientific Reports 7
(no. 7888), 1–21.
Xiao Y, Wen T, Hongsanan S, Jeewon R et al. 2018 – Multigene phylogenetics of
Polycephalomyces (Ophiocordycipitaceae, Hypocreales), with two new species from
Thailand. Scientific Reports 8, 18087.
Xu Y, Weiss LM. 2005 – The microsporidian polar tube: a highly specialised invasionorganelle.
International Journal for Parasitology 35, 941–953.
Yadav DR, Kim SW, Adhikari M, Um YH et al. 2015 – Three new records of Mortierella species
isolated from crop field soil in Korea. Mycobiology 43, 203–209.
Yang CL, Xu XL, Wanasinghe DN, Jeewon R et al. 2019 – Neostagonosporella sichuanensis gen.
et sp. nov. (Phaeosphaeriaceae, Pleosporales) on Phyllostachys heteroclada (Poaceae)
from Sichuan Province, China. MycoKeys 46, 119–150.
Yang H, Hyde KD, Karunarathna SC, Deng C et al. 2018a – New species of Camptophora and
Cyphellophora from China, and first report of sexual morphs for these genera. Phytotaxa
343, 149–159.
Yang J, Liu JK, Hyde KD, Jones EG et al. 2019 – Aquimonospora tratensis gen. et sp. nov.
(Diaporthomycetidae, Sordariomycetes), a new lineage from a freshwater habitat in
Thailand. Phytotaxa 397, 146–158.
Yang Q, Fan XL, Du Z, Tian CM. 2018b – Diaporthosporellaceae, a novel family of
Diaporthales (Sordariomycetes, Ascomycota). Mycoscience 59, 229–235.
Yang T, Groenewald JZ, Cheewangkoon R, Jami F et al. 2017 – Families, genera, and species of
Botryosphaeriales. Fungal Biology 121, 322–346.
Yoshida K, Schuenemann VJ, Cano LM, Pais M et al. 2013 – The rise and fall of the
Phytophthora infestans lineage that triggered the Irish potato famine. eLife 2, e00731.
Yu X, Hoyle RL, Guo F, Ratliff CM et al. 2019 – A Vavraia–like microsporidium as the cause of
deadly infection in threatenedand endangered Eurycea salamanders in the United States.
Parasites & Vectors 12, 108.
Yu XD, Dong W, Bhat DJ, Boonmee S et al. 2018 – Cylindroconidiis aquaticus gen. et sp nov., a
new lineage of aquatic hyphomycetes in Sclerococcaceae (Eurotiomycetes). Phytotaxa 372,
79–87.
Yuan HS, Qin WM. 2018 – Multiple genes phylogeny and morphological characters reveal
Dextrinoporus aquaticus gen. et sp. nov. (Polyporales, Basidiomycota) from southern China.
Mycological Progress 17, 773–780.
Zalar P, Hennebert GL, Gunde–Cimerman N, Cimerman A. 1997 – Mucor troglophilus, a new
species from cave crickets. Mycotaxon 65, 507–516.
Zamora JC, Svensson M, Kirschner R, Olariaga I et al. 2018 – Considerations and consequences
of allowing DNA sequence data as types of fungal taxa. IMA fungus 9, 167–175.
Zhang ZF, Liu F, Zhou X, Liu XZ et al. 2017 – Culturable mycobiota from Karst caves in China,
with descriptions of 20 new species. Persoonia 39, 1–31.
Zhao RL, Li GJ, Sanchez-Ramirez S, Stata M et al. 2017c – A six–gene phylogenetic overview of
Basidiomycota and allied phyla with estimated divergence times of higher taxa and a
phyloproteomics perspective. Fungal Diversity 84, 43–74.

1455
Zhao YJ, Hosaka K, Hosoya T. 2016 – Taxonomic re-evaluation of the genus Lambertella
(Rutstroemiaceae, Helotiales) and allied stroma-forming fungi. Mycological Progress 15,
1215–1228.
Zhu HY, Pan M, Bonthond G, Tian CM, Fan XL. 2019 – Diaporthalean fungi associated with
canker and dieback of trees from Mount Dongling in Beijing, China. Mycokeys 59, 67–94.
Zitouni-Haouar FEH, Alvarado P, Sbissi I, Boudabous A et al. 2015 – Contrasted genetic diversity,
relevance of climate and host plants, and comments on the taxonomic problems of the genus
Picoa (Pyronemataceae, Pezizales). PLoS ONE 10, e0138513.
Zmitrovich IV. 2018 – Conspectus systematis Polyporacearum v. 1.0. Folia Cryptogamica
Petropolitana 6, 3–145.

1456

You might also like