Mycological Contributions of Rolf Singer

UNIVERSITY OF
ILLINOIS LIBRARY
AT URBANA-CHAMPAIGN
BIOLOGY
'
2(101
SER
Vlycological Contributions of Rolf Singer:

Field Itinerary, Index to New Taxa,
ind List of Publications
Gregory M. Mueller and Qiuxin Wu, Editors
February 28, 1997

Publication 1482
PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY

utors to Ficldiana
Referee '
' n full. V
Vol. 2. Tl
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IHust e text (not
This paper meets the requirements of ANSI/NISO Z39.48-1992 (Permanence of Paper).

FIELDIANA
Botany
NEW SERIES, NO. 38
Mycological Contributions of Rolf Singer:

Field Itinerary, Index to New Taxa,
and List of Publications
Gregory M. Mueller and Qiuxin Wu, Editors

Department of Botany
Field Museum of Natural History
Roosevelt Road at Lake Shore Drive
Chicago, Illinois 60605-2496
T>iE LIBRARY OF THE
1 9 1997
Accepted December 6, 1995

Published February 28, 1997
Publication 1482
PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY

1997 Field Museum of Natural History
ISSN 0015-0746
PRINTED IN THE UNITED STATES OF AMERICA
Contents
Introduction and Acknowledgments v

Contributors vi
1 .
Summary of Rolf Singer's Contributions to Mycology 1
Gregory M. Mueller and Qiuxin Wu

2. The Field Itinerary of Rolf Singer 9
Betty A. Strack and Gregory M. Mueller
3. An Index to Genera, Species, and Infraspecific Taxa of 13
Basidiomycete Fungi Described by Rolf Singer
Qiuxin Wu, Gregory M. Mueller, and Clark L. Ovrebo
4. The Publications of Rolf Singer 101
Gregory M. Mueller, Qiuxin Wu, James H. Ginns, and
Scott A. Redhead
Index to Genera 113
Index to Species 115
111
Illustrations
1. Rolf Singer 2
2. Books and bound publications of Rolf Singer 3
3. Number of publications of Rolf Singer by age 3
4. Genera in which Singer and associates described at least 40

new taxa 3
5. Geographic distribution of Singer's type specimens 4
6. Herbaria with large holdings of Singer's type specimens 4
Tables
1. List of new genera published by Rolf Singer 5
2. Type localities and numbers of Singer's new taxa 6
3. Herbaria with holdings of Singer's type specimens 7
4. Rolf Singer's fieldwork, by country and year 1 1
5. Itinerary of Rolf Singer's fieldwork since 1941 11
IV
Introduction and Acknowledgments
Singer, 1906-1994, was one of the most influential figures in the

Rolf
history of mycology. This volume summarizes many of his con-
tributions to the study of fungi. Chapter 1 provides information on his
research program, and Chapter 2 presents his field itinerary. Much of this
volume consists of an index to all of the genera, species, and infraspecific
taxa described by Singer (2,452 specific and infraspecific epithets distrib-
uted in 246 genera). This index includes information on the type specimen,
place of deposition, and a literature citation for the type description. The
index is followed by a bibliography of his 439 publications.
The authors wish to extend their heartfelt thanks to the following people:
Michael Huft for creating the program to convert our dBase records into
WordPerfect files; Gail Kushino for help in searching for literature, proof-
reading, photocopying, and other miscellaneous but essential tasks; Eliz-
abeth Moore for keying in data and helping assemble the field itinerary;
Martha Singer and Meinhard Moser for checking the bibliography; and the
librarians at the Field Museum and the Gray and Farlow libraries at Har-
vard University. We also wish to thank the three anonymous reviewers
whose careful reading of this manuscript greatly improved its quality.
Contributors
Gregory M. Mueller Department of Botany, Field Museum of Natural History,

Roosevelt Road at Lake Shore Drive, Chicago, Illinois 60605-2496
Qiuxin Wu Department of Botany, Field Museum of Natural History, Roosevelt

Road at Lake Shore Drive, Chicago, Illinois 60605-2496
Betty A. Strack Department of Botany, Field Museum of Natural History, Roose-

velt Roadat Lake Shore Drive, Chicago, Illinois 60605-2496
Clark L.Ovrebo Department of Botany, University of Central Oklahoma, Ed-

mond, Oklahoma 73034-0177
James H. Ginns Crop Protection Program, Eastern Cereal and Oilseed Research
Centre, Research Branch, Agriculture and Agri-Food Canada, Ottawa,
Ontario, K1A OC6
Scott A. Redhead Crop Protection Program, Eastern Cereal and Oilseed Re-
search Centre, Research Branch, Agriculture and Agri-Food Canada,
Ottawa, Ontario, K1A OC6
VI
Summary of Rolf Singer's Contributions to Mycology
Gregory M. Mueller
Qiuxin Wu
Singer, 1906-1994 (Fig. 1), was one of vided him with a unique perspective. He devel-
Rolfthe most influential figures in the history of oped an unequaled, worldwide knowledge of the
mycology. This brief chapter distills the collec- Agaricales, and this, coupled with his keen mind
tions of facts presented in the rest of this volume and sharp eye, made him uniquely qualified to
in an attempt to synthesize the impact that Rolf influence mycology.
Singer had on our knowledge of fungi. Two in-
teresting works by Martha Singer, Rolf's wife and

companion of 60 years, provide additional infor- Systematics and Nomenclature
mation on his life (M. Singer, 1979, 1984). Sev-
eral memorials on Rolf Singer have also been
Singer is probably best known for his impact
published Mueller, 1994, 1995; M. Singer,
(e.g., on the systematics and nomenclature of Agari-
1994). These papers provide the historical back- cales and related taxa. His monumental The Agar-
drop to the discussions presented below. icales in Modern Taxonomy (Singer, 1986a) has
While the bulk of Singer's papers dealt with the appeared in four editions and is almost universally
systematics of Agaricales and related taxa, he also used as the "accepted" classification of mush-
profoundly influenced our knowledge of myco- rooms and their relatives. His monographs and re-
geography, fungal ecology, and ethnomycology. visions are often the only modern treatment for
Contributions to these four topics make up the
many genera.
vast majority of his 439 papers and books (Fig.
Singer's role in documenting fungal biodiver-
2). However, he also published on other topics, sity is also unequaled, and he revolutionized what
including fossil fungi (Singer, 1957e, 1958J, we know about Agaricales in South and Central
1990f), polypores (Singer, 194 la, 1943d, 1944b), America. Seventy-four of the 230 genera included
human pathogens (Singer, 1955g, 1958q), and in the fourth edition of The Agaricales in Modern
mountaineering (Singer, 1942e, f)- (Complete ci- Taxonomy, or nearly a third of the accepted gen-
tations to the papers cited here may be found in era, were proposed by Singer and his colleagues.
Chapter 4.) Singer's first five papers were pub- He also proposed genera in families not treated in
lished in 1922, when he was 16 years old. He The Agaricales in Modern Taxonomy, bringing
continued publishing at an impressive pace for the the total new genera to 82 (Table 1 ). Additionally,
next 70 years (Fig. 3). he published 2,452 specific and infraspecific epi-
During his career, Singer held academic posi- thets distributed in 246 genera (Fig. 4). Type lo-
tions on three continents and traveled extensively calities for these new taxa occur in 56 countries
throughout the Americas, Europe, and parts of or possessions (Table 2). However, 85% of them
Asia. He also made one trip to North Africa. A were originally described from the Americas, with
detailed itinerary of his collecting expeditions, Argentina, Bolivia, Brazil, Chile, Colombia, Mex-
along with a listing of the countries in which he ico,and the United States together accounting for
undertook fieldwork, appears in Chapter 2. These 75% (Fig. 5). Holotypes of these new taxa are
travels left a lasting impression on Singer and pro- housed in 40 herbaria throughout the world (Table
Singer's Contributions to Mycology 1

Figure 1 . Rolf Singer proofreading manuscript pages of the 4th edition of The Agaricales in Modern Taxonomy
in his office at the Field Museum of Natural History, circa 1985. (Courtesy of Field Museum of Natural History.)
3). Most of the specimens, however, can be found were often the first, or among the first, investi-
in a relatively few herbaria (Fig. 6). Chapter 3 gations on the subject. He and his colleagues pro-
consists of an index to the genera, species, and duced ground-breaking studies on the occurrence
infraspecific taxa described by Singer. This index and importance of mycorrhizae in various forest
includes information on the type specimen, place types in temperate and tropical South America
of deposition, and a literature citation for the type (e.g., Singer, 1959J, 1960k, 1963i, 1964g, 1971c,
description. 1983f, 1989d). His work along the Amazon and
Singer also influenced the systematics com- Rio Negro in Brazil helped document the different
munity through his work on botanical nomencla- forest types in the region and the role of fungi in
ture. He attended most of the International Botan- the health and functioning of these systems
ical Congresses from the 1950s on, and he was a (1978e, 1984b, c, 1986c, 1988c). He also carried
out one of the detailed
long-standing member of the Nomenclature Com- community
first analysis
mittee. His publications on the subject ranged from studies on mushrooms Americas (Singer,
in the
proposals for conservation of names (e.g., Singer, 1965i). Although he published few papers dealing
1946b, 1961j, 1967g) to major discussions of the solely with biogeography (e.g., Singer, 1955h,
implications of rule changes (e.g., 1948e, 19501, 1967f, 1988f), he included information on bio-
1960g, 1965g, 1983d, 1984e, 1986b, f, 1987d). geography in many of his monographs and re-
gional treatments.
Ecology Ethnomycology and Mushroom

Cultivation
Singer's contributions to our
knowledge of the
biogeography and ecology of fungi were also ex- Singer's mycological interests also included
tremely significant. His publications in these areas hallucinogenic and edible fungi. He played an ac-
Chapter One
Figure 2. Books and bound publications of Rolf Singer, as displayed during his 80th birthday testimonial dinner
at the Field Museum of Natural History in 1986. (Courtesy of Field Museum of Natural History.)
Pluteus
Mycena 1411
Marasmius 2971
(arasmlellus
Hydcopus
Galerina
n Crinlpellis
Crepidolus
Conocybe
Clitocybe
Figure 3. Number of publications of Rolf Singer by Figure Genera in which Singer and associates de-
4.
age. Singer's first scientific publications appeared in scribed at least 40 new taxa. Singer and associates de-
1922, when he was 16 years old. This publishing effort scribed 2,452 specific and infraspecific taxa. These were
peaked in his late 40s and early 50s, when he published classified into 246 genera. He described 40 or more taxa
103 works in a span of 10 years. In total, Singer pub- in 1 of these genera, which account for 50% of his new
1
lished 439 scientific papers or books. taxa.
Singer's Contributions to Mycology

i
"400
200
z
Figure 5. Geographic distribution of Singer's type 100
specimens. Type localities for Singer's new taxa occur

in 58 countries or areas. Over 85% of these taxa were
described from North, Central, and South America, with BAFC INPA MICH SCO
Herbaria Housing Singer's Type
Argentina, Bolivia, Brazil, Chile, Colombia, Mexico,
and the United States accounting for 75%. Descriptions Herbaria with large holdings of Singer's
Figure 6.
of 1.1% of the taxa do not include data on the country
type specimens. Singer's holotypes are preserved in 40
of origin. herbaria throughout the world. He consistently deposited
type material in the institution where he was working.
In some cases, parts of the holotype or paratypic mate-
tive role in the identification of the hallucinogenic rial for South American material are also housed in F,
FH, K, or MICH. Although Singer's holotypes are scat-
fungi used in Mexico and insubsequent physio- tered throughout the world, most are concentrated in the
logical, ecological, and cultural studies of these herbaria shown in this figure.
fungi (e.g., Singer, 1957f, 1958h, r). Other papers
on hallucinogenic fungi include Singer, 1959h,
1960J, and 1978g. He also was active in the study
of the cultivation and utilization of edible mush- Literature Cited
rooms (e.g., Singer, 1961g, 1964i, 1987a).
A summary of his publications does not fully Holmgren, P. K., N. H. Holmgren, and L. C. Barnett.
document the impact that Rolf Singer had on my- 1990. Index Herbariorum. Part I: The Herbaria of the
World, 8th ed. New York Botanical Garden, 693 pp.
cology. He was an active correspondent, and he
responded promptly to the numerous requests for Mueller, G. M. 1994. Rolf Singer's mycological legacy.
assistance and information. Numerous mycolo- Mcllvainea, 11(2): 7-9.
gists had the pleasurable and rewarding experi- M.
Mueller, G. 1995. Rolf Singer, 1906-1994. Mycol-
ence of working with him in the field, and he ogia, 87: 144-147.
facilitated the research of many mycologists, bot-
Singer, M. 1979. Glancing back. Beiheft Sydowia, 8:
anists, and zoologists working in the Neotropics. 14-25.
His presence in mycology will live on through
Singer, M. 1984. Mycologists and Other Taxa. J. Cramer,
the incredible legacy that he left for us. We
hope Braunschweig, Germany, 120 pp.
that the contributions included in this volume will
Singer, M. 1994. In memoriam Rolf Singer. Mcllvainea,
help current and future mycologists tap this
11(2): 4-6.
wealth of information.
Chapter One
TABLE 1 . List of new genera published by Rolf Singer.
Genus
TABLE 1. Continued.
Genus
TABLE 3. Herbaria with holdings of Singer's type specimens. 51
BAFC
The Field Itinerary of Rolf Singer
Betty A. Strack
Gregory M. Mueller
Singer maintained an incredibly active zil, Colombia, Peru, and Uruguay, as well as in
Rolffieldwork program, with mycological col- Mexico, the United States, and several countries
lecting trips undertaken to many areas of North, in Europe, including Austria, the Netherlands, and
Central, and South America, Europe, and Japan Sweden. After moving to Chicago in 1968, Singer
throughout his career. Because of the complexity continued an intensive field schedule in South
of his program and because he deposited
field America (including Amazonian Europe
Brazil),
specimens herbaria where he was working,
at the (especially Czechoslovakia, Austria, and Switzer-
we felt his field itinerary would be useful to doc- land), Mexico, and the United States, and he start-
ument when and where he collected. Table 4 lists ed fieldwork in Costa Rica and Japan. Other plac-
the countries where Singer undertook fieldwork, es where Singer collected in more recent years
and his field itinerary is given in Table 5. included Puerto Rico, Honduras, and Morocco.
Until 1941 Rolf Singer collected fungi in var- Table 5 lists most of Singer's fieldwork itiner-
ious parts of Europe and Asian Soviet Union. He ary from 1941 to 1992. Data in Table 5 were ob-
began gathering mushrooms as a boy in the for- tained from his field notebooks and from label
ests around his parents' home in the small village data of Singer collections housed at F (herbarium
of Schliersee in southern Germany. From 1928 to abbreviations follow Holmgren et al., 1990). In-
1931 he primarily collected in Austria while he formation before 1941 was either lacking or too
was studying for his Ph.D. at the University of difficult to accurately interpret. In addition to pro-
Vienna. He also undertook the first
mycological viding a historical record, Table 5 lists the time
expeditions to the western part of the Caucasus when he was by year, to help readers
in the field
Mountains in 1928 and 1929. In 1934 he moved locate the herbaria in which he deposited speci-
to Spain, then France, collecting fungi in both mens from specific localities, and by month of
countries. Singer then lived in the Soviet Union each year, to provide information on seasonality
(1935-1940), where he undertook mycological of fruiting. Since Singer routinely deposited spec-
trips around Leningrad (St. Petersburg) and ex- imens in the institution of the country in which
peditions to the Caucasus and Altai Mountains, he was working, information on the countries
Siberia, and Karelia. where he collected may help researchers locate
Singer moved to the United States in 1941 and herbaria most likely to have Singer specimens in
worked at the Farlow Herbarium at Harvard Uni- its holdings. In other cases, he deposited his col-
versity until 1948. In addition to collecting in the lections at his home institution. For example,
northeastern United States, Singer became ac- specimens from Singer's 1977 and 1978 Amazon
quainted with subtropical fungi while carrying out expeditions in Brazil are located in Manaus, Bra-
fieldwork in southern Florida. From 1948 to 1968 INPA, whereas many of Singer's specimens
zil, at
Singer held positions in Tucuman and Buenos Ai- collected in 1974 in Czechoslovakia are housed
res,Argentina, and Santiago, Chile. In addition to in Chicago at F, his home institution. Holotypes
collecting throughout Argentina and Chile, he un- of taxa described by Singer are preserved in 40
dertook fieldwork during this time in Bolivia, Bra- herbaria worldwide (see Table 3, page 7).
The Field Itinerary of Rolf Singer

Some countries (e.g., Argentina, Brazil, and the an assigned These letters correspond
letter prefix.
United States) are subdivided into smaller areas to the general areas where he collected, which in
in Table 5 because these large countries have di- turn are related to where the collections might
verse habitats and may have Singer collections in have been deposited. For the most part, the letters
more than one herbarium. Most of his collections representing specific areas, with the most likely
from subtropical Argentina and Brazil are depos- herbaria for housing those collections given in pa-
ited in LIL and BAFC, while specimens from rentheses, are: A= Asia (low numbers at W, high
temperate Argentina are primarily in BAFC. Most numbers B = tropical South America (LIL,
at F);
tropical Brazilian specimens are housed at INPA. BAFC, INPA, F); C = Europe (low numbers in
Many of Singer's collections were made on lo- European herbaria, high numbers mostly at F); F
calday excursions from his then current home.
= Florida, USA (mostly at FH); M
= Mexico and
The parts of the countries where Singer had ex- southern South America (LIL, BAFC, SGO, some
tended residences along with his home institutions at F); N = North America except Florida and
are presented here. These are southern Germany Mexico (MICH and F); S = Buenos Aires region,
(until 1928, and from 1931 to 1933); Vienna, Argentina (LIL, BAFC); and T
= subtropical
Austria (W, 1928-1931
and 1933); Barcelona, South America, Tucuman region, Argentina (LIL,
Spain (BCB, 1934-1935), and Paris, France (PC, BAFC, few at F). He used four additional small
1935); Leningrad, USSR (LE, 1935-1940); Cam- series of numbers: MA
= Mexico (F), H = Ha-
bridge, Massachusetts, USA (FH, 1941-1948); waii, USA (F), V = Virginia, USA (FH), and W
Tucuman, Argentina (LIL, 1948-1961); Buenos = White Mountains, New Hampshire, USA (FH).
Aires, Argentina (BAFC, 1961-1967); Santiago, Herbarium abbreviations are from Index Herba-
Chile (SGO, 1967); Manaus, Brazil (INPA, 1977- riorum (Holmgren et al., 1990).
1979); and Chicago, USA (F, 1968-1994). Rec- Even though this itinerary remains incomplete,
ords of these local collecting trips, especially the over 50 years of Rolf Singer's diverse field pro-
earlier ones (e.g., Massachusetts area, United gram is presented here. This compilation provides
States, from 1941 to 1948), may not be listed in information on the geographic breadth of his col-
Table 5. lections and taxonomic work.
Rolf Singer used what at first appears to be a
complicated numbering system for his collections.
However, if one understands the system, it is usu- Literature Cited
ally possible to predict where his collections
Holmgren, P. K., N. H. Holmgren, and L. C. Barnett.
might be housed. He organized his collections and 1990. Index Herbariorum. Part I: The Herbaria of the
corresponding field notebooks by numbers with World, 8th ed. New York Botanical Garden, 693 pp.
10 Chapter Two
TABLE 4. Rolf Singer's fieldwork, by country and TABLE 5. Itinerary of Rolf Singer's fieldwork since
year. 1941.*
Country Year(s)
Argentina 1948-1952, 1955-1966, 1969,

1988-1990
Austria 1928-1931, 1960, 1964, 1970,
1976, 1979, 1985, 1987
Bolivia 1956
Brazil 1951, 1952, 1956, 1957, 1960,
1961, 1964, 1966, 1977-1981,
1983, 1985
Canada 1984
Chile 1963, 1965, 1967
Colombia 1960, 1968
Costa Rica 1981, 1983, 1984, 1986, 1989
Czechoslovakia 1970, 1974, 1979
Denmark 1964
Ecuador 1973
France 1935?, 1970, 1971
Germany Until 1928, 1931-1933, 1970, 1987
Great Britain 1960,1970,1971
Honduras 1991
Iceland 1985
Italy 1970
Japan 1983, 1984
Mexico 1957, 1969, 1982, 1988
Morocco 1992
The Netherlands 1960
Norway 1964
Peru 1958
Portugal 1961
Puerto Rico 1987
Spain 1934, 1961, 1979
Sweden 1964
Switzerland 1964, 1970, 1971, 1976, 1980, 1984
Uruguay 1952, 1965, 1966
USA 1941-1947, 1953, 1958, 1968-1976,
1979-1992
USSR 1928, 1929, 1935-1940
Yugoslavia 1970
TABLE 5. Continued TABLE 5. Continued
Year
An Index to Genera, Species, and Infraspecific Taxa
of Basidiomycete Fungi Described by Rolf Singer
Qiuxin Wu
Gregory M, Mueller
Clark L. Ovrebo
the following index, genus, species, variety, iting type specimens in the institution where he
In and form names published by Rolf Singer are was working, even though it was not clearly stat-
listed alphabetically with literature citations and ed in his publication. Herbarium abbreviations are
type specimen information. This index is intended from Index Herbariorum (Holmgren et al., 1990,
to treat specimen-based taxa that Singer described New York Botanical Garden, Bronx. 693 pp.).
in the literature. New genus names are included.
New combinations, nomina nova, and supraspe- ACURTIS
cific names are not listed here. Literature citations
sericem ssp. antarcticus Singer, 1959n: 587
(i.e., year and a suffix: page number e.g., 1969a:
HOLOTYPE: Australia. Macquarie Island: Half
319) correspond to the listings in Chapter 4,
Moon Bay, 31 January 1951, Bunt (LIL)
where full citations are given. Holotype informa-
tion, including type locality, date, collector, col- AERUGINOSPORA
lection number, and herbarium of deposit, is from
hiemalis Singer & Clemen9on, 1971b: 118
Singer's original publications. In the few cases of
HOLOTYPE: Switzerland. Vaud: Botanical Gar-
discrepancies found between Singer's publica-
den Lausanne, 15 February 1971, Singer C
tions, we
consulted his unpublished field note-
5251 (F)
books and appended a short note or discussion to
the listing. Many of Singer's taxa were described
AGARICUS
before enforcement of certain rules by the Inter-
national Code of Botanical Nomenclature 1
nebularum Singer, 1969a: 165
(i.e.,
HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16
January 1935 for Latin diagnosis and 1 January
1958 for typification). Taxa described after 1935 April 1967, Singer M 6511 (SCO)
valdiviae Singer, 1965i: 161
without a Latin diagnosis are annotated with "no
HOLOTYPE: Chile. Valdivia: Hueycolla, 30
Latin diagnosis." Some taxa described before March 1963, Singer M 3237 (BAFC)
1958 have data on the geographic origin of the
type specimens but do not have a designated ho- AGROCYBE
lotype. These data are presented following "Type
information" to separate them from those having aegerita var. rugosovenosa Singer, 1953b: 19
HOLOTYPE: Argentina. Tucumdn: Rio de los
complete holotype information, which follows Sosas, 26 February 1952, Singer T 1875 (LIL)
"HOLOTYPE." A question mark (?) after an her- 225
allocystis Singer, 1969a:
barium name indicates that the herbarium is the HOLOTYPE: Argentina. Buenos Aires: Sierra
possible place of deposit, based on Singer's un- de la Ventana, 15 November 1962, Singer S
published field notes and Singer's habit of depos- 348 (BAFC)
Index to Genera, Species, and Infraspecific Taxa 13

AGROCYBE
coprophila Singer, 1941e: 99 diemii Singer, 1954b: 120, 121

Type information: USSR. Altai Mtns., Singer & HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
Vasilieva (LE?) pi National Park, Quetrihue, 13 May 1952,
firma var. tucumana Singer, 1959b: 398 Singer M 562 (LIL)

HOLOTYPE: Argentina. Tucuman: Rio de los foetens Singer, 1953b: 15
Sosas, 4 February 1955, Singer T 21 30 (LIL). HOLOTYPE: Argentina. Tucuman: Pie del Per-
The type information for this taxon is identical iquillo, 30 December 1951, Singer & Helber-
to that of Agrocybe tucumana Singer (1973d: ger T 1672 (LIL)
76), except for the herbarium of deposit, humboldtii Singer, 1963b: 241
which is BAFC. HOLOTYPE: Colombia. Cundinamarca: above
insignis Singer, 1941b: 85 Pacho, Los Robedales, 26 July 1960, Singer
Type information: USSR. Terskiense, Lobik B 3527 (BAFC)
(LE?) lilloi Singer in Singer & Digilio, 1951e: 245
karelica Singer, 19415: 85 HOLOTYPE: Argentina. Tucuman: Sierra de
Type information: Finland. Karelia: near Kivacz, San Ciudad Universitaria, 6 November
Javier,
May 1935, Freindling (LE?) 1950, Grassi [Singer no.] T 1016 (LIL) [Pub-
lazoi Singer, 1969a: 222 lished in 1952]
HOLOTYPE: Chile. Valparaiso: Vina del March, nana Singer, 1941b: 85
28 July 1966, Lazo VI- 17 (SCO)
Type information: USSR. Kazakhstan, April 1915,
microspora Singer, 1977g: 205 Fedtschenko (LE?)
HOLOTYPE: USA. Illinois: Morton Arboretum,
16 May 1970, Singer N 2100 (F) AMPAROINA Singer, 1958a: 110
paradoxa Singer, 1973d: 76
HOLOTYPE: Argentina. Tucuman: near Tafi del Type species: Marasmius spinosissimus Singer
Valle, 29 January 1962, Singer T 3767 (BAFC)
heteracantha Singer, 1976J: 58
procera Singer, 1969a: 224 HOLOTYPE: Colombia. Valle: Cali, Buenaven-
HOLOTYPE: Chile. Valparaiso: Vina del Mar,
tura, San Joaquin, 19 April 1968, Singer B
March 1966, Lazo VI- 1 (SGO) 6250 (F)
tucumana Singer, 1973d: 76
HOLOTYPE: Argentina. Tucuman: Rio de los AMYLOFLAGELLULA Singer, 1966a: 14
Sosas, 4 February 1955, Singer T 2130
(BAFC). The type information for this species Type species: Cyphella pulchra Berk. & Br.
is identical to that of Agrocybe firma var. tuc-
umana (1959b: 398), except for the herbarium ANTHRACOPHYLLUM

of deposit.
viscosa Singer, 1969a: 223 paxilloides Singer, 1955d: 227
HOLOTYPE: Chile. Santiago: Maipii, 8 Septem-
HOLOTYPE: Brazil. Pernambuco: Recife, Dois
ber 1966, Lazo RI-8 (SGO) Irmaos, 10 April 1955, Jose de Silva & Batis-
ta 2225 (LIL)
xerophytica Singer, 1959b: 399
HOLOTYPE: Argentina. Tucuman: above San
Pedro de Colalao, 21 January 1955, Singer T APHYLLOTUS Singer, 1973d: 29
2053 (LIL)
Type species: A. campanelliformis Singer
ALNICOLA campanelliformis Singer, 1973d: 29
devia Singer, 1950a: 150 HOLOTYPE: Colombia. Cundina marca: Futa-
HOLOTYPE: Argentina. Buenos Aires: Punta gasuga, 1 April 1968, Singer B 6034 (F)
1
Lara, 9 June 1949, Singer Sill (LIL)

diplocystis Singer, 1950a: 151 ARCANGELIELLA
HOLOTYPE: Argentina. Tucuman: above Tafi
crassa Singer & Smith, 1960h: 74
del Valle, 1
May 1949, Singer T 503 (LIL)
HOLOTYPE: USA. California: Stanislaus Na-
lignicola Singer, 1939a: 67
tional Forest, 25 July 1931, Patty s.n. (NY)
Type information: USSR. Kazakhstan: Alma
Ata, Nevodovski (LE?)
ARMILLARIELLA
AMANITA 12
affinis Singer, 1989a:
aestivalis Singer ex Singer, 1959b: 390 HOLOTYPE: Costa Rica. Heredia: Varablanca,
HOLOTYPE: USA. Michigan: Cheboygan Coun- near Hacienda "Tirol", 11 June 1984, Singer
ty, Burt Lake, 29 July 1953, Singer N
899 (F) B 14615 (F)
14 Chapter Three
ARMILLARIELLA
clavuligera Singer in Singer & Digilio, 195 le: ASTEROTUS Singer, 1943c: 161
70
HOLOTYPE: Argentina. Tucuman: Santa Rosa, Type species: Panus dealbatus Berk.
17 December 1950, Singer T 1060 (LIL)
argentinus Singer in Singer &
Digilio, 195 le:
[Published in 1952]
107
densifolia Singer in Singer & Digilio, 195 le:
HOLOTYPE: Argentina. Tucuman: Rio de los
72
Sosas, 27 January 1951, Singer T 1136 (LIL).
HOLOTYPE: Argentina. Tucuman: Quebrada de
[Published in 1952].
Lules, 2 April 1949, Singer T 367 (LIL). [Pub-
lished in 1952]
graminum Singer, 1969a: 57
HOLOTYPE: Argentina. Neuquen: Cerro Colo-
ditopa Singer, 1950c: 191 rado, 9-10 March 1959, Singer M 1733
HOLOTYPE: Argentina. Tucuman: Anta Muer-
(BAFC)
ta, 10 April 1949, Singer & Digilio [Singer
no.] T 403 (LIL). The type information is
identical to that of Nothoclavulina ditopa AUSTROBOLETUS

Singer (1970a: 18).
graciliqffinis Singer, 1989a: 126
griseomellea Singer, 1969a: 40
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, HOLOTYPE: Brazil. Amazonas: Road Manaus-
1-4 May 1967, Singer M 6574 (SCO)

Caracarai, km 45, 31 March 1980, Webber
[Singer no.] B 12088 (INPA)
minuta Singer & Digilio, 1954b: 106, 107
neotropicalis Singer, Garcia & Gomez, 199 la:
Type information: Argentina. Neuquen: Nahuel
53, 88
Huapi National Park, Quetrihue, May 1952,
M
576/ M667? (LIL) HOLOTYPE: Costa Rica. Guanacaste: Bagaces,
Singer
7 km NW of town, October 1982, Gomez
montagnei Singer, 1956f: 182
18708 (F)
HOLOTYPE: Argentina. Neuquen: Quetrihue,
13 May 1952, Singer M 513 (LIL)
olivaceus Singer, 1983f: 132
var. umbrinobrunnea Singer, 1956f:

HOLOTYPE: Brazil. Amazonas: Road Manaus-
montagnei
Caracarai, km 125, 20 July 1978, Singer &
183
Araujo [Singer no.] B 11323 (INPA)
puiggarii ssp. tigrensis Singer, 1970a: 8 AUSTROGASTER Singer, 1962a: 57

HOLOTYPE: Argentina. Buenos Aires: Punta
Lara, 1 June 1949, Singer S 58 (LIL) Type species: Austrogaster marthae Singer
saviczii Singer, 1938e: 6
baeospermum Singer, 1969a: 360
Type information: USSR. Savicz (LE?) HOLOTYPE: Argentina. Rio Negro: Cerro Otto,
sparrei Singer, 1956f: 183
February 1934, Burkart 4400 (BAFC)
HOLOTYPE: Chile. Juan Fernandez Island:
marthae Singer, 1962a: 57
Masa Tierra, Quebrado Piedra Agujereada, 12
HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
March 1955, Sparre F 33 (R, LIL)
pf National Park, February, Singer M 3040
sparrei var. elaeodes Singer, 1969a: 45 (LIL)
HOLOTYPE: Argentina. Neuquen: Puerto Man-
zano, 18 May 1952, Singer 741 (LIL) M
sparrei var. xantha Singer, 1969a: 45
BAEOSPORA Singer, 1938a: 191, 193
HOLOTYPE: USA. Michigan: Hell, 23 Septem-
species: Colly bia sensu Quel
ber 1957, Singer M 1620 (MICH) Type myosura Fr.
& Ricken
viridiflava Singer, 1989a: 12
HOLOTYPE: Spain. Catalonia: Mountains near
mundula Singer, 1989a: 90
Tossa, 24 October 1979, Singer C 9601 (F)
HOLOTYPE: Amazonas: Igarape do Tar-
Brazil.
yungensis Singer, 1970a: 12 umazinho, 14 May 1980, Singer B 12168
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car-
(INPA)
men Pampa, 19 February 1956, Singer B 1268
oligophylla Singer, 1938a: 194
(LIL)
Type information: USSR. Altai Mtns.: Czuiskie
Alpy, August 1937, Singer & Vasilieva (LE?)
ARTHROSPORELLA Singer, 1970a: 17 pallida Singer, 1978h: 40
HOLOTYPE: Ecuador. Napo: Shushufindi, 15
Type species: Armillariella ditopa Singer May 1973, Singer B 7451 (F)

BOLBITIUS
BOLBITIUS chilensis Singer, 1967b: 69

HOLOTYPE: Chile. Valdivia, 27 April 1967, R.
exiguus Singer, 19361: 433 & M. Singer & Hernandez [Singer no.] 6933
Type information: Spain. Catalonia: Pyrenees (SAM)
Mtns., July 1934.
frostii ssp. floridanus Singer, 1945i: 799
mesosporus Singer, 1977g: 219 Type information: USA. Florida: Sebring.
HOLOTYPE: Ecuador. Tungurahua: Banos, 27
fuligineotomentosus Singer, 1973d: 101
April 1973, Singer B 7148 (F) HOLOTYPE: Colombia. Valle: Rio Pance, 3
tucumanensis Singer, 1959b: 398 May 1968, Singer B 6958 (F)
granulosiceps Singer, 1945i: 798
la Higuera, 25 January 1955, Singer T 2081 Type information: USA. Florida: Miami.
(LIL). Also published in Singer, 1961b: 69.
griseus ssp. pinicaribaeae Singer, 1945i: 797
Type information: USA. Florida: Coral Gables.
BOLETELLUS guadelupae Singer & Fiard, 1976i: 445
ananas var. crassotunicatus Singer, 1983f: 152 HOLOTYPE: Guadeloupe. Matouba, 31 July
HOLOTYPE: 1975, Fiard s. n. (F)
Nicaragua. Slilma Sia, May 1976,
heterodermus Singer, 1989a: 124
Ivory S/23 (F)
ananas var. minor Singer, 1983f: 153 HOLOTYPE: Costa Rica. Puntarenas: Rio Baru,
HOLOTYPE: Brazil. Amazonas: Road Manaus- August 1983, Gomez 21548 (F)
Caracaraf, km 115, 4 April 1978, Singer B hypocarycinus Singer, 1945i: 798

10913 (INPA) Type information: USA. Florida: Alachua Coun-
ty, Gainesville.
coccineus var. amarus Singer, 1992a: 8
michoacanus Singer, 1977g: 256
HOLOTYPE: Costa Rica. Guanacaste: Bagaces,
7 km NW
HOLOTYPE: Mexico. Michoacan: Parque Na-
of town, Gomez & Alfaro 22101
(F)
cional near Mirador, 9 August 1969, Singer M
8998/11 (F)
fibuliger Singer, 1983f: 154
patouillardii Singer, 1947a: 55
HOLOTYPE: Venezuela. Dto Federal: Parque
HOLOTYPE: Cambodia, Reserve Forestiere de
Nacional El Avila, 27 July 1972, Dumont,
Cain &
Samuels [Dumont no.] VE-6201 (NY)
Campong Chhuong, July, Petelot 222 (FH)
rubellus ssp. bicoloroides Singer, 1945i: 798
ivoryi Singer, 1983f: 163
Type information: USA. Florida: Alachua Coun-
HOLOTYPE: Belize. Augustine: Forest Station,
ty-
18 June 1976, Ivory 5/725 (F)
rubellus ssp. caribaeus f. serotinus Singer,
linden Singer, 1945a: 134
1947a: 53
Type information: Liberia. Miamu, Under 408 Type information: USA. Florida: Dade County.
(FH) rubellus ssp. caribaeus Singer, 1945i: 798
pictiformis var. fallax Singer, 1945a: 132 Type information: USA. Florida: Miami.
HOLOTYPE: USA. Florida: Alachua County, rubellus ssp. consobrinus Singer, 1945i: 798
Hammock, 14 July 1943, Singer F
Kelley's
Type information: USA. Florida: Miami.
2716 (FH)
rubellus ssp. dumetorum f. crassotunicatus
Singer, 1947a: 51
BOLETINUS HOLOTYPE: USA. Florida: Dade County, Ma-
asiaticus Singer, 1938d: 164 theson Hammock, 8 October 1942, Singer F
976 (FH)
Type information: USSR. (LE?)
rubellus ssp. dumetorum Singer, 1945i: 798
oxydabilis Singer, 1938d: 160
information: USSR. Altai Mtns.: Lake Te- Type information: USA. Florida: Miami.
Type
rubellus ssp. fraternus f. subdumetorum Singer,
lezkoje, September 1937, Singer (LE?)
1947a: 49
BOLETOCHAETE 358
HOLOTYPE: USA. Florida: Highlands County,
Singer, 1944a:
Highlands Hammock State Park, 8 September
Type species: Boletus spinifer Pat. & Baker 1942, Singer F 648a (FH)
rubricitrinus var. fairchildianus Singer, 1945i:
brunneosetosa Singer, 1944a: 359
Har-
798
Type information: Liberia. Nengbe, April,
ley (FH?) Type information: USA. Florida: Miami.
splendidus ssp. moseri Singer & Kuthan,
BOLETUS 1976h: 151
HOLOTYPE: Czechoslovakia. Slovakia: Rako-
austrinus Singer, 1945i: 798 va, near Cadca, 31 August 1975, J. Kuthan
Type information: USA. Florida: Miami. 75/S 141 (F)
16 Chapter Three
BOLETUS
subsolitarius Singer, 19451: 798 luteofuscum Singer, 1953d: 115

Type information: USA. Florida: Miami. HOLOTYPE: Brazil. Rio Grande do Sul: Sao
subvelutipes glabripes Singer, 1947a: 70
f. Leopoldo, 31 October 1951, Singer B 27
HOLOTYPE: USA. Florida: Alachua County, (LIL)
Gainesville, 30 May 1943, Singer F 2169 luteofuscum var. maior Singer, 1969a: 53
(FH) HOLOTYPE: Chile. Valparaiso: Vina del Mar,
weberi Singer, 1945i: 797 16 July 1966, LawVI- 3 (SGO)

psilocybe Murrill &
Singer, 1944a: 363
Gainesville.
ty, Type information: USA. Florida: Juniper Springs,
October, Murrill (FH?)
BONDARZEWIA Singer, 1940: 4 terrigenum Singer, 1950c: 192
HOLOTYPE: Argentina. Tucuman: between San
Type species: Polyporus montanus Quel. Pablo and Parque Aconquija, 2 April 1949,
Singer (LIL)
perniciosa Singer, 1953c: 884
May 1952, Singer 608 (LIL) M CALOCYBE
BRAUNIELLULA Smith & Singer, 1958c: 928 africana Singer, 1977g: 268
HOLOTYPE: Zaire. Panzi Kivu, Goossens-Fon-
Type species: Brauniellula nancyae Smith tana 5334 (BR)
alneti Singer, 1977g: 267
leucosarx Smith & Singer, 1958c: 928
HOLOTYPE: Argentina. Jujuy: Lagunas de
HOLOTYPE: USA. Idaho, Payette National For-
Yala, 14 February 1966, Singer (F)
est, 31 July 1958, Smith 59316 (MICH?)
atropapillata Singer, 1977g: 269
HOLOTYPE: Ecuador. Napo: Lago Agrio, 8
CALATHINUS May 1973, Singer B 7284 (F)
substriatulus Singer, 1943a: 156 bipigmentata Singer, 1977g: 267

HOLOTYPE: Brazil. Pernambuco: Dois Irmaos,
Type information: USSR. Altai Mtns. (Kurai),
13 July 1960, Singer (F)
August.
caucasica Singer, 1961b: 47
Type information: USSR. Caucasus: Krasnodar
CALLISTODERMATIUM Singer, 198 la: 506
Yatyrgrarta, 29 July 1935, Vassilieva (LE,
KAZ)
Type species: Callistodermatium violascens Singer
coniceps Singer, 1977g: 268
violascens Singer, 198 la: 506 HOLOTYPE: Venezuela. Dto Federal: Parque
HOLOTYPE: Amazonas: 30 km N of
Brazil. Nacional El Avila, 27 July 1972, Dumont and
Manaus, 3 June 1977, Singer B 9926 (F) VE 6205 (NY)
others
cyanea Singer ex Redhead & Singer, 1978c:

CALLISTOSPORIUM Singer, 1944a: 363 501
HOLOTYPE: Brazil. Sta Catharina: Porto Novo,
Type species: Gymnopilus palmarum Murr. 1928, Rick s. n. (FH). Basionym: Calocybe
cyanea Singer, 1946g: 126 (nom. invalid).
amazonicum Singer, 1977g: 263
HOLOTYPE: Brazil. Rio Grande do Sul: Gua- cyanella Singer ex Redhead &
Singer, 1978c:
March 1956, 501
pore, near Guajara mirim, 10
(BAFC)
Singer
HOLOTYPE: USA. Florida: Dade County, near
Miami, Matheson Hammock, Singer F 734
at
chrysophorum Singer, 1961b: 51
HOLOTYPE: USA. Florida: Alachua County, 8 (FH). Basionym: Calocybe cyanella Singer,
June 1943, Singer F 2227 (F) 1946g: 124 (nom. invalid).
fibrillosa Singer, 1950c: 192
favrei Singer, 1977g: 262
HOLOTYPE: Switzerland. Vaud Bois de Chene: HOLOTYPE: Argentina. Tucuman: Taff del Val-
Chene pr, Nyon, 18 October 1942, Favreano le, Parque Aconquija, 8 May 1949, Singer T
(Favreano) 546 (LIL)
galerinoides Singer, 1948c: 29 heterospora Singer, 1961b: 46
HOLOTYPE: USA. Florida: Alachua County, HOLOTYPE: USA. Florida, Dade County, Ma-
Sugarfoot Hammock, 5 July 1943, Singer F theson Hammock, 9 November 1942, Singer
2598 (FH) F 1452c (FH)

CALOCYBE
rubra Rick in Singer ex Redhead Singer, & alnetorum Singer, 1975c: 866
1978c: 501 HOLOTYPE: Argentina. Jujuy: Lagunas de
HOLOTYPE: Brazil. 1928, Rick 585 (FH). Bas- Yala, 14 February 1966, Singer T 5140a (F)
ionym: Calocybe rubra Rick in Singer, 1948. austrochilensis Singer, 1969a: 86

Pap. Mich.Acad. Sci. Arts & Lett. 32: 126 HOLOTYPE: Chile. Llanquihue: Puella, 20
(nom. invalid) March 1959, Singer M 1948 (LIL)
Candida var. stipitata Singer, 1955a: 765
CALYPTELLA HOLOTYPE: Argentina. Catamarca, Santa

Rosa, Vallis Rio Campo, 18 January 1954
bonairensis Singer, 1973d: 27 (LIL)
HOLOTYPE: Argentina. Buenos Aires: La Pla- castaneipes Singer, 1975c: 856
ta, 20 June 1962, Singer S 206 (BAFC) HOLOTYPE: Colombia. Boyaca: between Ra-
epibrya Singer, 1973d: 25 miriqui and Rondon, 30 July 1960, Singer B
HOLOTYPE: Argentina. Tucuman: Sierra de 3632 (F)
San Javier, 14 May 1961, Singer T 3743 (LIL) dendrophora Singer, 1955a: 763
flava Singer, 1977g: 270 HOLOTYPE: Argentina. Neuquen: Puerto Man-
HOLOTYPE: Argentina. Jujuy: Lagunas de zano, 18 May 1952, Singer M 746 (LIL)
Yala, 9 February 1965, Singer (F) diplocystis Singer, 1975c: 868
pteridophytorum Singer, 1973d: 26 HOLOTYPE: Peru. Cuzco: Urubamba, Machu
HOLOTYPE: Colombia. Valle: Dagua, 27 April Picchu, 20 May 1958, Singer B 3025 (LIL)
1968, Singer B 64 13 (F) elongatispora Singer, 1975c: 851
CAMAROPHYLLUS Yala, 11 February 1965, Singer T 4024 (F)
floridana Singer, 1950e: 249

adonis Singer, 1952c: 172 HOLOTYPE: USA. Florida: Newnan's Lake, 31
HOLOTYPE: Argentina. Tierra del Fuego: Rio July 1947, Singer F 3428 (FH)
Grande, Estancia Nueva Argentina, Lago Fag-
gigantospora Singer, 1975c: 874
nano, 18 February 1950, Singer M
351 (LIL)
HOLOTYPE: Australia. New South Wales: be-
grinlingii Singer, 1973d: 2 tween Major's Crk. & Araluen, March 1970,
HOLOTYPE: Zaire. Brazzaville: Rte. de Linzo-
McVean s. n. (F)
lo, no date, Grinling 71106 (RPC). Isotype: F.
peullensis Singer, 1969a: 85
hieronymi Singer, 1977g: 277 HOLOTYPE: Chile. Llanquihue: Puella, 20
HOLOTYPE: Bolivia. Nor-Yungas: La Paz, San
March 1959, Singer 1894 (LIL) M
Geronino, 7 February 1956, Singer B 968 (F)
podocarpi Singer, 1975c: 864
laccarioides Singer, 1952c: 170 HOLOTYPE: Argentina. Jujuy: 15 km NE of
HOLOTYPE: Argentina. Tierra del Fuego: Rio Posta de Lozano, 20 February 1966, Singer T
Grande, Estancia Nueva Argentina, 16 Feb- 5307 (F)
ruary 1950, Singer M 300 (LIL) tenuitunicata Singer, 1955a: 764
paraiboensis Singer, 1977g: 278 HOLOTYPE: Argentina. Catamarca: Santa
HOLOTYPE: Brazil. Parafba: near Joao Pessoa,
Rosa, Suncho, Vallis Rio Campo, 18 January
1 1
July 1960, Singer B 3334 (BAFC) T 1750
1952, Singer (LIL)
patagonicus Singer, 1969a: 9 witteana Singer, 1975c: 876
HOLOTYPE: Chile. Llanquihue: Puerto Mont y
HOLOTYPE: Zaire. Pare National Albert, 17
San Augustin, 8 May 1967, Singer M 6790
June 1953, Fredericq 9440 (BR)
(SCO)
CAMPANELLA CANTHARELLULA Singer, 1936e: 281
aberrant Singer, 1975c: 855 Type species: Cantharellus umbonatus Gmelin

HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co-
roico, 12 February 1956, Singer B 1116 (LIL) infundibuliformis Singer, 1969a: 64
HOLOTYPE: Argentina. Rio Negro: Laguna
aequatorialis Singer, 1975b: 240
HOLOTYPE: Ecuador. Tungurahua: Rio Verde,
Frias, 3 May 1964, Singer 3861 (BAFC) M
26 April 1973, Singer B 7128 (F). Also pub-
lished in Singer, 1975c: 869. CHAETOCALATHUS Singer, 1942a: 518.
aeruginea Singer, 1975c: 854 [Published in 1943].
HOLOTYPE: Argentina. Tucuman: S. M. Tu-
cuman, 22 January 1955, Singer T 2078 (LIL) Type species: Agaricus craterellus Dur. & Lev.
18 Chapter Three
CHAETOCALATHUS
aurantiacus Singer, 1976a: 57 CLAVARIA

HOLOTYPE: Bolivia. Pando: Manuripi, Con-
quista, 25 March 1956, Singer B 2201 (F) albidobisa Singer, 1969a: 390
carnelioruber Singer, 1942a: 526 HOLOTYPE: Chile. Concepcion: Boca del Bio-
Type information: Guiana. Bartica, 9 December Bio, 11 November 1967, Singer 7814 M
1923, Under (FH). [Published in 1943]. (SCO)
eurysporus Singer, 1978f: 195 alliodora Bond &
Singer, 1936J: 724
HOLOTYPE: Panama. Panama: near "Altos de No type information given.
Pacora", 30 June 1975, Dumont, Carperter & floridana Singer, 1945d: 425
Mori [Dumont no.] P 1540 (NY). Also see HOLOTYPE: USA. Florida: Dade County, 17
September 1942, Singer F 733 (FH)

1989a: 67.
pachytrichus Singer, 1942a: 522 leucoporphyria Singer, 1947c: 205

HOLOTYPE: Phillippine Islands. Laguna: Mt. Type information: Spain. Catalonia: Riu de Sant
Makiling, July 1916, Baker 4356 (FH). [Pub- Maurici, July 1934, Singer,
lished in 1943].
wettsteinii Singer, 1931: 515
Type information: USSR. Caucasus.
CHAETOTYPHULA
xanthoporphyria Singer, 1947c: 205
amylochaete Singer, 1978h: 1 Type information: Spain. Catalonia: Riu de Sant
HOLOTYPE: Ecuador. Zamora, 21 km from Za- Maurici, July 1934, Singer.
mora on the Zamora-Zanxaza Rd., 31 July
1975, Dumont EC 1784 (NY)
CLAVULINA
columbiana Singer, 1978h: 2
HOLOTYPE: Colombia. Antioquia: Medellin,
angustisperma Singer, 1969a: 387
20 July 1974, Dumont and others CO 1744
HOLOTYPE: Argentina. Rio Negro: Nahuel
(NY or F?)
Huapi National Park near Santa Maria, 19
montana Singer, 1978h: 3
April 1965, Singer M
5061 (BAFC)
Yala, 9 February 1965, Singer T 3920 (BAFC)
CLITOCYBE
CHAMAEMYCES
aberrantissima Singer, 1953d: 118
paraensis Singer, 1989a: 100 HOLOTYPE: Brazil. Rio Grande do Sul: Pared,
HOLOTYPE: Brazil. Para: Belem, 7 June 1966, 2 November 1951, Singer B 52 (LIL)
Singer B 4243 (BAFC) acicola Singer ex Singer, 1990b: 6
HOLOTYPE: USSR. Kurai: Altai Mtns., 15 Au-
CHAMONIXIA gust 1938, Singer A 103 (LE)
brevicolumna Smith & Singer, 1959f: 219 aconquijensis Singer in Singer & Digilio,
HOLOTYPE: USA. Idaho, Payette National For- 1951e: 38
est, 26 August 1958, Smith 60519 (MICH) HOLOTYPE: Argentina. Tucuman: Tafi del Val-
le, Parque Aconquija, 30 March 1951, Singer
CHEIMONOPHYLLUM Singer, 1955f: 417 T 1461 (LIL). [Published in 1952].
& aprilis Singer in Singer & Digilio, 1951e: 27

Type species: Agaricus candidissimus Berk.
Curt.
HOLOTYPE: Argentina. Tucuman: Tafi del Val-
le, Parque Aconquija, 14 April 1951, Singer
T 1478 (LIL). [Published in 1952].
CHLOROPHYLLUM
asema Singer, 1973d: 11
molybdites var. luteolosperma Singer, 1946g: HOLOTYPE: Argentina. Tucuman: near Aguad-
139 ita, 25 February 1951, Singer T 1328 (LIL)
HOLOTYPE: USA. Florida: Alachua County, augustinensis Singer, 1955a: 766
Gainesville, 16 May 1943, Singer F 1953a HOLOTYPE: Trinidad. St. Augustine, 9 Febru-
(FH). [Published in 1948].
ary 1950, Wilson [Dennis no.] 481 (K)
bresadoliano-affinis Singer, 1943a: 35
CHROOGOMPHUS Type information: USSR. Altai Mtns.
rutilus ssp. michoacanensis Singer & Kuthan, brevibasidiata Singer, 1943a: 45
1976e: 85 Type information: USSR. Altai Krai: Artubash,
HOLOTYPE: Mexico. Michoacan: San Jose de aimak Turoczak. Published as Clitocybe (Om-
Purua, 8 August 1969, Singer M 8986 (F) phalina) brevibasidiata.

CLITOCYBE
catalaunica Singer, 1943a: 30 obscuratipes Singer, 1969a: 32

Type information: Spain. Catalonia: Vail d'Aran, HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
Salardu. September. 1934. Also published in pi National Park, above Puerto Manzano, 19
Singer, 1947c: 242.

March 1963, Singer M
3088 (BAFC)
caucasica Singer, 1978f: 230 oligophylla Singer in Singer & Digilio, 195 le:
HOLOTYPE: USSR. Swanetia: Central Cauca- 40
sus, 20 August 1928, Singer (W) HOLOTYPE: Argentina. Tucuman: Cerro Alto
coluiana Singer, 1973d: 11 del Taficillo, 4 March 1951, Singer T 1381
HOLOTYPE: Colombia. Cauca: Gabriel Lopez, (LIL). [Published in 1952].
San Pedro, 5 May 1967, Singer B 6967 (F) pleurotus Singer, 1952c: 182
HOLOTYPE: Argentina. Tierra del Fuego: Rio
cystidiosa Singer, 1967c: 356
HOLOTYPE: Zaire. Brazzaville: 'Route de Li- Grande, Estancia Nueva Argentina, 10 Feb-
foula', November 1965, Grinling 51,110

ruary 1950, Singer M
175 (LIL)
(BAFC) polylepidicola Singer, 1989a: 9

duckeana Singer, 1989a: 8
Yala, 10 February 1965, Singer T 4025
HOLOTYPE: Brazil. Amazonas: 30 km N of
(BAFC)
Manaus, 12 July 1978, Singer (INPA). No col-
lection number praeandina Singer, 1955c: 147
given.
HOLOTYPE: Argentina. Tucuman: Sierra de
espinosae Singer, 1969a: 33 San T 2215 (LIL)
Javier, 1955, Singer
HOLOTYPE: Chile. Santiago: Quebrada Diego,
6 July 1967, Singer M 7109 (SGO) scandens Singer in Singer &
Digilio, 195 le: 36
HOLOTYPE: Tucuman: Sierra de
Argentina.
exilis Singer, 1989a: 9
San Javier, Anta Muerta, 24 April 1949, Sing-
er, Digilio & Grassi [Singer no.] T 490 (LIL).
Manaus, 6 April 1977, Singer B 9618 (INPA)
fimbriata Singer, 1978f: 231 spodoxutha Singer, 1953b: 3
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Or- HOLOTYPE: Argentina. Tucuman: near Tapia,
iente, 16 March 1956, Singer B 2008 (LIL, 6 January 1952, Singer T 1708 (LIL)
BAFC) steppicola Singer, 1943a: 33
hyacinthina Singer, 1943a: 46 Type information: USSR. Oirotia: aimak Kosch-
Type information: USSR. Oirotia: Aimak Czoia. agatsch, August.
Published as Clitocybe (Omphalina) hyacin-
subhygrophana Singer, 1943a: 27
thina.
Type information: USSR. Altai Mtns.
kabulensis Singer, 1955a: 766 subhygrophanoides Singer, 1952c: 181
HOLOTYPE: Afghanistan. Tangi gharu: near HOLOTYPE: Argentina. Tierra del Fuego: Rio
Kabul, 12 March 1950, Gilli LXXXVI (W) Grande, Estancia Nueva Argentina, 13 Feb-
laricicola Singer, 1943a: 28 ruary 1950, Singer M
245 (LIL)
Type information: USSR. Altai Mtns. subleptoloma Singer, 1952c: 179
lignicola Singer, 1973d: 12
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 12 Grande, Estancia Nueva Argentina, 10 Feb-
March 1956, Singer B 1900 (LIL) ruary 1950, Singer M
169 (LIL)
melliolens Singer in Singer &
Digilio, 195 le:
subsquamulosa Singer, 1943a: 35
34 Type information: USSR. Altai Mtns., July-Au-
gust.
HOLOTYPE: Argentina. Buenos Aires: Garibal-
di, 30 May 1949, Singer S 38 (LIL). [Pub-
subulifera Singer, 1955a: 767
HOLOTYPE: Argentina. Tucuman: De "Tapia"
lished in 1952].
vs. mer, 17 February 1951, Singer T 1220
monticola Singer, 1943a: 30
(LIL)
Type information: Spain. Catalonia: Pyrenees
tournefortiae Singer, 1989b: 794
Mtns., Aiguamoix, July, 1943. Also published
HOLOTYPE: Henderson Islands. (South Pacif-
in Singer, 1947c: 221.
ic), Northwest Beach, Schuster 00007 (F)
montivaga Singer, 1978f: 232
tungurahuae Singer, 1978f: 233
HOLOTYPE: Ecuador. Tungurahua: Banos, 1
HOLOTYPE: Ecuador. Tungurahua: Rio Topo,
May 1973, Singer B 7234 (F) 30 April 1973, Singer B 7215 (F)
nothofageti Singer, 1954b: 103 vasilievae Singer, 1941b: 84
HOLOTYPE: Argentina. Neuqu6n: Nahuel Hua- Type information: USSR. Oirotia: Altai Mtns.,
pf National Park, 15 May 1952, Singer 640 M August 1937, Singer & Vasilieva (LE). Also
(LIL) published in Singer, 1943a: 26.
20 Chapter Three
CLITOCYBE
vulgaris Singer, 1961b: 48 embrapaensis Singer, 1989a: 22

HOLOTYPE: USA. Massachusetts, Cambridge, HOLOTYPE: Brazil. Amazonas: Reserva Ducke,
Singer s.n. (FH) 6 October 1978, Singer B 1 1403 (INPA)
flavomycelina Singer in Singer &
Digilio, 1951e:
CLITOCYBULA 51
HOLOTYPE: Argentina. Tucumdn: Rio de los
azurea Singer, 1973d: 18
Sosa, 27 January 1951, Singer T 1128 (LIL).
HOLOTYPE: Brazil. Sao Paulo: Serra de Can-
tareira, 17 December 1964, Singer B 4115
(BAFC) fuegiana Singer, 1952c: 186

mellea Singer, 1954b: 111
Grande, Estancia Nueva Argentina, 10 Feb-
157 (LIL)
pf National Park, Quetrihue, 18 May 1952,
heimii Singer, 1937: 234
Singer M 751 (LIL)
Type information: USSR, near Chosta, Septem-
ber 1936, Vasilieva (LE?)
CLITOPILUS
inconspicua Singer, 1989a: 22
hobsonii var. chilensis Singer, 1969a: 333 HOLOTYPE: Brazil. Amazonas, 30 km N of
HOLOTYPE: Chile. Santiago: Cerro Robles, 19 Manaus, 31 October 1977, Singer B 10233
June 1967, Singer M
6978 (SCO) (INPA)
incrustatus Singer, 1948c: 38 ixotrama Singer, 1973d: 20
HOLOTYPE: USA. Florida: Alachua County, HOLOTYPE: Bolivia. Pando: Manuripi, 25
Gainesville, 19 May 1943, Singer F 1981 March 1956, Singer B 2202 (BAFC)
(FH) Hndquistii Singer in Singer & Digilio, 195 le:
incrustatus var. macrosporus Singer, 1969a: 57
334 HOLOTYPE: Argentina. Buenos Aires: Punta
HOLOTYPE: Chile. Valdivia: Hueycolla, 4 Lara, 9 June 1945, Singer S 116 (LIL). [Pub-
May
1967, Singer M 6720 (SGO) lished in 1952].
rhodotrama Singer in Singer & Digilio, 1951e: lodgeae Singer, 1989a: 23

421 HOLOTYPE: Costa Rica. Heredia, Sta. OTS
near "La Selva", 23 December 1979, Lodge
HOLOTYPE: Argentina. Jujuy: Termas de
294 (F)
Reyes, 5 January 1947, Garolera & Digilio
7681 (LIL, FH). [Published in 1952]. neotropica Singer, 1961b: 54
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 15
venososulcatus Singer, 1946a: 557
February 1956, Singer B 1173 (LIL). Singer
HOLOTYPE: USA. Florida: Dade County, Ma-
stated that this was a "nom. nov.", but it
prob-
theson Hammock, 29 October 942, Singer F
1
ably was a new species.

1344 (FH)
nobilior Singer, 1989a: 23
HOLOTYPE: Amazonas: Reserva
Brazil.
COLLYBIA B 11077 (INPA)
Ducke, 31 May 1979, Singer
acicola Singer in Singer &
Digilio, 195 le: 60 paranaguae Singer, 1955a: 768
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Brazil. Parana, Paranagua, 27 Sep-
San Javier, Cerro Alto del Taficillo, 6 January tember 1952, Singer B 406 (F)
1950, Singer T 830 (LIL). [Published in
peladae Singer, 1969a: 70
1952]. HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
albelloavellanea Singer, 1955a: 767 Chiveria, 7 May 1967, Singer M 6777 (SGO)
HOLOTYPE: Brazil. Parana, Paranagua, 27 Sep- phoebophila Singer in Singer & Digilio, 195 le:
tember 1952, Singer B 409 (F) 53
alkalivirens Singer, 1948c: 27 HOLOTYPE: Argentina. Tucumdn: Sierra de
HOLOTYPE: USA. Virginia: Giles County, Mt. San Javier, Anta Muerta, 16 April 1950, Sing-
Lake Biological Station, 23 June 1947, Singer er T 964 (LIL). [Published in 1952].
& Wilson 337 (FH) phylladophila Singer in Singer & Digilio,
amazonensis Singer, 1989a: 21 1951e: 55
HOLOTYPE: Brazil. Amazonas, Road Manaus- HOLOTYPE: Argentina. Tucuman: San Javier,
Caracarai,km 45, 10 August 1978, Singer B 24 April 1951, Singer T 1526 (LIL). [Pub-
11376 (INPA) lished in 1952].
batistae Singer, 1965e: 23 phylladophiloides Singer, 1989a: 24
HOLOTYPE: Brazil. Pernambuco: CaMaragibe, HOLOTYPE: Brazil. Amazonas: Lago Janauarf,
6 July 1960, Singer B 3102 (BAFC) 18 March 1983, Singer B 12444 (F)

COLLYBIA
rionegrensis Singer, 1989a: 24 fuscimarginata var.bispora Singer, 1977c: 126

HOLOTYPE: Brazil. Amazonas, Ponta Negra HOLOTYPE: Czechoslovakia. Prague, Podbaba,
near Manaus, 15 January 1978, Singer B 13 August 1974, Pouzar & Singer [Singer
10528 (INPA) no.] C 5557 (F)
sleumeri Singer, 1953b: 4 halophila Singer, 1959b: 397
HOLOTYPE: Argentina. Catamarca: Cerro Ne- HOLOTYPE: Afghanistan. West of Butschak:
gro, between Minas Capillitas and Cerro Ne- along road from Kabul to Nimla, 22 April
gro, 3 March 1952, Sleumer & Sparre [Singer 1951, Gilli CXXX1II/10 (LIL)
no.] T 1912 (LIL) hornana Singer &
Hausknecht, 1989c: 87
solitaria Singer, 1989a: 25 HOLOTYPE: Lower Austria: Dobern-
Austria.
HOLOTYPE: Brazil. Amazonas, 30 km N of dorf near Horn, 8 June 1987, Hausknecht S
Manaus, 29 June 1977, Singer B 10028 (INPA?) 7670 (WU 6201)
subhybrida Singer, 1965i: 151 izonetae Singer, 1989a: 103
HOLOTYPE: Argentina. Neuque"n: Quetrihue, HOLOTYPE: Brazil. Amazonas: 30 km N of
14 May 1952, Singer M 573 (LIL, BAFC) Manaus, 23 March 1978, Singer B 10855 (INPA)
syringea Singer, 1955a: 768 javanica Singer, 1989a: 103
HOLOTYPE:
Brazil. Parana, Paranagua, 27 Sep- HOLOTYPE: Indonesia. Java, von Hohnel 3860
tember 1952, Singer B 404 (F) (FH)
tetrachroa Singer in Singer &
Digilio, 1951e: kuhneriana Singer, 1969a: 212
52 HOLOTYPE: Argentina. Tucuman: near Arroyo
HOLOTYPE: Argentina. Tucuman: Tafi del Val- Mista, 27 October 1949, Singer T 722 (LIL)
le, Parque Aconquija, 5 March 1949, Singer lentispora Singer, 1950a: 134
T 199 (LIL). [Published in 1952]. HOLOTYPE: Argentina. Tucuman: San Pablo,
tucumanensis Singer in Singer & Digilio, 1951e:
27 March 1949, Singer T 316 (LIL)
47 leporina var. tetraspora Singer &
Hausknecht,
HOLOTYPE: Argentina. Tucuman: Sierra de 1988d: 112
San Javier, Anta Muerta, 16 April 1950, Sing- HOLOTYPE: Austria. Vienna: Lobau, near
er T 963 (LIL). [Published in 1952]. Kreuzgrundtraverse, 29 September 1984,
urubambae Singer, 1961b: 55 Hausknecht & Mrazek S 1441 (WU). Isotype: F
HOLOTYPE: Peru. Machu Picchu: Rio Urubam- lobauensis Singer & Hausknecht, 1988d: 107
ba, 20 May 1958, Singer B 3024 (LIL). HOLOTYPE: Austria. Vienna: Lobau, near
velutipes aestivalis Singer, 1922a:

f. 40 Kreuzgrundtraverse, 15 October 1982, Haus-
No type information. knecht S 910 (WU). Isotype: F
magnispora var. levis Singer in in Singer & Di-

CONOCYBE gilio, 195 le: 306
affinis Singer, 1969a: 207 San Anta Muerta, 20 October 1949,
Javier,
HOLOTYPE: Argentina. Tucuman, 10 Novem- Singer T 715 (LIL). [Published in 1952].
ber 1950, Singer T 1020 (LIL)
magnispora var. striatella Singer in Singer &
alba Singer in Singer & Digilio, 1951e: 295
Digilio, 1951e: 305
HOLOTYPE: Argentina. Tucuman: Tapia, 15 HOLOTYPE: Argentina. Tucuman, Singer S 43
April 1951, Singer T 1484 (LIL). [Published (LIL). [Published in 1952].
in 1952].
mazatecomm Singer, 1957a: 362
albocinerea Singer, 1941e: 99
Type information: Mexico. Oaxaca: Huautla de
HOLOTYPE: USSR. Oirotia: Kurai, 14/15 Au- Jimnez, 10-11 July 1957 (MICH). Two spec-
gust 1937, Singer & Vasilieva A 116 (LE). imens were cited: M
1503 and 1523, but M
Also published in 1950a: 130. no type was designated.
amazonica Singer, 1989a: 103 1953b: 18
megalospora ssp. nivalis Singer,
HOLOTYPE: Brazil.Amazonas: 30 km N of HOLOTYPE: Catamarca, between
Argentina.
Manaus, 11 May 1977, Singer B 9862 (INPA) Minas Capillitas and Cerro Negro, 3 March
diemii Singer, 1969a: 212 1952, Sleumer & Sparre [Singer no.] T 1933
HOLOTYPE: Argentina. Neuquen: Puerto Man- (LIL)
zano, 19 March 1963, Singer M 3075 (BAFC) mesospora subalpina Singer, 1989a: 104
var.
excedens van pseudomesospora Singer & Haus- HOLOTYPE: Switzerland. Vaud: above Pont de
knecht, 1992b: 95 Nant, 6 July 1984, Singer C 9682 (F)
HOLOTYPE: Austria. Vienna: Lobau, near Op- microsperma Singer, 1992b: 103
pidum Mhiilleiten, 20 June 1986, Hausknecht HOLOTYPE: Brazil. Bahia: Itabuna, 6 June
S 1424 (WU 5384). Isotype: F 1978, Singer B 11058 (INPA)
22 Chapter Three
CONOCYBE
minima Singer & Hausknecht, 1992b: 93 tenera f. excentrica Singer, 1950a: 140
HOLOTYPE: Austria. Pulkau: Feldberg near HOLOTYPE: Argentina. Buenos Aires: Garibal-
Grobreipersdorf, 7 September 1988, Haus- di, 30 May 1949, Singer S 44 (LIL)
knecht S 1794 (WU 7186). Isotype: F tenerrima Singer in Singer & Digilio, 195 le:
missionum Singer in Singer & Digilio, 195 le: 292
308 HOLOTYPE: Argentina. Tucumdn: Garden of
HOLOTYPE: Argentina. Misiones: Cataratas del the Instituto Lillo, 24 February 1951, Singer
Iguazii, 26 November 1949, Singer & Digilio T 1320 (LIL). [Published in 1952].
M 104 (LIL). [Published in 1952]. tenerrima var.monticola Singer, 1969a: 206
mixta Singer in Singer & Digilio, 195 le: 296 HOLOTYPE: Argentina. Tucuman: Taff del Val-
HOLOTYPE: Argentina. Tucuman: Taff del Val- le, 10 January 1950, Singer T 842 (LIL)
10 January 1950, Singer T 843 (LIL).
le,
tetraspora Singer, 1969a: 209
[Published in 1952]. HOLOTYPE: Argentina. Buenos Aires: San Cle-
myosura Singer, 1989a: 104 mente de Tuy, 21 October 1965, Singer S 552
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri- (BAFC)
beralta, 31 March 1956, Singer B 2403 (LIL) tuxtlaensis Singer, 1989a: 105
neoantipus var. carinthiaca Singer & Haus- HOLOTYPE: Mexico. Veracruz, Estacion Biol-
knecht, 1988d: 117 ogica de las Tuxtlas, 23 June 1969, Singer M
HOLOTYPE: Austria. Karnten: St. Veil a. d. 8185 (F)
Glan, near Tanzenberg, 26 September 1986, xerophytica Singer in Singer & Digilio, 195 le:
Forstinger and others (WU). Isotype: F 294
neoantipus var. tucumana Singer & Haus- HOLOTYPE: Argentina. Tucuman: Tapia, 15
knecht, 1988d: 115 April 1951, Singer T 1494 (LIL). [Published
HOLOTYPE: Argentina. Tucuman, 20 October
in 1952].
1949, Singer T 714 (LIL) xylophila Singer, 1969a: 208

proxima Singer in Singer & Digilio, 195 le: 293 HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16
HOLOTYPE: Argentina. Tucuman: Cerro Alto April 1967, Singer M 6531 (SGO)
del Taficillo, 4 March 1951, Singer T 1339
(LIL). [Published in 1952]. COPELANDIA
radicata Singer, 1953b: 18
HOLOTYPE: chlorocystis Singer & Weeks, 1979a: 469
Argentina. Tucuman: Anta Muer-
HOLOTYPE: USA. Florida: Dade County, Red-
ta, 11 January 1952, Singer T 1718 (LIL)
lands Fruit and Spice Park, Homestead, July
reticulatorugosa Singer in Singer & Digilio, 1976, Hearn (F)
1951e: 299
HOLOTYPE: Argentina. Tucuman: Taff del Val-
13 January 1950, Singer T 869 (LIL).
COPRINITES Poinar Jr. & Singer, 1990f: 1099
le,
(Fossil genus)
roberti Singer & Hausknecht, 1992b: 97 Type species: Coprinites dominicana Poinar, Jr.
HOLOTYPE: Austria. Maissau, 18 September & Singer

1984, Hausknecht S 1228 (WU 9220)
dominicana Poinar, Jr. & Singer, 1990f: 1099
stictospora Singer, 1989a: 104
HOLOTYPE: Dominican Republic. Santiago:
Cordillera septentrional, between Santiago
Manaus, 25 May 1977, Singer B 97 15 (INPA)
and Puerto Plata, The La Toca mine, AF-9- 1 1
striatipes var. sejuncta Singer & Hausknecht,
(Fossil species)
1988d: 119
HOLOTYPE: Argentina. Catamarca: Suncho,
near Santa Rosa, 20 January 1952, Singer T
COPRINUS
1792 (LIL) carbonicola Singer, 1969a: 176
subvelata Singer, 1950a: 139 HOLOTYPE: Argentina. Rio Negro: Nahuel
HOLOTYPE: Argentina. Tucuman: Taff del Val- Huapf National Park, Brazo Tristeza, 7 May
le, Parque Aconquija, Sierra de San Javier, 12 1964, Singer M
3975 (BAFC)
March 1949, Singer T 202 (LIL) herbivorus Singer, 1973d: 70
subxerophytica Singer & Hausknecht, 1992b: HOLOTYPE: Argentina. Jujuy: Lagunas de
80 Yala, 9 February 1965, Singer T 3929 (F)
HOLOTYPE: Austria. Vienna: Lobau, Leinerau hylaeae Singer, 1978h: 49
near Mittelwasser, 27 May 1984, Hausknecht HOLOTYPE: Ecuador. Napo: Lago Agrio, 17
S 1132 (WU 5223). Isotype: F May 1973, Singer B 7524 (F)

COPRINUS
hypsizygus Singer, 1969a: 177 kerrii Singer &Araujo, 1983f: 191

7 May 1965, Singer M 5576-5776a (BAFC) Caracarai, km 130, 24 March 1978, Kerr &
phylladophilus Singer, 1969a: 180 Singer [Singer no.] B 10861 (INPA)

HOLOTYPE: Argentina. Buenos Aires: Sierra kivaczensis Singer, 1949b: 95
de Tandil, Finca Pascuala, 16 November 1962, Type information: Finland. Karelia: Kivacz, 4
Singer S 365 (BAFC) September 1935 (LE?)
saccosporm Singer, 1969a: 179 megaspoms Singer, 1949b: 95
HOLOTYPE: Chile. Osorno: Lago Puyehue, 14 Type information: Finland. Karelia: Kivacz, 16
September 1967, Singer M

7507 (SGO) September 1935 (LE?)
sclerobasidium Singer, 1969a: 179 sphagneti Singer, 1949b: 95
HOLOTYPE: Argentina. Neuquen: Lago Lacar, Type information: Finland. Karelia: Kivacz, 30
Pucara, 6 November 1966, Singer 6014 M September 1935 (LE?)
(BAFC) umbilicatus Singer &
Araujo, 1983f: 195
subrenispermus Singer, 1969a: 182 HOLOTYPE: Brazil. Amazonas: Road Manaus-
HOLOTYPE: Chile. Santiago: Lagunas de Acu- Caracarai, km 125, 20 July 1978, Singer &
leo, 24 September 1967, Singer M 7789 Araujo [Singer no.] B 11313 (INPA)
(SGO) velutinellus Singer, 1949b: 96
uspallatae Singer, 1969a: 178 Type information: Finland. Karelia: Kivacz, 31
HOLOTYPE: Argentina. Mendoza: Upsallata, August 1935 (LE?)
14 January 1966, Singer (BAFC). No collec-
tion number given.
CRATERELLUS
CORIOLUS boyacensis Singer, 1963b: 239
HOLOTYPE: Colombia. Boyaca: Tunja, Arca-
fernandezianus Singer, 1959c: 374, 376 buco, Finca del Senor Luis Antonia Porras, 29
B 3598 (BAFC)
July 1960, Singer
Masa Quebrada Pangal, 16 February
Tierra,
1955, Sparre F 7 (S). Isotype: LIL?
CREPIDOTUS
CORTINARIUS &
acanthosyrinus Singer in Singer Digilio,
aberrans Signer, 1949b: 96 1951e: 411
Type information: Finland. Karelia: Kivacz, Au- HOLOTYPE: Argentina. Tucuman: Tapia, 3 July
gust 1936 (LE?) 1949, Singer T 591 (LIL). [Published in
amazonicm Singer & Araujo, 1983f: 193 1952].
HOLOTYPE: Brazil. Amazonas: Road Manaus- albidus var. bisporus Singer, 1973a: 436, 478
Caracaraf, km 45, 5 May 1980, Singer & Ar- HOLOTYPE: Brazil. Rio de Janeiro: Angra dos
aujo [Singer no.] B 12139 (INPA) Reis, 31 September 1952, Singer B 440 (FH)
arcticus Singer, 1938e: 15 albidus var. boliviae Singer, 1973a: 434, 478
Type information: USSR. (LE?) HOLOTYPE: Argentina. Jujuy: Valle del Rio
boyacensis Singer, 1963b: 242 Lozano, 18 February 1966, Singer T 5274 (F)
Type information: Colombia. Boyaca: Tunja, Ar- ampullicystis Singer, 1973a: 460, 478
cabuco, Finca del Senor Luis Antonio Porras, HOLOTYPE: Mexico. Veracruz: 7 km S of
no date, (BAFC). no collector named. Montepio, Estacion Biologica de las Tuxtlas,
campinaranae Singer & Araujo, 1983f: 196

19 June 1969, Singer M
8032 (F)
HOLOTYPE: Brazil. Amazonas: Road Manaus- applanatus var. subglobiger Singer, 1973a: 383,
Caracaraf, km 125, 22 May 1978, Singer & 478
Araujo [Singer no.] B 11021 (INPA) HOLOTYPE: Mexico. Oaxaca: San Augustustfn,
flavornatus Singer, 1931: 539 10 July 1969, Singer M 8400 (F)
Type information: USSR. Caucasus: Kha. aristoteliae Signer, 1959c: 398, 399
galeriniformis Singer & Araujo, 1983f: 194 HOLOTYPE: Chile. Juan Fernandez Island:
HOLOTYPE: Brazil. Amazonas: Road Manaus- Masa Tierra, Lower Quebrada Pangal, 24
Caracarai, km 125, 22 May 1978, Singer & March 1955, Sparre F 68 (S). Isotype: LIL?
Araujo [Singer no.] B 11026 (INPA) australis Singer, 1973a: 418, 478
gigasporus Singer, 1938e: 15 HOLOTYPE: Argentina. Misiones: Iguazii, Re-
Type information: USSR. Leningrad, December fugio Yaguaret6, 16 April 1957, Singer M
1935, Singer (LE?) 1030 (BAFC)
24 Chapter Three
CREPIDOTUS
austroandinus Singer, 1969a: 327 icterinm Singer, 1973a: 395, 479

HOLOTYPE: Chile. Llanquihue: Puella, 20 HOLOTYPE: Argentina. Tucuman: San Pedro
March 1959, Singer M 1868 (BAFC) de Colalao, 23 January 1955, Singer T 2058
bullulifer Singer, 1965i: 182 (LIL)
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, igapoensis Singer, 1989a: 121
28 March 1963, Singer M
3203 (BAFC) HOLOTYPE: Brazil. Amazonas: Igarape do Tar-
calolepis var. heterocystis Singer, 1973a: 469, umazinho, 12 March 1980, Singer B 12019
478 (INPA)
HOLOTYPE: Mexico. Veracruz: 7 km S of krieglsteineri Singer, 1988b: 69
Montepio, Estaci6n Biologica de las Tuxtlas,
HOLOTYPE: USA. Florida: Coral Gables, 28
19 June 1969, Singer 8029 (F)M September 1970, Gill (F)
calolepis ssp. polycystis Singer, 1973a: 475, 478

latifolius var. appendiculatus Singer, 1973a: 360,
HOLOTYPE: Costa Rica. San Jose: Cerro de las 479
Vueltas, 1 December 1925-1 January 1926, HOLOTYPE: Argentina. Tucuman: Horco Mol-
& Valeria n. le,30 April 1960, Singer T 3623 (BAFC)
Standley (BPI)s.
latifolius var. veraecrucis Singer, 1973a: 360,

calolepis ssp. tigrensis var. januarius Singer,
1973a: 474, 478
479
HOLOTYPE: Mexico. Veracruz: 3 km S of
San Javier, Ciudad Universitaria, 16 January Montepio, 30 July 1969, Singer 8761 (F) M
1955, Singer T 2020 (LIL) levisporus Singer, 1973a: 442, 479
HOLOTYPE: Mexico. Veracruz: Tlilapa, 24 June
catamarcae Singer, 1973a: 422, 478
HOLOTYPE:
1969, Singer M
8195 (F)
Argentina. Catamarca: Rio Chac-
luridus var. luridus Singer, 1973a: 419, 479
ras, 11 February 1952, Singer T 1821 (BAFC)
HOLOTYPE: Mexico. Morelos: of Tepoz-NW
citricolor Singer in Singer & Digilio, 1951e: 27 June 1969, Singer
tlan, 8247 (F)M
411 luridus var. minutus Singer, 1973a: 421, 479
HOLOTYPE: Argentina. Tucuman: Quebrada de HOLOTYPE: Mexico. Veracruz: 7 km S of
Lules, 18 December 1951, Singer T 1656
Montepio, Estacion Biologica de las Tuxtlas,
(LIL). [Published in 1952]. 19 June 1969, Singer M
8032a (F)
cristulatus Singer, 1957a: 369 luridus var. oaxacae Singer, 1973a: 420, 479
HOLOTYPE: Mexico. Oaxaca: Huautla de Jim- HOLOTYPE: Mexico. Oaxaca: Rancho del Cura
nez, 10 July 1957, Singer M
1516 (MICH). below Huautla, 7 July 1969, Singer 8317 M
Type information is from an annotation slip (F)
attached to the original publication. martini Singer, 1955a: 775
decipiens Singer, 1973a: 365, 478 HOLOTYPE: Panama. Zona Canalis: Barro Col-
HOLOTYPE: Mexico. Morelos: NW of Tepoz- orado Island, 7 July 1952, Martin & Welden
tlan, 18 July 1969, Singer M 8467 (F) 7494 (F)
defibulatus Singer in Singer & Digilio, 195 le: mexicanus Singer, 1973a: 413, 479
410 HOLOTYPE: Mexico. Veracruz: 7 km S of
HOLOTYPE: Montepio, Estaci6n Biologica de las Tuxtlas,
Argentina. Buenos Aires: Santa
Catalina, 7 June 1949, Singer S 90 (LIL).
8059 (F)
[Published in 1952]. molliformis Singer, 1973a: 441, 480

HOLOTYPE: Mexico. Morelos: NW
of Tepoz-
epigloeus Singer, 1973a: 462, 479 8262 (F) M
tlan, 1 July 1969, Singer
nephrodes var. meristocystis Singer, 1954b:
Montepio, Estacion Biol6gica de las Tuxtlas,
19 June 1969, Singer 8026 (F)M 146, 147
eucalypticola Singer, 1978h: 73 20 May 1952,
pf National Park, Quetrihue,
HOLOTYPE: Ecuador. Pichincha: above Quito,
M 792 (LIL)
Singer
5 May 1973, Singer B 7249 (F)
nephrodes var. minimus Singer, 1954b: 147
guzmanii Singer, 1973a: 415, 479 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
HOLOTYPE: Mexico. Veracruz: Coyame, 23
pf National Park, Isla Victoria, 17 May 1952,
June 1969, Singer M 8174 (F) Singer M 731 (LIL)
harperi Singer, 1959n: 586 palmarum Singer in Singer & Digilio, 195 le:
HOLOTYPE: USA. Virginia: Bedford, May 406
1919, Harper 1177 (F) HOLOTYPE: Argentina. Misiones: Cataratas del
harperi var. minor Singer, 1973a: 363, 479 Iguazu, 27 November 1949, Singer & Digilio
HOLOTYPE: Mexico. Dto. Federal: Cerro Oco- [Singer no.] M 135 (LIL). [Published in
piazco, 27 June 1969, Singer M 8216 (F) 1952].

CREPIDOTUS
parlatorei Singer in Singer & Digilio, 195 le: truncatus var. microcarpus Singer, 1973a: 378,
412 480
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Argentina. Tucuman: Rio Las Pa-
San Javier, 6 January 1950, Singer T 825 vas, 3 km W of Santa Rosa, 17 December
(LIL). [Published in 1952]. 1950, Singer T 1056 (LIL)
pilosiceps Singer, 1989a: 121 tuxtlae Singer, 1973a: 439, 480
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Mexico. Veracruz: 3 km S of
Manaus, 18 January 1978, Singer B 10556 Montepio, 22 June 1969, Singer M 8138 (F)
(INPA)
variisporus Singer in Singer & Digilio, 1951e:
podocarpi Singer in Singer & Digilio, 1951e:
413
413 HOLOTYPE: Argentina. Tucuman: Sierra de
HOLOTYPE: Argentina. Tucuman: Sierra de San Javier, Anta Muerta, 31 October 1949,
San Javier, 6 January 1950, Singer T 623
Singer T 741 (LIL). [Published in 1952].
(LIL). [Published in 1952].
velutinoaffinis Singer, 1961b: 78
polylepidis Singer, 1961b: 79 HOLOTYPE: Venezuela. Merida: Sierra de San-
HOLOTYPE: Venezuela. Merida: Sierra de San-
to Domingo, 31 July 1958, Dennis 1801 (K)
to Domingo, 24 July 1958, Dennis 1726 (K)
xanthocephalus Singer, 1973a: 396, 480
quitensis var. basicrinitus Singer, 1973a: 363,
480
le, Quebradita del Mastil, 17 February 1965,
Singer T 4083 (F)
le, Valle de San Javier, 15 August 1946, Di-
gilio & Grassi 791 (LIL) xanthophaeus Singer, 1973a: 455, 480
HOLOTYPE: Venezuela. Dto. Federal: Caracas
roseolus Singer, 1973a: 350, 480
HOLOTYPE:
Botanic Garden, 1
July 1958, Dennis 1128 (K)
Argentina.: Tucuman, Tucuman,
Jardin del Instituto M. Lillo, 20 February yungicola Singer, 1973a: 443, 480
1955, Singer T 2186 (LIL)
HOLOTYPE: Bolivia. La Paz: Nor-Yungas,
roseus Singer, 1947g: 87 Capillaria, 22 February 1956, Singer B 1335
Type information: USA. Florida: Dade Co., Mi- (BAFC). B 1422 cited as type in Latin diag-
nosis.
ami, Matheson Hammock, 30 October 1942,
Singer (FH). [Published in 1948].
rubriceps Singer, 1973a: 397, 480 CRINIPELLIS
HOLOTYPE: Brazil. Sao Paulo: Sierra de Can-
tareira, Reserva Forestal, 17 December 1964,
albipes Singer, 1976a: 36
Singer B 4098 (F) HOLOTYPE: Mexico. Veracruz: 7 km S of
schusteri Singer, 19895: 796
HOLOTYPE: Henderson Islands (South Pacific), 20 June 1969, Singer 8083 (F)M
13 May 1987, Schuster 00008 (F)
alcalivirens Singer, 1976a: 37
serotinus Singer, 196 Ib: 79
HOLOTYPE: Colombia. Cundinamarca: Salto
HOLOTYPE: Argentina. Tucuman: Arroyo Mis- de Tequendama, 24 July 1960, Singer B 3510
ta, 15 June 1955, Singer T 2782 (LIL)
(F)
sphaerosporus var. austrocedri Singer, 1969a:
austrorubida Singer, 1976a: 43
325
Sosas, 18 January 1950, Singer T 917 (LIL)
carecomoeis var. litseae Singer, 1942a: 489
stercorarius Singer in Singer & Digilio, 1951e:
HOLOTYPE: Philippines. Laguna: Mt. Makil-
409
HOLOTYPE: ing, 5 September 1917, Catalan 4464 (FH).
Argentina. Tierra del Fuego: Rio
M
400 (LIL). [Published carecomoeis subelata Singer, 1942a: 488
var.
ruary 1950, Singer
in 1952]. HOLOTYPE: New Caledonia. Col d'Amien,
& 1907, Le Rat 97 (FH). [Published in 1943].

sublevisporus Singer in Singer Digilio, 1951e:
410 catamarcensis Singer, 1953b: 12
HOLOTYPE: Argentina. Tucuman, 18 Decem- HOLOTYPE: Argentina. Catamarca: Valle de
ber 1948, Singer &
Digilio [Singer no.] T 25 Suncho, above Alto de la Junta, 17 January
(LIL). [Published in 1952]. 1952, Singer T 1745 (LIL)

tigrensis var. valdiviensis Singer, 1965i: 183 commixta Singer in Sing.& Digilio, 1953b: 11
HOLOTYPE: Chile. Valdivia: Rio Hueycolla, 30 HOLOTYPE: Argentina. Tucuman: Tapia, 6 Jan-
March 1963, Singer M 3243 (BAFC) uary 1952, Singer T 1710 (LIL)
26 Chapter Three
CRINIPELLIS
commixta var. junta Singer, 1976a: 33 mexicana Singer, 1976a: 29

HOLOTYPE: Argentina. Tucuman: Arroyo Mis- HOLOTYPE: Mexico. Veracruz: Jalapa, 12-20
ta, 15 June 1955, Singer T 2791 (LIL) December 1909, W. & E. Murrill 142 (NY)
coroicae Singer, 1976a: 27 mirabilis Singer, 1942a: 497
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- HOLOTYPE: New Caledonia. Between Col
roico, 26 January 1956, Singer B 558 (F)

d'Amien and Mgropo, 1907, Le Rat 101 (FH).
dicotyledonum Singer in Singer & Digilio, 1951e: [Published in 1943].
221
missionensis Singer in Singer & Digilio, 195 le:
HOLOTYPE: 226
Argentina. Tucuman: Rio de los
HOLOTYPE: Argentina. Misiones: Cataratas del
Sosas, 27 January 1951, Singer T 1140 (LIL).
27 November 1949, Singer & Digilio
Iguazii,
[Singer no.] M
123 (LIL). [Published in
dipterocarpi Singer, 1942a: 496
1952].
Type information: Vietnam. Tonkin, Eberhardt 483
patouillardii Singer, 1942a:
(FH). [Published in 1943].
HOLOTYPE: Japan. Bonin Island, Wright 525
eggersii var. epiphyllus Singer, 1967a: 48 (FH). [Published in 1943].
HOLOTYPE: Bolivia. La Paz: Nor Yungas,
phyllophila Singer, 1957a: 358
Charobamba, 30 January 1956, Singer B 747 HOLOTYPE: Mexico. Oaxaca: Huautla de Ji-
(LIL) menez, 10-12 July 1957, Singer 1507 M
eggersii var. flavipes Singer, 1976a: 47 (MICH)
HOLOTYPE: Bolivia. Pando: Madre de Dios, 28 piceae Singer, 1939a: 64
March 1956, Singer B 2310 (LIL) Type information: USSR. Altai Mtns.: near Ku-
eggersii var. lilaceiceps Singer, 1976a: 49 rai, August, Singer & Vasilieva (LE?)
HOLOTYPE: Ecuador. Napo: Lago Agrio, 9 podocarpi Singer in Singer & Digilio, 195 le:
May 1973, Singer B 7316 (F) 225
foliicola Singer, 1955a: 771 HOLOTYPE: Argentina. Tucuman: Cerro Alto
HOLOTYPE: USA. Florida: Highlands County, del Taficillo, 6 January 1950, Singer T 833
Highlands Hammock State Park, 2 September (LIL). [Published in 1952].

1942, Singer F 444a (F) pseudostipitaria ssp. occidentalis Singer, 1942a:
galeropsidoides Singer, 1989a: 66 470
HOLOTYPE: Brazil. Amazonas: Road Manaus- HOLOTYPE: Guadeloupe. Plateau des Rivieres,
Caracaraf, km 45, 20 December 1977, Aguiar Duss 492 (FH). [Published in 1943].
[Singer no.] B 10380 (INPA) pseudostipitaria ssp. occidentalis var. mesites

gracilis Singer, 1976a: 14 Singer, 1942a: 471
HOLOTYPE: Brazil. Pernambuco: Sao Louren- HOLOTYPE: Martinique. Pare du College de St.
co,Mata do Sao Joao , 16 July 1960, Singer Pierre, August 1900, Duss 1877 (FH). [Pub-
B 3412 (F) lished in 1943].
herrerae Singer, 1976a: 23 pseudostipitaria ssp. orientalis Singer, 1942a:

HOLOTYPE: Mexico. Veracruz: 7 km S of 472
HOLOTYPE: Vietnam. Tonkin: La Pho, 23 July
21 June 1969, Herrara [Singer no.] 8125 M 1909, Demange 275 (FH). [Published in
1943].
(F)
insignis Singer, 1976a: 45 purpurea Singer, 1976a: 44

HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri-
HOLOTYPE: Colombia. Valle: Cali, Saladito,
15 April 1968, Singer B 61 5 la (F)
beralta, 29 March 1956, Singer B 2320 (F)
rustica Singer, 1978h: 25
iopus Singer, 1939a: 64 HOLOTYPE: Ecuador. Tungurahua: Rio Topo,
Type information: USSR. Kazakhstan: in the bo-
30 April 1973, Singer B 7197 (F)
tanical garden of Alma Ata, May 1937, Nev-
odovski (LE?) sapindacearum Singer, 1976a: 39
HOLOTYPE: Brazil. Paraiba: Joao Pessa, 11
macrosphaerigera Singer, 1953d: 138 July 1960, Singer B 3377 (F)
HOLOTYPE: Brazil?. 24 January 1946, Sehnem
schini Singer, 1976a: 20
1917 (C8339) (LIL) HOLOTYPE: Argentina. Catamarca: Hill above
megalospora Singer in Singer & Digilio, 195 le: Estancia Santa Rosa, near Rio Campo, 20 Jan-
222 uary 1952, Singer T 1793 (LIL)
HOLOTYPE: Argentina. Tucuman: Sierra de septotricha Singer, 1942a: 468
San Javier, Anta Muerte, January, Singer T HOLOTYPE: Guadeloupe. Basse Terre, 14 July
1666 (LIL). [Published in 1952]. 1901, Duss 513 (FH). [Published in 1943].

CRINIPELLIS
sinensis Singer, 1989a: 67 CYSTOAGARICUS Singer, 1947b: 85

HOLOTYPE: China. Yunnan, between "Yung-
bei" and "Tschuhsiung (Tsuyung)", 13 May Type species: Nolanea strobilomyces Murr.
1915, Handel-Mazzetti 6298 (WU)
jujuyensis Singer, 1973d: 65
siparunae Singer, 1942a: 505 HOLOTYPE: Argentina. Jujuy: Lagunas de
Type information: USSR. Leningrad, Botanical T 5012/21
Garden, 10 June 1939, Singer (LE). [Pub-
sachaensis Singer, 1978h: 47
lished in 1943]. This species was also pub-
lished in 1943a: 115.
HOLOTYPE: Ecuador. Napo: Sacha Norte, Sa-
cha 4, 10 May 1973, Singer B 7357 (F)
tenuipilosa Singer in Singer & Digilio, 195 le:
228
CYSTODERMA
Lules, 24 January 1951, Singer T 1119 (LIL).
amianthinum f. olivaceum Singer, 1945f: 111
tucumanensis Singer, 1976a: 40 Type information: USSR. Siberia, Singer (LE)
HOLOTYPE: Argentina. Tucuman: Yerba Bue- amianthinum var. sublongisporum Singer, 1945f:
na, 12 February 1955, Singer T 2158 (LIL)
112
HOLOTYPE: USA. New York: Newcomb, Hun-
tingtonWild Life Forest, September 1941,
CYPTOTRAMA Singer, 1960a: 375
Singer 97 (FH)
Type species: Cyptotrama macrobasidium Singer

australe Smith &
Singer, 1945f: 97
dennisii Singer, 1973d: 34 theson Hammock, 22 October 1942, Singer F
HOLOTYPE: Mexico. Veracruz, 30 July 1969, 1184 (FH)
Singer M 8763 (F) austrofallax Singer, 1969a: 174
depauperatum Singer, 1978h: 43 HOLOTYPE: Chile. Valdivia: near Hueycolla, 4
HOLOTYPE: Colombia. Antioquia: Medellin, May 1967, Singer M
6682 (SGO)
Boqueron, 20 July 1974, Dumont CO 1750 caucasicum Singer, 1945f: 105
(F) Type information: USSR. Western Caucasus:
hygrocyboides Singer, 1969a: 90 near Guseripl (LE)
HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16 fallax Smith &
Singer, 1945f: 116
April 1967, Singer 6518 (SCO) M Type information: USA. Smith 17272 (MICH,
macrobasidium Singer, 1960a: 376 FH)
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, near granulosum var. typicum f. robustum Smith &
Coroico, 27 January 1956, Singer B 577 (LIL) Singer, 1945f: 87
niveum Singer, 1989a: 91 HOLOTYPE: USA. Michigan: Milford, 15 Sep-
HOLOTYPE: Amazonas: Igarape de Tar-
Brazil. tember 1938, Smith 10967 (MICH)
umazinho, 19 March 1980, Singer B 12039 paradoxum Smith &
Singer, 1948d: 454
(INPA) HOLOTYPE: USA. Oregon: Mt. Hood National
pauperum Singer, 1989a: 91 Forest, 19 October 1947, Smith 27987
HOLOTYPE: Brazil. Amazonas: Road Manaus- (MICH)
Itacoatiara, km 65, 6 October 1977, Maas ponderosum Smith & Singer, 1945f: 92
[Singer no.] B 10180 (F) HOLOTYPE: USA. Tennessee: Great Smoky
platense Singer, 1969a: 92 Mountain National Park, Mt. Le Conte, 26
HOLOTYPE: Argentina. Buenos Aires: Punta July 1942, Hesler 14390
(TENN)
Lara, May 1966, Singer S 739 (BAFC)
1
subpurpureum Smith & Singer, 1948d: 457
HOLOTYPE: USA. Oregon: Mt. Hood National
CYSTANGIUM & Forest,Cedar Creek at Skyline Road, 25 Sep-
Singer Smith, 1960h: 67
tember 1947, Smith 26824 (MICH)
Type species: Secotium sessile Massee & Rod-
way ex Rodway CYSTOLEPIOTA Singer in Singer & Digilio,
1951e: 281. [Published in 1952].
depauperatum Singer & Smith, 1960h: 69
HOLOTYPE: Chile. Llanquihue': Puella, 20
Type species: Cystolepiota constricta Singer
March 1959, Singer M 1921 (LIL)
pineti Singer, 1985: 417 albogilva Singer, 1989a: 98
HOLOTYPE: Mexico. Veracruz: 5 km N of Vi- HOLOTYPE: Brazil. Amazonas: Ponta Negra, 1
gos, 30 October 1982, Singer 9514 (F) M January 1978, Singer B 10445 (INPA)
28 Chapter Three
CYSTOLEPIOTA
amazonica Singer, 1989a: 99 DECONICA

acutiuscula Singer, 1941b: 86
Manaus, 18 May 1977, Singer B 9643 (INPA).
Type information: Finland. Karelia: near Kivacz
Although the type information is identical to in Caucaso Cr., Singer (LE?)
that of Marasmiellus amazoniensis Singer
alnetorum Singer in Singer & Digilio, 195 le:
(1989a: 26), the two species were based on
332
different collections judging from Singer's un-
published notebooks. See note under M. ama-
Yala, 8 April 1950, Singer T 955 (LIL). [Pub-
zoniensis.
lished in 1952].
aurantiaca Singer, 1973d: 65
HOLOTYPE: Chile. Santiago: near Ranque, 6
M 7122 (SCO) DENNISIOMYCES Singer, 1955d: 225
July 1967, Singer
australis Singer, 1969a: 167 Type species: Dennisiomyces glabrescentipes
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- Singer
pf National Park, Camino del Brazo Blest, los
March 1963, Singer 3153 M fuscoalbus Singer, 1989a: 88
Cantaros, 23
HOLOTYPE: Brazil. Amazonas: Manaus, 29
(BAFC) June 1977, Singer B 9896 (INPA)
brunneotingens Singer in Singer & Digilio, 1951e:
glabrescentipes Singer, 1955d: 225
283 HOLOTYPE: Brazil. Pernambuco: Recife, Dois
HOLOTYPE: Argentina. Tucuman: Quebrada de Irmaos, Jose de Silva & Batista 2223 (LIL)
Lules, 23 April 1950, Antonuccio [Singer no.]
rionegrensis Singer, 1989a: 88
T 977 (LIL). [Published in 1952]. HOLOTYPE: Brazil. Amazonas: Road Manaus-
cinereofusca Singer, 1982d: 37 Caracaraf, km 45, 11 June 1977, Singer B
HOLOTYPE: Costa Rica. Guanacaste: Parque 10071 (INPA)
Nacional Santa Rosa, 25 July 1981, Gomez &
Singer B 12453 (F) DERMOLOMA
constricta Singer in Singer & Digilio, 1951e:
283 aposcenwn Singer, 1989a: 88
HOLOTYPE: Mexico. Veracruz: 7 km S of Es-
tacion Biol6gica de Los Tuxtlas, 28 July 1969,
San March 1949, Singer T 173
Javier, 1 (LIL).
Singer M 8708 (F)
coryleti Singer & Clemen9on, 1971b: 120
luteifolia Singer in Singer & Digilio, 195 le: HOLOTYPE: Czechoslovakia. Kremsier, 30
282 September 1970, Singer C 5230 (F)
HOLOTYPE: Huanuco: Tingo Maria, 31
Peru.
pataguae Singer, 1989a: 89
October 1948, Carpenter 112 (LIL). [Pub- HOLOTYPE: Chile. Colchagua: Pumanque, 7
lished in 1952]. August 1967, Lazo PV-130 (SCO)
marthae Singer, 1969a: 168 yungense Singer, 1973d: 61
HOLOTYPE: Argentina. Neuquen: Quetrihue, 7 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 30
April 1963, M. Singer & Diem [Singer no.] January 1956, Singer B 700 (LIL, BAFC)
M 3296 (BAFC)
potassiovirens Singer, 1989a: 99 DESCOLEA Singer, 1950i: 256
Type species: Descolea antarctica Singer
Manaus, 31 November 1977, Singer B 10247
(INPA) antarctica Singer, 1950i: 257
pumanquensis Singer, 1973d: 66 HOLOTYPE: Argentina. Tierra del Fuego, 18
HOLOTYPE: Chile. Colchagua: near Pumanque, February 1950, Singer M 144a (LIL)
25 July 1967, Lazo PU-104 (SCO)
rosea Singer, 1969a: 168 DIACANTHODES Singer, 1945c: 141
Type Daedalea philippinensis Pat.
zano, Rio Bonito, 6 April 1963, Singer M species:
3292 (BAFC)
DICTYOPANUS
rubra Singer in Singer & Digilio, 195 le: 282
HOLOTYPE: Argentina. Tucuman: Sierra de pusillus var. pseudorhipidium Singer, 1953b: 14
San Javier, 30 March 1951, Singer T 1455 HOLOTYPE: Argentina. Tucuman: Anta Muer-
(LIL). [Published in 1952]. ta, 16 February 1952, Singer 1868 (LIL)

ECCILIA
ECCILIA aurantiaca Singer, 1974b: 33, 101

HOLOTYPE: Bolivia. Pando: Manuripi, Con-
undata var. ombrophila Singer, 1947c: 219 quista, 25 March 1956, Singer B 2199 (F)
Type information: collected in 1934.
calocera var. claudopus Singer, 1974b: 38, 101
HOLOTYPE: New Zealand. North Island: Wai-
ELASMOMYCES temata County, Waitarua, Waitakere Ranges,
July 1973, Beaver s. n. (F)
camphoratus Singer & Smith, 1960h: 62 dealbata Singer, 1955a: 763
HOLOTYPE: USA. Washington: Lake Quinault,
HOLOTYPE: Panama. Chiriqui: Casita Alta
5 November 1925, Brown s.n. (MICH)
above Boquete, 31 July 1952, Martin & Wei-
odoratus Singer & Smith, 1960h: 57
den 8070 (F)
HOLOTYPE: USA. Washington: Mt. Rainier
dumontii Singer, 1974b: 61, 102
National Park, 23 July 1948, Smith 29303
HOLOTYPE: Venezuela. Merida: El Pino, 16
(MICH) km E of Apartaderos, 19 July 1971, Dumont
roseipes Singer & Smith, 1960h: 64 and others VE 2480 (NY)
HOLOTYPE: USA. California: near Trinidad, no
echinata Singer, 1950e: 253
date, White 171 (MICH) HOLOTYPE: Argentina. Tucuman: Los Sosas,
zellerianus Singer & Smith, 1960h: 61 19 January 1950, Singer T 919 (LIL)
HOLOTYPE: USA. Washington: Lake Quinault,
fendleri Singer, 1974b: 32, 101
26 October 1925, Brown s.n. (MICH) HOLOTYPE: Brazil. Sao Paulo: Reserva Flores-
tal do Horto, Florestal, Serra de Cantareira, 17
ENDOLEPIOTULA Singer, 1963a: 260 December 1964, Singer & Furtado B4118 (F)
filopes Singer & Fidalgo, 1965f: 11
Type species: Endolepiotula ruizlealii Singer
HOLOTYPE: Brazil. Sao Paulo: Sao Paulo,
ruizlealii Singer, 1963a: 260 Parque do Estado, 1961, Furtado SP 60965
HOLOTYPE: Argentina. Mendoza: Santa Rosa, (SP)
Las Catitas, 10 March 1941, Ruiz Leal 7. 616 furfurella Singer, 1974b: 87, 103
(Ruiz Leal's herbarium). Fragments of the HOLOTYPE: Venezuela. Merida: Parque Na-
type at BAFC and MICH. cional Sierra Nevada, 25 July 1971, Dumont
and others VE 2991 (NY)
ENDOPTYCHUM heliconiae Singer, 1974b: 58, 101
HOLOTYPE: Venezuela. Sucre: Trail from Los
depressum Singer & Smith, 1958n: 216 Pocitos to Rio Grande Arriba, 13 July 1972,
HOLOTYPE: USA. Idaho, Seven Devils' Mtns., Dumont and others VE 5029 (NY)
23 August 1954, Smith 46579 (MICH) intermedia ssp. pulverulenta var. dicotyledonea
Singer, 1974b: 95, 104
ENTOLOMA HOLOTYPE: Trinidad. River Estate, Diego
Martin, 29 September 1949, Dennis 95 (K)
pyrenaica Singer, 1947c: 219 intermedia pulverulenta var. dennisiana
ssp.
Type information: Spain. Catalonia: Ribera de
Singer, 1974b: 95, 104
Sant Maurici, 1934.
HOLOTYPE: Venezuela. Merida: La Montana
above Merida, 30 July 1971, Dumont and oth-
EPICNAPHUS Singer, 1960f: 274, 279 ers VE 3454 (NY)
intermedia ssp. pulverulenta var. tetraspora
Type species: Epicnaphus phalaropus Singer
Singer, 1974b: 97, 104
phalaropus Singer, 1960f: 275, 279 HOLOTYPE: Venezuela. Trujillo: between Es-
HOLOTYPE: Pando: Manuripi, Con-
Bolivia. cuque, and La Mesa de San Pedro, 31 July
quista, 24 March 1956, Singer B 2175 (LIL) 1971, Dumont and others VE 3516 (NY)
mainsii Singer, 1974b: 60, 101
FAVOLASCHIA HOLOTYPE: Belize. El Cayo: Valentin, 25 June
1936, Mains s. n. (BPI)
alsophilae Singer, 1974b: 73, 102 meridae Singer, 1974b: 84, 103
HOLOTYPE: Colombia. Valle: Dagua road near HOLOTYPE: Venezuela. Merida: Parque Na-
km 25, 28 April 1968, Singer B 6443 (F) Laguna Negra, 18 July
cional Sierra Nevada,
andina Singer, 1974b: 68, 102 1971, Dumont and VE 2317 (NY)
others
HOLOTYPE: Colombia. Cundinamarca: Paramo montana Singer, 1974b: 70, 102
Chisaca-Sumapas, 8 May 1968, Singer B 7022 HOLOTYPE: Argentina. Jujuy: Lagunas de
(F) Yala, 11 February 1966, Singer T 5056 (F)
30 Chapter Three
FAVOLASCHIA
oligogloea Singer, 1974b: 78, 103 bisphaerigera var. grisea Singer, 1947c: 243
HOLOTYPE: Ecuador. Napo: Lago Agrio, 8 Type information: Spain. Catalonia, between
May 1973, Singer B 7278 (F) Bossost and Portillo, October 1934.
oligopora Singer, 1974b: 25, 101 deusta Singer & Clemenson, 197 Ib: 123
HOLOTYPE: Colombia. Valle: Call, Buenaven- HOLOTYPE: USA. Michigan: Cheboygan
tura, San Joaquin, 22 April 1968, Singer B County, Bryant Road, 24 July 1968, Clemen-
6279 (F) qon 680724-0 (LAU)
pantherina Singer, 1974b: 74, 103 dusenii var. griseilamellata Singer, 1952c: 220
HOLOTYPE: Argentina. Misiones: Iguazii, Re- HOLOTYPE: Argentina. Tierra del Fuego, Es-
fugio Yaguarete, 15 April 1957, Singer M tancia Nueva Argentina, 10 February 1950,
1003 (F)
Singer M 162 (LIL)
pterigena Singer, 1950e: 255 marthae Singer & Clemen9on, 1971b: 122
HOLOTYPE: Argentina. Tucuman: Sierra Mu- HOLOTYPE: Italy. Savoja: Val Feret, Frebouze,
noz, 13 January 1950, Singer T 862 (LIL) 8 June 1971, M. Singer [Singer no.] C 5383
pterigena var. boliviana Singer, 1974b: 56, 101 (F)
metuloidigera Singer, 1989a: 90
men Pampa, 1
February 1956, Singer B 817 HOLOTYPE: Argentina. Jujuy: Lagunas de
(F)
Yala, 9 February 1965, Singer T 3910 (F)
pterigena purpurea Singer, 1974b: 57, 101
var.
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Ca- tenuisperma Singer, 1969a: 146
taratas de San Juan, 28 January 1956, Singer
National Park, Arroyo Frutillas, 24 April
B 603 (F) pi
1965, Singer M 3620 (BAFC)
pubemla Singer, 1974b: 71, 102
HOLOTYPE: Venezuela. Dto. Federal: Mt. La tetrasphaerigera Singer, 1965e: 38
Naiguata, 25 June 1971, Dumont and others
HOLOTYPE: Brazil. Rio Grande do Sul: Gua-
VE 710 (NY) nabara, Floresta de Tijuca, 28 October 1961,
B 4046 (BAFC)
roseogrisea Singer, 1974b: 82, 103 Singer
HOLOTYPE: Colombia. Tolima: road from Ca- tilieti Singer, 1943a: 63
jamarca to Calana, 11 April 1968, Singer B Type information: USSR. Moldavia, August-
6035 (F) September.
sabalensis var. geonematis Singer, 1974b: 36,
101
FISTULINELLA
May 1973, Singer B 7441 (F)
alfaroae Singer & Gomez, 199 la: 77, 88
subamyloidea Singer, 1974b: 89, 104 HOLOTYPE: Costa Rica. Limon: Valle de la Es-
HOLOTYPE: Venezuela. Sucre: NW of Irapa, 9
Dumont and others VE 4425 (NY) trella, 5 August 1986, Singer, Gomez & Alfaro
July 1972,
[Singer no.] B 14621 (F)
teapae Singer, 1974b: 64, 102
HOLOTYPE: Mexico. Rio Teapa, between Ixta- campinaranae Singer, 1978a: 436
comitan and Solusuchiapa, 1
August 1969,
Singer M
8798 (F) Caracaraf, km 45, 18 July 1977, Singer B
10109 (INPA)
thwaitesii ssp. dybowskyana Singer, 1945g: 211
Type information: Uganda [Protectorate]. Near campinaranae var. scrobiculata Singer, 1983f:
Tropical Inst. at Mulange, Dummer (Patouilla) 148
thwaitesii ssp. zenkeriana Singer, 1945g: 211 HOLOTYPE: Brazil. Amazonas: Road Manaus-
Type information: Liberia. Oulah, August 1926, Caracarai, km 45, 31 March 1980, Webber &
Under (FH?) Singer [Singer no.] B 12089 (INPA)
varariotecta Singer, 1945g: 201 conica var. belizensis Singer, 1983f: 140
Type information: British West Indies. Grand HOLOTYPE: Belize. Augustine: Forest Station,
Etang, Thaxter (FH) 17 June 1976, Ivory S/114 (F)
violascens Singer, 1974b: 88, 103 &
wolfeana Singer Garcia, 199 la: 82, 88
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car- HOLOTYPE: Mexico. Hidalgo: Minas Viejas,
men Pampa, 17 February 1956, Singer B 1227 31 July 1981, Garcia 658 (F)
(F)
FAYODIA FLAMMULA
agloea Singer & Passauer, 1979c: 299
HOLOTYPE: Austria. Upper Austria, 2 May freindlingiae Singer, 1936g: 170, 171
1976, Moog/Passauer (W) Type information: USSR. (LE?)
BIOL08Y
191 HALL Index to Genera, Species, and Infraspecific Taxa 31
APR ?8
FLAMMULA
lenticeps Singer, 1941e: 97 antarctica Singer, 1962d: 13

Type information: Finland. Karelia, August, HOLOTYPE: Antarctica. Peninsula, Costa de
Freindling & Singer (LE?) Danco, 5 January 1958, Corte 1.024/CE
maackiae Singer, 1941e: 97 50.134 (IAA: Institute Antartico Argentine,
Type information: USSR. Khaborovsk: Oriens no longer exists).
Extr., September, Ljubarski (LE?) arenaria Singer, 1953d: 124
semiimbricata Singer, 1937: 240, 241 HOLOTYPE: Brazil. Rio Grande do Sul: Pareci,
Type information: USSR. Western Caucasus: In 2 November 1951, Singer B 70 (LIL)
valley Flum, Malaja Laba, spring 1936, Vas- asteliae Singer, 1969a: 315
iljeva (LE?) HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
30 April 1967, Singer M 6557 (SGO)
FULVIDULA austrocalyptrata Smith & Singer, 1964h: 82
alpina Singer, 1937: 239
pi National Park, Puerto Blest, path to Cn-
Type information: USSR. Caucasus: Mt.
taros, 23 April 1958, Pujols & Gamundi 50
Goultchi, In valle Urzuacski., July-August
(LIL). Isotype: MICH
1936, Vasilieva.
austrocalyptrata var. olivaceolamellata Singer,
janthinosarx Singer, 1941e: 92
1964h: 83
Type information: USSR. Altai Mtns.: Katun,
HOLOTYPE: Chile. Llanquihue: Puella, 21
July 1937, Singer (LE?)
March 1959, Singer 1945 (LIL)M
berteroana Singer, 1969a: 306
GALERINA HOLOTYPE: Argentina. Rio Negro: Nahuel
aberrant Smith & Singer, 1955e: 582 Huapi National Park, Brazo Tristeza, 6 May
HOLOTYPE: USA. Michigan: Cheboygan 1964, Singer M
3998 (BAFC)
County, Whitemore Lake, Mud Lake Bog, 25 boliviana Singer, 1964h: 58
September 1933, Smith 33-1024 (MICH) HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co-
acicola Smith & Singer, 1955e: 562 tapata, 28 January 1956, Singer B 628a
HOLOTYPE: USA. Oregon: Mt. Hood National (MICH)
Forest, Little Crater Lake, 5 October 1946, borealis Smith & Singer, 1958i: 470
Smith 24101 (MICH) HOLOTYPE: USA. Idaho: Granite Creek, near
agloea Smith & Singer, 1955e: 574 Nordman, 15 October 1956, Smith 54772
HOLOTYPE: USA. Washington: Mt. Rainier (MICH)
National Park, Carbon River, 2 October 1952,
bullulifera Singer in Singer & Digilio, 195 le:
Smith 40462 (MICH) 375
aimara Singer, 1964h: 112 HOLOTYPE: Argentina. Tucuman: Tafi del Val-
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- T
le, Parque Aconquija, 10 July 1949, Singer
tapata, 25 January 1956, Singer B 502 600 (LIL). [Published in 1952].
(MICH) & 470
californica Smith Singer, 1958i:
allospora Smith & Singer, 1955e: 585 HOLOTYPE: USA. California: Trinidad, 4 De-
HOLOTYPE: USA. Michigan: Cheboygan Coun-
cember 1956, Smith 56299 (MICH)
ty, Mud Lake Bog, 12 October 1953, Smith
44038 (MICH)
cascadensis Smith &
Singer, 1955e: 571
alpestris Singer, 1974a: 372
National Park, 19 October 1952, Smith 41144
HOLOTYPE: Switzerland. Valais, Glacier Lake
below Glacier de Moiry, 14 August 1971,
(MICH)
Singer 5494 (F)
castaneipes Smith & Singer, 1958i: 472
andina Singer, 1964h: 144
HOLOTYPE: USA. Oregon: Grants Pass, 12
November 1956, Smith 55523 (MICH)
25 January B 509 castanescens Smith & Singer, 1958i: 471
tapata, 1956, Singer
(MICH) HOLOTYPE: USA. Idaho: Granite Creek, Nord-
& man, 9 October 1956, Smith 54347 (MICH)
anelligera Smith Singer, 1958i: 475
HOLOTYPE: USA. Idaho: Payette Lakes, South cedretorwn var. austroandina Singer, 1964h:
fork of Lake Fork, 17 September 1956, Shaf- 281
fer (Smith no:) 53494 (MICH) HOLOTYPE: Argentina. Rio Negro: Nahuel
angustifolia Smith & Singer, 1964h: 317 Huapi National Park Rio Frias, path
,
to Paso
HOLOTYPE: USA. Colorado: Lizard Head de los Nubes, 16 March 1959, Singer M 1849
Pass, 10 August 1956, Smith 52000 (MICH) (LIL)
32 Chapter Three
GALERINA
cedretorum var. bispora Smith & Singer, 1964h: cerina var. yungensis Smith & Singer, 1964h:
280 78
HOLOTYPE: Smith 56162 (MICH). Type local- HOLOTYPE: Bolivia. La Paz: Nor-Yungas, near
ity not given. Catarata, San Juan, 24 January 1956, Singer

cedretorum \ar.filiformis Smith & Singer, 1964h: B 624 (MICH)
282 cingulata Singer, 1964h: 218
HOLOTYPE: USA. Washington: Mt. Rainier HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car-
National Park, near Green Lake, 17 October men Pampa, February, Singer B 1536 (MICH)
1952, Smith 41032 (MICH) cinnamomea Smith & Singer, 1958i: 473
cedretorum var. variabilis Smith & Singer, 1964h: HOLOTYPE: USA. Oregon: Grants Pass, 10
284 November 1956, Smith 55422 (MICH)
HOLOTYPE: USA. Colorado: San Juan Mtns., columbiana Singer, 1973d: 89
52126 (MICH)
13 August 1956, Smith HOLOTYPE: Colombia. Boyaca: Arcabuco, 29
cerina Smith & Singer, 1955e: 563 July 1960, Singer B 3608 (F)
Type information: USA. 7/88 (MICH) conspicua Singer, 1969a: 318

cerina var. ampullicystis Smith & HOLOTYPE: Argentina. Rio Negro: Nahuel
Singer, 1955e:
565 Huapf National Park, between Laguna Frias
and Paso de las Nubes, 24 March 1963, Singer
HOLOTYPE: USA. Michigan, 23 June 1953,
Singer N 142 (F)
M 3186 (BAFC)
cerina var. brachycystis Smith & Singer, 1955e: coquimbensis Singer, 1969a: 310
569
HOLOTYPE: USA. April 1967, Singer M
6532 (SCO)
Michigan: Pinckney, George
decipiens Smith & Singer, 1955e: 577
Reserve, 21 July 1933, Smith 33-606 (MICH)
HOLOTYPE: USA. Michigan: Emmet County,
cerina var. bresadolae Smith & Singer, 1955e: Mackinaw City Hardwoods, 7 June 1953,
568 N 431 (F)
Singer
HOLOTYPE: USA. Washington: Mt. Rainier &
decipiens var. separans Smith Singer, 1955e:
National Park, Green Lake, October 1952,
579
Smith 40994 (MICH)
HOLOTYPE: USA. Michigan: Emmet County,
cerina var. cerina f. bispora Smith & Singer, Wilderness Park, 29 July 1953, Smith 41739
1955e: 565 (MICH)
HOLOTYPE: USA. Michigan: near Whitemore dicranorum Smith & Singer, 1958i: 474
Lake, June 1951, Smith 44136 (MICH) HOLOTYPE: USA. Maine: S of Sinclair, 27 Au-
cerina var. contorticystis Smith & Singer, 1955e: gust 1956, Bigelow 4421 (MICH)
dimorphocystis Smith & Singer, 1955e: 558
570
HOLOTYPE: USA. Michigan: Luce County, HOLOTYPE: USA. Michigan: Luce County,
Tahquamenon Falls State Park, 17 August Falls State Park, 1953, Smith
Tahquamenon
1953, Singer N 1326 (F) 41401 (MICH)
cerina var. longicystis Smith & Singer, 1955e: dimorphocystis var. nexapensis Singer, 1964h:
567 46
HOLOTYPE: USA. Washington: Mt. Rainier HOLOTYPE: Mexico. Dto. Federal,near San Pe-
National Park, Carbon River, Smith 40453 dro de Nexapa, 21 July 1957, Singer 1566 M
(MICH) (MICH)
cerina var. luteovelata Smith & Singer, 1955e: discernibilis Singer, 1973d: 90
566 HOLOTYPE: Colombia. Valle: Cali, 15 April
HOLOTYPE: USA. Washington: Olympic Na- 1968, Singer B 6150 (F)
Hoh
tional Forest, River, 7 May 1939, Smith dominici Singer apud Dennis ex Singer, 1964h:
13191 (MICH) 125
cerina var. luteovelata f. occidentalis Smith & HOLOTYPE: Venezuela. MeYida: Sierra de San-
Singer, 1964h: 69 Domingo, Dennis 2506 (LIL)

to
HOLOTYPE: USA. Washington: Snoqualmie elaeophylla Singer, 1969a: 312

Pass, 26 September 1954, Smith 47903 HOLOTYPE: Chile. Santiago: Quebrada Diego,
(MICH) near Carmen, 6 July 1967, Singer 7115 M

cerina var. nebularum Smith & Singer, 1964h: (SGO)
76 emmetensis Smith & Singer, 1955e: 580
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Mt. HOLOTYPE: USA. Michigan: Emmet County,
Uchumachi, 25 February 1956, Singer B 1471 between Brutus and Pellston, June- July 1953,
(MICH) Singer N 83 (F, MICH)

GALERINA
emmetensis var. intermedia Smith & Singer, karstenii Smith & Singer, 1964h: 319
1955e: 581 HOLOTYPE: Finland. 9 September 1865, Kar-
HOLOTYPE: USA. sten (H)

Michigan: near Brutus, 6
41515 (MICH)
July 1953, Smith laeta Singer, 1964h: 185
evelata var.fulvipes Smith & Singer, 1964h: 86 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co-
HOLOTYPE: USA. Washington: near Longmi- tapata, 28 January 1956, Singer B 626
re, 11 October 1954, Smith 48836 (MICH)
(MICH)
fallax Smith &
Singer, 1955e: 561 larigna Singer, 1941e: 93
HOLOTYPE: USA. Smith 16239 (MICH) Type information: USSR. Altai Mtns., July,
& Singer & Vasilieva (LE?)

fallax f. subfiliformis Smith Singer, 1964h:
85 latispora Smith & Singer, 1955e: 591
HOLOTYPE: Canada. Quebec: Laurentide Park,
National Park, Castle Peak, 20 September
Mare du Sault, 2 September 1956, Smith
near
1952, Smith 40099 (MICH)
61758 (MICH)
latispora var. mexicana Singer, 1964h: 312
filiformis Smith & Singer, 1958i: 474 HOLOTYPE: Mexico. Dto. Federal: Paso de
HOLOTYPE: USA. California: Trinidad, 10 De-
M 1567 (MICH)
Cortes, 21 July 1957, Singer
cember 1956, Smith 56556 (MICH)
leucobryicola Smith & Singer, 1958i: 476
fuegiana Singer, 1953a: 242 HOLOTYPE: USA. Michigan: Washtenaw
County, Pinckney Recreation Area, 29 May
1955, Smith 49533 (MICH)
1950, Singer M 327 (LIL)
ruary
longinqua Smith & Singer, 1958i: 477
funariae Smith & Singer, 1955e: 594 HOLOTYPE: Macquarie Island: N of
Australia.
HOLOTYPE: USA. Washington: Kalaloch, Green Gorge, 29 March 1954, Bunt (LIL)
12063 (MICH)
April, Smith
luteolosperma Singer, 1955e: 570
gamundiae Singer, 1964h: 194 HOLOTYPE: USA. Michigan: Trout Lake, 4
HOLOTYPE: Argentina. Neuquen: Villa La An- October 1953, Smith 43722 (MICH)
gostura, Pujals & Gamundi 366 (LIL). Dupli-
BAFC.
cate material at
machangaraensis Singer, 1978h: 67
HOLOTYPE: Ecuador. Pichincha: near Quito,
glacialis Singer, 1941e: 92 Rio Machangara, 20 May 1973, Singer B
Type information: USSR. Altai Mtns., August,
7545 (F)
Singer (LE?)
macquariensis Smith & Singer, 1958i: 478
hypnorum var. evelata Singer, 1946g: 146
HOLOTYPE: Australia. Macquarie Island, Bunt
Type information: USA. New Hampshire: White 1950-51 (MICH)
Mtns., Mount Washington, August 1945, mainsii Smith & 479
Singer, 19581:
Singer WM-3 or WM-21 (FH). [Published in HOLOTYPE: USA. Montana: Glacier National
1948].
Park, Logan Pass, 20 August 1941, Mains
hypsizyga Singer, 1964h: 174 6116 (MICH?)
HOLOTYPE: Argentina. Minas
Catamarca:
mainsii var. tetraspora Singer, 1974a: 382
Capillitas, 1-3 March 1952, Singer T 1916 HOLOTYPE: Switzerland. Valais, along River
(LIL) C
Borgne de Ferpecle, 11 July 1971, Singer
inconspicua Singer, 1954b: 139, 140 5471 (F)
majalis Singer, 1989a: 116
National Park, Quetrihue, 16 May 1952, HOLOTYPE: USA. Illinois: Cook
pi
County, 26
Singer M 662 (LIL). Also published in Smith
May 1984, Singer N 4121 (F)
& Singer, 1955e: 592. marthae Singer, 1969a: 309
inconspicua var. mixta Singer, 1969a: 319 HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
HOLOTYPE: Argentina. Rio Negro: Nahuel 7 May 1965, Singer M 5589 (BAFC)
Huapi National Park, near Santa Maria, 19 megalocystis Smith & Singer, 1955e: 588
5064 (BAFC) HOLOTYPE: USA. No-
California: Trinidad, 30
inconspicua var. sublevis Singer, 1969a: 319 vember 1935, Parks & Smith 3680 (MICH)
HOLOTYPE: Argentina. Neuquen: Puerto Man- meridionalis Singer &
Clemenson, 1971b: 127
zano, 8 April 1963, Singer 3324 (BAFC) M HOLOTYPE: France. Alpes maritimes: 3 km
infernalis Singer, 1964h: 268 west of Valouris, 15 December 1970, Singer
HOLOTYPE: Argentina. Tucuman: Infiernillo, 3 C 5347 (F)
January 1960, Singer B 3046 (LIL) mesites Smith & Singer, 1955e: 587
jaapii Smith & Singer, 1955e: 574 HOLOTYPE: USA. Michigan: Emmet County,
Type information: Germany. See Fungi Selecti Mackinaw City Hardwoods, October, Smith
Exsiccati 10, VIII, 1903 44054 (MICH)
34 Chapter Three
GALERINA
minor Singer, 1953d: 125 perrara Singer, 1962d: 16

HOLOTYPE: Rio Grande do Sul: Pareci,
Brazil. HOLOTYPE: Antarctica. Peninsula, Costa de
2 November 1951, Singer B 65 (LIL) Danco, 17 February 1961, Corte 044 (IAA:
mollis Smith &
Singer, 1955e: 590 Institute Antartico Argentine, no longer ex-
HOLOTYPE: USA. Washington: Mt. Rainier ists)
National Park, Green Lake, 17 October 1952, phlegmacoides Singer, 1969a: 313
Smith 41074 (MICH) HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
montivaga Singer, 1969a: 306 Chi vena, 10 May 1965, Singer M 5711
HOLOTYPE: Argentina. Tucuman: Taff del Val- (BAFC)
le, 12 December 1958, Singer M
3161 (LIL)
physospora Singer in Singer & Digilio, 1951e:
mycenoides mamillata Singer, 1950h: 480
f. 381
HOLOTYPE: USSR. Altai Mtns. (LE). No Latin HOLOTYPE: Argentina. Tucuman: Taff del Val-
diagnosis. le, Parque Aconquija, 10 July 1949, Singer T
nana bohemica Singer, 1950h: 467
f. 597 (LIL). [Published in 1952].
Type information: USSR. Altai Mtns. (LE). No
polytrichorum Singer, 1964h: 194
Latin diagnosis. HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co-
naucorioides Smith &
Singer, 1955e: 584 tapata, 28 January 1956, Singer B 619
HOLOTYPE: USA. Michigan: Marquette Coun- (MICH)
Huron Mtns., September, Smith 33-995
praetervisa Singer, 1964h: 352
ty,
(MICH) HOLOTYPE: Argentina. Neuquen: Rincon, 18

nigripes Smith & Singer, 1958i: 480 March 1963, Singer 3064 (BAFC)M
HOLOTYPE: USA. Oregon: McLeod, near
proxima Singer, 1964h: 200
Grant's Pass, 15 November 1956, Smith 55691
HOLOTYPE: Argentina. Tierra del Fuego, Feb-
(MICH)
ruary, Singer M 187 (LIL)
nordmaniana Smith & Singer, 1958i: 480
pseudocamerina var. pargoloviana Singer, 1964h:
HOLOTYPE: USA. Idaho: Granite Creek, near
120
Nordman, 9 October 1956, Smith 54260
(MICH) Type information: USSR. Leningrad, 1937,
Singer (LE). Syntype: MICH
nubigena Smith & Singer, 1964h: 60
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Cer- pseudocerina Smith & Singer, 1958i: 483
B HOLOTYPE: USA. Colorado: San Juan Mtns.,
ro Uchumachi, 25 February 1956, Singer
Trout Lake, 14 August 1956, Smith 52182
1469 (MICH)
(MICH)
oligocalyptrata Singer, 1989a: 117
HOLOTYPE: Colombia. Boyaca: Arcabuco, 28 pubescentipes Smith & Singer, 1964h: 320
HOLOTYPE: USA. Idaho: Seven Devils' Mtns.,
July 1960, Singer B 3571 (F)
12 September 1956, Smith 53374 (MICH)
oreina Smith & Singer, 1955e: 593
HOLOTYPE: USA. Washington: Mt. Rainier pucarensis Singer, 1969a: 305
National Park, Snow Lake, Smith 40344 HOLOTYPE: Argentina. Neuquen: Pucard, 6
(MICH)
November 1966, Singer 6023 (BAFC)M
papillata Singer in Singer & Digilio, 195 le: pulchra Smith & Singer, 1964h: 101
382
National Park, near Green Lake, 19 October
HOLOTYPE: Argentina. Tucuman: Cerro Alto
1952, Smith 41205 (MICH)
(LIL). [Published in 1952]. quinteroensis Singer, 1969a: 316

HOLOTYPE: Chile. Valparaiso: Quintero, 27
patagonica Singer, 1954b: 140, 141
June 1967, Singer M 7004 (SGO)
pi, Gebiet, 16 May 1952, Singer M
677 (LIL) radicellicola Singer, 1964h: 353
& HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
payettensis Smith Singer, 1958i: 481
HOLOTYPE: USA. Idaho: Payette Lakes, Lake pf National Park, Puerto Blest, Los Cantaros,
Fork Creek, South Fork, 17 September 1956, 23 March 1963, Singer M
3154 (BAFC)
Smith 53491 (MICH) riparia Singer, 1953a: 249
peladae Singer, 1969a: 314 Type information: Argentina. Tierra del Fuego
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, (LIL). No type indicated. Type may be among
6 May 1967, M. Singer [Singer no.] M 6735 Singer's collections 314-315,M 240 and M M
(SCO) 352.
perangmta Smith & Singer, 1958i: 482 rostrata Smith & Singer, 1958i: 484
HOLOTYPE: USA. California: Humboldt State HOLOTYPE: USA. California: Orick, Prairie
Park, 30 November 1956, Smith 56007 Creek State Park, 5 December 1956, Smith
(MICH) 56364 (MICH)

GALERINA
sahleri var. sequoiae Smith & Singer, 1964H: 65 subtibiicystis Singer, 1953d: 146
HOLOTYPE: USA. California: Orick, 3 Decem- HOLOTYPE: Brazil. Rio Grande do Sul: Taim-
ber 1935, Smith 3745 (MICH) besinho, 5 November 1951, Singer B 102
saltensis Singer in Singer &
Digilio, 195 le: 382 (LIL)
HOLOTYPE: Argentina. Salta: Corralitos, 24 subtruncata Smith & Singer, 1958i: 486
March 1951, Singer T 1418 (LIL). [Published HOLOTYPE: USA. Washington: Mt. Rainier
in 1952]. National Park, St. Andrew's Creek, 24 Sep-
sanctixaverii Singer, 1973d: 91 tember 1952, Smith 40199 (MICH)
HOLOTYPE: Argentina. Tucuman: San Javier, taimbesinhoensis Singer, 1953d: 148
28 February 1962, Singer T 3776 (BAFC) HOLOTYPE: Brazil. Rio Grande do Sul: Taim-
semiglobata Singer, 1953d: 147 besinho, 5 November 1951, Singer B 101
HOLOTYPE: Brazil. Rio Grande do Sul: Taim- (LIL)
besinho, 5 November 1951, Singer B 100 tatrensis Singer, 1989a: 118
(LIL) HOLOTYPE: Czechoslovakia. Slovakia: Skal-
sequoiae Singer, 1989a: 118 nate Pleso, 5 September 1974, Singer C 5881
HOLOTYPE: USA. Samuel Taylor
California: (F)
Park, 6 February 1982, Mackler and others thujina Smith & Singer, 1955e: 595
[Singer no.] N 5005 (F) HOLOTYPE: USA. Michigan: Luce County,
simocyboides Singer, 1969a: 311 Tahquamenon Falls State Park, 1 1 September
HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16 1953, Smith 42589 (MICH)

April 1967, Singer 6529 (SCO)M triscopa f. longicystis Smith & Singer, 1958i:
stylifera var. badia Smith & Singer, 1958i: 486 489
HOLOTYPE: USA. Idaho: Kaniksu National HOLOTYPE: USA. Idaho: Payette Lakes, 30
Forest, Granite Creek, 6 October 1956, Smith
August 1954, Bigelow [Smith no.] 47110
54112 (MICH)
(MICH)
stylifera var. caespitosa Smith & Singer, 1964h: triscopa var. tetrascopa Smith & Singer, 1964h:
134 105
HOLOTYPE: USA. Michigan: Oakland County, HOLOTYPE: USA. Michigan, Smith 42157
Haven Highland Recreation Area, Octo-
Hill,
(MICH)
ber 1951, Smith 41223 (MICH)
tsugae Smith & Singer, 1955e: 586
stylifera var. velosa Smith &
Singer, 1958i: 487 HOLOTYPE: USA. Michigan: Cheboygan Coun-
HOLOTYPE: USA. Idaho, 1954, Smith 47201
ty, Mud Lake Bog, Smith 44070 (MICH)
(MICH) tundrae Smith & Singer, 1955e: 584
subarctica Smith & Singer, 1964h: 54 HOLOTYPE: USA. Washington: Mt. Rainier
HOLOTYPE: USA. Alaska: near Anchorage, 9
National Park, below Gobler's Knob, 12 Oc-
October 1959, Wells 1959 # 3 (MICH)
tober 1952, Smith 40858 (MICH)
subbadia Smith &
Singer, 1955e: 572 & 560
turfosa Smith Singer, 1955e:
HOLOTYPE: USA. Oregon: Blue River, 16 Oc-
HOLOTYPE: USA. Michigan: Whitemore Lake,
tober 1937, Smith 7845 (MICH)
September 1933, Smith 33-1101 (MICH)
subbullulifera Singer, 1953d: 137
uchumachiensis Singer, 1964h: 150
HOLOTYPE: Brazil. Rio Grande de Sul: Estacio
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Cer-
Sao Salvador, 9 November 1951, Singer B 126
ro Uchumachi, 25 February 1956, Singer B
(LIL)
1470 (MICH)
subceracea Smith &
Singer, 1955e: 559
velutipes Singer, 1964h: 284
HOLOTYPE: USA. Michigan: Wilderness Park,
7 October 1953, Smith 43847 (MICH)
HOLOTYPE: Argentina. Misiones: Puerto Igua-
& zu, 12 April 1957, Singer M 932 (LIL)

subcerina Smith Singer, 1958i: 485
HOLOTYPE: USA. Maine: Aroostook County, vexans Smith & Singer, 1955e: 583
near Guerette, 25 August 1956, Bigelow 4379 HOLOTYPE: USA. Michigan, Pt. Aux Chenes
and Wilderness Park, Smith 42824 (MICH)
(MICH)
subglabripes Smith & Singer, 1955e: 589
viatica Singer, 1964h: 231
National Park, Carbon River, 2 October 1952, men Pampa, 17 February 1956, Singer B 1247
Smith 40459 (MICH) (LIL)
subpapillata Singer & Dennis, 1964h: 221 victoriae Singer, 1954b: 140
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car- HOLOTYPE: Argentina. Neuque"n: Nahuel Hua-
men Pampa, 17 February 1956, Singer B 1240 pi National Park, Isla Victoria, 17 May 1952,
(LIL) Singer M 710 (LIL)
36 Chapter Three
GALERINA
vinaceobrunnea Smith & Singer, 1955e: 573 alutaceum Singer, 1978b: 155
HOLOTYPE: USA. Michigan: Burt Lake, Mil- HOLOTYPE: Switzerland. Singer C 5116 (F).
ler's Bog, 23 September 1949, Smith 34149 Isotype: LAU
(MICH) angustifolium Singer, 1959b: 379
vittaeformis var. albescens Smith & Singer, 1964h: HOLOTYPE: USA. Florida: Dade County, Mi-
333 ami, Brickell Hammock, 25 October 1942,
HOLOTYPE: USA. Smith 40128 (MICH) Singer F
1285 (FH)
vittaeformis f. bispora Smith & Singer, 1964h: bethlehemicum Singer, 1989a: 14

334 HOLOTYPE: Brazil. Para: Belem, 10 June 1966,
HOLOTYPE: USA. Washington: Mt. Rainier Singer B 4274 (BAFC)
National Park, 30 September 1952, Smith brunneum Singer, 1964e: 78
40340 (MICH) HOLOTYPE: La Paz: Nor-Yungas, 23
Bolivia.
vittaeformis var. pachyspora Smith & Singer, February 1956, Singer B 1399 (BAFC)
1958i: 488 bryogeton Singer, 1961b: 50
HOLOTYPE: Australia. Macquarie Island: West HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Ca-
March 1953, Bunt (LIL, MICH)
Point, 31
(LIL)
vittaeformis var. subannulata Singer, 1953a:
246 calongei Singer, 1970a: 67
HOLOTYPE: HOLOTYPE: Colombia. Valle: Municipalidad
Argentina. Tierra del Fuego: Rio
Dagua, 14 April 1968, Singer B 6122 (F)
ruary 1950, Singer 429 (LIL) M candidum Singer, 1989a: 14
HOLOTYPE: Brazil. Amazonas, Road Manaus-
yungicola Singer, 1964h: 114
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- Caracarai, km 45, 10 August 1978, Singer B
B 932 (MICH) 11372 (INPA)
roico, 5 February 1956, Singer
cheilocystidiatum Singer, 1970a: 68
HOLOTYPE: Colombia. Valle: between Cisne-
GALEROPSIS ros and Cordoba, 23 April 1968, Singer B
6300 (F)
andina Singer, 1963c: 107
HOLOTYPE: Argentina. Mendoza: San Carlos, collybiomorphum Singer, 1978b: 155
Arroyo de Cruz de Piedras, 20 January 1941,
Ruiz Leal 7287 (Ruiz Leal's herbarium). A du- Manaus, 18 May 1977, Singer B 9662 (INPA)
costaricense Singer, 1989a: 15
plicate of the holotype in BAFC.
andina 1963c: 107
var. gracilior Singer,
HOLOTYPE: Costa Rica. Heredia, above San
HOLOTYPE: Argentina. Mendoza: San Carlos, Jose de la Montana, 29 June 1981, Singer B
12520 (F)
Laguna del Diamante, 20 January 1941, Ruiz
Leal 7233 (Ruiz Leal's herbarium) cotapatae Singer, 1964e: 87
February 1956, Singer B 821 (BAFC)

GASTROBOLETUS
daguense Singer, 1989a: 15
& HOLOTYPE: Colombia. Valle: Com El Vergel,
fascifer Singer Smith, 1964J: 310
HOLOTYPE: Zimbabwe. Umtali 28
District,
dennisii Singer, 1964e: 81
May 1961, Chase 7549 (K)
& HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 9
laricinus Singer Both, 1977b: 60
HOLOTYPE: USA. New York: Olcott, 1 Octo- February 1956, Singer B 1056 (BAFC)
ber 1975, Both 1858 (BUF). Isotype: FH depauperatum Singer, 1951b: 601
HOLOTYPE: Argentina. Tucuman: Santa Rosa,
Along Rio de las Pavas, 17 December 1950,
GERRONEMA Singer, 195 Ib: 599
Singer T 1067 (LIL)
elasticum Singer, 195 Ib: 599
Type species: Gerronema melanomphax Singer
albogriseolum Singer, 1989a: 13 ta, 19 December 1949, Singer T 786 (LIL)
HOLOTYPE: Brazil. Amazonas, Road Manaus- elasticum ssp. amazonicum Singer, 1964e: 63
Caracaraf, km 125, 20 July 1978, Singer B HOLOTYPE: Bolivia. El Beni: Vaca Diez, 14
11291 (INPA) March 1956, Singer B 1978 (BAFC)
alexandri Singer, 1961b: 51 flammeum Singer, 1989a: 16
HOLOTYPE: USA. Michigan: Luce County, HOLOTYPE: Brazil. Amazonas, Road Manaus-
near Tahquamenon Falls, 23 June 1953, Sing- Caracarai, km 106, 4 April 1978, Singer B
er N 127 (F) 10916 (INPA)

GERRONEMA
hypolepargum Singer, 1978h: 9 xanthodictyon Singer, 1964e: 65

HOLOTYPE: Ecuador. Tungurahua: Rio Verde, HOLOTYPE: Bolivia. El Beni: Vaca Diez, 7
28 April 1973, Singer B 7162 (F) April 1956, Singer B 2546 (BAFC)
infundibuliforme Singer, 1964e: 79 yalae Singer, 1989a: 18
HOLOTYPE: Brazil. Rio Grande do Sul: Gua- HOLOTYPE: Argentina. Jujuy: Lagunas de
pore, Guajaramirim, 10 March 1956, Singer B Yala, 11 February 1965, Singer T 4011 (F)
1803 (BAFC)
josserandii Singer, 1961b: 49 GLOEOCANTHARELLUS Singer, 1945c: 140
HOLOTYPE: Netherlands. Terschelling, 16 July
1955, Jansen s.n. (L)
Type species: Cantharellus purpurascens Hes-
laccarioides Singer, 1989a: 16
ler
June 1977, Singer B 9936 (INPA)
leptosarcum Singer, 1964e: 61 GLOIOCEPHALA
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 6
March 1956, Singer B 1602 (BAFC) allomorpha Singer, 1976a: 290
majus Singer, 1964e: 64 HOLOTYPE: Colombia. Valle: Coir. El Vergel,
HOLOTYPE: Bolivia. Pando: Madre de Dios, 5 Municip. Dagua, km 25 N of Dagua road, 28
April 1956, Singer B 2495 (BAFC) April 1968, Singer B 6451 (F)
marchantiae Singer & Clemencon, 197 Ib: 119 alvaradoi Singer, 1976a: 300
HOLOTYPE: Switzerland. Valais: near the HOLOTYPE: Argentina. Jujuy: Lagunas de
Borgne de Ferpecle, 11 July 1971, Singer C Yala, 9 February 1965, Singer T 3909 (F)
5445 (F) anastomosans Singer, 1976a: 303
mariae Singer, 1989a: 17 HOLOTYPE: Ecuador. Napo: Shushufindi, 13
HOLOTYPE: Brazil.Amazonas: Manaus, 25 May 1973, Singer B 7392 (F)
April 1978, Souza [Singer no.] B 10997 capillata Singer, 1976a: 285
(INPA) HOLOTYPE: Mexico. Chiapas: between Finca
melanomphax Singer, 1951b: 602 Sospiro and El Pozo, 4 August 1969, Singer
HOLOTYPE: Argentina. Tucuman: near Los So- M 8979A (F)
sas, 1
January 1951, Singer T 1094 (LIL) cinnamomea Singer, 1976a: 289
melleum Singer & Clemen9on, 1971b: 120 HOLOTYPE: Mexico. Chiapas: between Finca
HOLOTYPE: Switzerland. Valais: near the Sospiro and El Pozo, 4 August 1969, Singer
Borgne de Ferpecle, 11 July 1971, Singer C M 8971 (F)
5453 (F) confusa Singer, 1960f: 269, 278
minutissimum Singer, 1969a: 52 HOLOTYPE: Brazil. Rio de Janeiro: Angra dos
HOLOTYPE: Chile. Llanquihue: Puella, 22 Reis,October 1952, Singer
1 B 426 (F)
March 1959, Singer 1998 (BAFC) M culmicola Singer, 1989a: 66
moseri Singer, 1983a: 277 HOLOTYPE: Ecuador. Pastaza: near road Am-
HOLOTYPE: Austria. Vienna: Prater, 2 October bato-Puyo, 24 July 1975, Dumont EC 1424
1904, von Hohnel (FH) (NY)
pantoxanthum Singer, 1989a: 17 inobasis Singer, 1960f: 263, 277
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Bolivia. Pando: Madre de Dios,
Manaus, 25 May B 9713 (INPA)
1977, Singer Santa Teresa, 21 July 1956, Singer B 2296
schusteri Singer, 1972a: 178 (LIL)
HOLOTYPE: Antarctica. Anvers Island, Norsel lamellosa Singer, 1960f: 265, 278
Point, February 1969, Schuster 69-3031 (BPI) HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car-
subpallidum Singer, 1959b: 378 men Pampa, 26 February 1956, Singer B
HOLOTYPE: Argentina. Tucuman: Rio de los 15285 (LIL)
Sosas, 1 March 1956, Singer T 2248 (LIL) longicrinita Singer, 1976a: 286
sucrense Singer, 1989a: 18 HOLOTYPE: Colombia. Valle: Cali, Buenaven-
HOLOTYPE: Venezuela. Sucre, 9 km N of road tura, San Joaqufn, 22 April 1968, Singer B
Rincon-Carupano-El Palar, 6 July 1972, Du- 6282 (F)
mont and others VE 4018 (NY) longifimbriata Singer, 1960f: 266, 278
theophili Singer, 1973d: 14 HOLOTYPE: Argentina. Tucuman: Sierra de
HOLOTYPE: Mexico. Veracruz, 7 km S of Es- San Javier, Ciudad Universitaria towards Es-
tacion Biologica de las Tuxtlas, 21 June 1969, tanciaSan Javier, 9 January 1960, Singer T
Singer M 8121 (F) 3520 (LIL)
38 Chapter Three
GLOIOCEPHALA
longisperma Singer, 1976a: 291 smithii var. xanthobasis Singer, 1946g: 150
HOLOTYPE: Brazil. Sao Paulo: Serra de Can- No type information. [Published in 1948].
tareira, Reserva Forestal, 17 July 1964, Singer subroseus var. homobasis Singer, 1946g: 150
B 4103 (F) No type information. [Published in 1948].
lutea Singer, 1976a: 301 vinicolor ssp. californicus Singer, 1946g: 149
HOLOTYPE: Ecuador. Napo: Sacha 4 (Sacha No type information. [Published in 1948].
Norte), 10 May 1973, Singer B 7333 (F)
mycenoides Singer, 1969a: 148 GOMPHUS

July 1967, Singer M 7161 (SCO) mamorensis Singer, 1983f: 17
palmarum Singer, 1960f: 264, 277 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Guayaramerin, 6 March 1956, Singer B 1614
Guayaramerin, 8 March 1 956, Singer B 1 744 (F)
(LIL) nothofagorum Singer, 1967b: 69
podocarporum Singer, 1976a: 294 HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
HOLOTYPE: Argentina. Jujuy: 15 km NE of 7 May 1967, Singer & Hernandez [Singer
20 February 1966, Singer T 5300
Yala, (F) no.] M 6776 (SAM)
quitensis Singer, 1976a: 299
HOLOTYPE: Ecuador. Pichincha: Quito, Que- GRIFOLA
brada de Miraflores, 24 April 1973, Singer B
7113 (F) gargal Singer, 1969a: 381
295 HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
religiosa Singer, 1976a:
15 km NE of Mirador, 6 May 1967, Singer M
HOLOTYPE: Mexico. Dto. Federal,road from
San Pedro de Nexapa to the Paso de Colon, 6934 (SCO)
16 July 1969, Singer MA- 10 (F)
sessilis Singer, 1976a: 303 GYMNOMYCES
HOLOTYPE: Ecuador. Pichincha: Quito, Que-
brada de Miraflores, 24 April 1973, Singer B alveolatus Singer & Smith, 1960h: 51
7119
HOLOTYPE: USA. Oregon: N of Corvallis, 23
(F)
May 1929, Zeller 6862 (NY)
spathularia Singer, 1960f: 271, 279
W cinnamomeus Singer & Smith, 1960h: 51
HOLOTYPE: Argentina. Salta: Oran, 3-4 km
of Rfo Blanco Bridge, North of Oran, 17
HOLOTYPE: USA. California: Berkeley, 7 Jan-
March 1955, Singer T 2308A (LIL) uary 1905, Gardner s.n. (NY)
compactus Singer & Smith, 1960h: 53
HOLOTYPE: USA. Oregon: near Sandy, 17
GOMPHIDIUS June 1937, Zeller 8286 (NY)
ferruginascens Singer & Smith, 1960h: 54

helveticus Singer, 1950d: 198
HOLOTYPE: USA. Idaho: McCall, 18 August
Type information: Switzerland. Vri Andermatt, 1958, Smith 60269 (MICH)
August 1950, Singer
parksii Singer & Smith, 1960h: 55
leptocystis Singer, 1946g: 148 HOLOTYPE: USA. California: near Los Gatos,
Type information: Canada. Vancouver (MICH). no date, Parks 477 (Z-27) (NY)
[Published in 1948]. roseomaculatus Singer & Smith, 1960h: 50
ochraceous ssp. muscigenus Singer, 1946g: 149 HOLOTYPE: USA. California: Saratoga, March
No type information. [Published in 1948]. This 1919, Parks Z-19 (NY)
subspecies was also published in Singer,
1949a: 472.
GYMNOPILUS
ochraceous ssp. muscigenus Singer, 1949a: 472
No type information. This subspecies was also chilensis Singer, 1969a: 303
published in Singer, 1946g: 149. HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
rutilus ssp. alabamensis Earle ex Singer, 1946a: 1967, Singer M 6679 (SGO)
535 excentriciformis Singer, 1969a: 302
subsp. novum for Comphidius alabamensis Earle HOLOTYPE: Chile. Llanquihue: 10 km N of
n. subnudum Colaco, 9 May 1967, Singer M 6805 (SGO)
septentrionalis Singer, 1946g: 149 humicola Harding ex Singer, 1961b: 76
No type information. [Published in 1948]. HOLOTYPE: USA. Michigan: Cheboygan
smithii Singer, 1946g: 150 County, east shore of Burt Lake, 3 July 1953,
No type information. [Published in 1948]. s.n. (F)

GYMNOPILUS
microloxm Singer, 1978h: 64 HEIMIOMYCES Singer, 1942c: 127

7474 (F) B Type species: Agaricus (Collybid) rheicolor Berk.
May 1973, Singer
& Curt.
pachycystis Singer, 1989a: 116
HOLOTYPE: Costa Rica. Alajuela: San Ramon, brunneipes Singer in Singer & Digilio, 195 le:
August 1981, Gomez 17916 (F) 242
pratensis Singer in Singer &
Digilio, 195 le: HOLOTYPE: Argentina. Tucuman: Las Len-
371 guas, 20 February 1951, Singer T 1309 (LIL).
HOLOTYPE: Argentina. Salta: Campo Quijano, [Published in 1952].
23 March 1951, Singer T 1415 (LIL). [Pub- pruinatipes Singer in Singer & Digilio, 1951e:
lished in 1952]. 243
radicicola Singer, 1978h: 65 HOLOTYPE: Argentina. Tucumdn: Riio de los
HOLOTYPE: Ecuador. Napo: Lago Agrio, 9 Sosas, 27 January 1951, Singer T 1129 (LIL).
May 1973, Singer B 7318 (F) [Published in 1952].
submarasmioides Singer, 1978h: 66
HOLOTYPE: Ecuador. Tungurahua: Rio Topo, HEMIMYCENA
amazonica Singer, 1989a: 67
GYRODON Manaus, 3 April 1978, Singer B 10887 (INPA)
adisianus Singer, 1989a: 123 angusta Singer, 1961b: 60
HOLOTYPE: Brazil. Amazonas: Ilha Marchan- HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 28
taria, 18 March 1983, Singer and others January 1956, Singer B 631 (LIL)
[Singer no.] B 12434 (F) crispuliformis Singer, 1973d: 35
exiguus Singer & Digilio, 1960i: 154 HOLOTYPE: Colombia. Valle: Cali, Buenaven-
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio tura, 20 April 1968, Singer B 6263 (F)
Yariza, 16 February 1956, Singer B 1216 depauperata Singer, 1943a: 122
(LIL) Type information: USSR. Altai Mtns. Central,
monticola Singer, 1957c: 256 July-August.
HOLOTYPE: Argentina. Tucuman: Taff del Val- diplocystis Singer, 1989a: 68
le, La Quebradita, 10 March 1950, Singer T HOLOTYPE: Brazil. Amazonas: Road Manaus-
928 (LIL) Caracaraf, km 45, 10 August 1978, Singer B
proximus Singer, 1945b: 244 11365 (INPA)
HOLOTYPE: USA. Florida: Alachua County, drepanocladi Singer, 1943a: 123
Devil's Millhopper, 29 June 1943, Singer F Type information: USSR. Altai Mtns. Central.
2425 (FH)
epibiotica Singer, 1961b: 61
proximus f. minor Singer & Gomez, 1990a: 53 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 13
HOLOTYPE: Costa Rica. Puntarenas: San Vito, February 1956, Singer B 1146 (LIL)
Wilson Botanical Garden, August 1986, Go-
epibrya Singer, 1943a: 121
mez 25144 (F) Type information: USSR. Altai Mtns. Central,
sibiricus Singer, 1938d: 174 August.
Type information: USSR. Altai Mtns.: Lake Te- guanacastensis Singer, 1989a: 68
lezkoje, September, Singer & Vasilieva (LE?) HOLOTYPE: Costa Rica. Guanacaste: Parque
Nacional Santa Rosa, 25 July 1981, Singer B
GYROPORUS 12464 (F)
herrerae Singer, 1973d: 36
phaeocyanescens Singer & Ivory, 1983f: 37
HOLOTYPE: Mexico. Dto. Federal, near San
HOLOTYPE: Belize. Augustine: Forest Station,
Pedro, 16 July 1969, Singer & Herrera s.n.
4 November 1976, Ivory S/358 (F)
(F)
longicystis Singer, 1961b: 61

HEBELOMA HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 30
gomezii Singer, 1983f: 188 January 1956, Singer B 707 (LIL)
HOLOTYPE: Costa Rica. Guanacaste: Parque nothofagi Singer, 1969a: 105
Nacional Santa Rosa, 25 July 1981, Singer & HOLOTYPE: Argentina. Rio Negro: Laguna
Gomez [Singer no.] B 12445 (F) Frias, 21 March 1963, Singer M
3104 (BAFC)
moseri Singer, 1969a: 287 patagonica Singer, 1969a: 106
HOLOTYPE: Argentina. Neuquen: Quetrihue, HOLOTYPE: Chile. Santiago: Cerro Robles, 19
15 April 1963, Moser s. n. (BAFC) June 1967, Singer M 6988 (SGO)
40 Chapter Three
HEMIMYCENA
pleurotiformis Singer, 1989a: 69 culmicola Singer, 1989a: 39

Brazil. HOLOTYPE: Colombia. Cundinamarca: Salto
Manaus, 3 June 1977, Singer B 9911 (INPA) de Tequendama, 25 July 1960, Singer B 3 503d
pseudoconidiophora Singer, 1938a: 196 (BAFC)

Type information: USSR. Altai Mtns.: Kurai, delasotae Singer, 1989a: 39
July, Singer & Vasilieva (LE?) HOLOTYPE: Argentina. Tucuman: Quebrada de
subtropicalis Singer, 1959b: 381
los Sosas, 18 December 1948, Singer & Di-
HOLOTYPE: Argentina. Tucuman: Tafi del Val- gilio [Singer no.] T 16 (LIL). Deposited as H.
le, Parque Aconquija, 17 February 1955, Sing- angustata (Berk.) Singer, see Singer & Digi-
er T 2185 lio, Lilloa 25: 110. 1953.
(LIL)
truncicola Singer, 1961b: 62 dimorphocystis Singer ex. Singer, 1969a: 63
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
San Javier, 12 January 1957, Singer T 2872 4 May 1967, Singer M 6880 (SCO)
(LIL) espeletiae Singer, 1989a: 40
HOLOTYPE: Colombia. Tolima: Santa Isabel,
Valle Rio Totarito, 6 February 1980, Boekhout
HERICIUM 589 (MEDEL)
caucasicum Singer, 1929b: 77 hydrogeton Singer, 1989a: 40

HOLOTYPE: Brazil. Amazonas: Una Marchan-
Type information: USSR. Caucasus. B 12213
taria, 1 July 1981, Singer (F)
izonetae Singer, 1989a: 41
HIATULA HOLOTYPE: Brazil. Amazonas: Igarap do Tar-
umazinho, 13 March 1983, Singer B 12424
melanoloma Singer, 1938e: 12 (F)
Type information: USSR. Leningrad, 1935, Vas- nigra var. microspora Singer, 1955a: 769
ilieva (LE?) HOLOTYPE: Panama. Zona Canalis: Corund,
13 August 1952, Martin & Welden 8353 (F,
HIATULOPSIS Singer & Grinling, 1967c: 364 IA)

patagonica Singer ex. Singer, 1969a: 61
Type species: Lepiota amara Beeli HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
pi National Park, between Laguna Frias and
aureoflava Singer, 1989a: 98 Paso Perez Rosales, 4 May 1961, Gamundi
HOLOTYPE: Brazil. Amazonas: Road Manaus- (BAFC)
Caracaraf, km 45, 28 February 1978, Singer pergelatinosa Singer, 1978h: 19, 21
B 10733 (INPA) HOLOTYPE: Ecuador. Napo: Lago Agrio, 8
HOHENBUEHELIA pycnophylli Singer, 1955a: 770
HOLOTYPE: Argentina. Jujuy: Tilcara, 23 Feb-
abietina Singer & Kuthan, 1980a: 61 ruary 1953, Sleumor (F)
HOLOTYPE: Czechoslovakia. Moravia: Besky- recedens Singer &
Kuthan, 1980a: 66
dy, Nature preserve Salajka near Bila., 21 July HOLOTYPE: Mexico. Dto. Federal: between
1974, Kuthan & Singer [Singer no.] C 5626 San Pedro de Nexapa and Paso de Corte, 16
(F) July 1969, Singer MA 0013 (F)
amazonica Singer, 1989a: 38 spegazzinii Singer ex. Singer, 1969a: 58

HOLOTYPE: Brazil. Amazonas: Manaus, 27 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
May 1977, Singer B 9727 (F) pf National Park, above Puerto Manzano, 1 1
bullulifera Singer in Singer & Digilio, 1951e:

April 1963, Singer M 3378 (BAFC). Invalidly
published in Sydowia 8: 117. 1954.
119
HOLOTYPE: tropicalis Singer, 1989a: 42
Argentina. Tucuman: Rio de los
Sosas, 27 January 1951, Singer T 1136a
Manaus, 3 April 1978, Singer B 70S97(INPA)
(LIL). [Published in 1952].
calongei Singer, 1973d: 30 HYBOGASTER Singer, 1964a: 13

HOLOTYPE: Colombia. Valle: Dagua, 14 April
1968, Singer B 6116a (F) Type species: Hybogaster giganteus Singer
campinaranae Singer, 1989a: 38 giganteus Singer, 1964a: 13
HOLOTYPE: Brazil. Amazonas: Road Manaus- HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
Caracaraf, km 125, 20 July 1978, Singer B Chiveria, between Union and Mirador, 29
1 1 32 7 (INPA) March 1963, Singer M 3229 (BAFC)

HYDROPUS
HYDROPUS chiriquiensis Singer, 1982e: 68

HOLOTYPE: Panama. Chiriqui: Cerro Respin-
africanus Singer, 1946g: 127 go, 6 km NW of Cerro Punta, 2 July 1975,
HOLOTYPE: Liberia. Du River, Firestone No. Dumont and others PA- 1805 (NY)
3, 27 July 1926, Under 130 (FH). [Published chlorinodorus Singer, 1982e: 25
in 1948]. HOLOTYPE: Brazil. Amazonas: Mata Ducke, N
of Manaus, 1 November 1977, Freire [Singer
albopubescens Singer, 1982e: 63
HOLOTYPE: Brazil. Amazonas: Road Manaus- no.] B 10254 (F)
citrinus Singer, 1978h: 34
Caracarai, at Estacao Experimental de Silvi-
cultura 11 1977, B HOLOTYPE: Ecuador. Napo: Lago Agrio, 16
Tropical, July Singer
10085 (F) May 1973, Singer B 7492 (F)
confertifolius Singer, 1982e: 50
albus Singer, 1978H: 39
HOLOTYPE: Amazonas: Road Manaus-
Brazil.
Caracaraf, 23 July 1978, Singer B 11338
(INPA)
altus Singer, 1982e: 74
corneri Singer, 1982e: 86
Manaus, 10 October 1977, Singer B 10190 (F) Manaus, Reserva Ducke, 28 June 1978, Sing-
amazonicm Singer, 1973d: 49 er B 11251 (INPA)
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 7 cystidiosus Singer, 1989a: 86
March 1956, Singer B 1639 (BAFC) HOLOTYPE: Brazil. Amazonas: 30 km N of
angustispermus Singer, 1982e: 59 Manaus, 13 July 1977, Singer B 10089
HOLOTYPE: Bolivia. Pando: Madre de Dios, (INPA)
Las Piedras, 5 April 1956, Singer B 2515 (F) decipiens Singer, 1982e: 115
beniensis Singer, 1982e: 17 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Bolivia. El Beni:Vaca Diez, Manaus, 13 July 1977, Singer B 10094
Guayaramerin, 12 March 1956, Singer B 1910 (INPA)
(LIL) depauperatus Singer, 1973d: 51
HOLOTYPE: Brazil. Para: Belem, 10 June 1966,
bisporus Singer, 1982e: 96
HOLOTYPE: Brazil. Amazonas: 30 km N of Singer B 4263 (BAFC)
dissiliens Singer, 1982e: 73
Manaus, 3 June 1977, Singer B 9909 (INPA)
bolivianus Singer, 1973d: 50
Manaus, 9 February 1978, Singer B 10652
(INPA)
dubius Singer, 1982e: 58
brunneobasis Singer, 1978h: 33 HOLOTYPE: Brazil. Amazonas: 30 km N of
Manaus, 30 September 1977, Singer B 10153
(F)
caespitosus Singer, 1982e: 32 dusenii ssp. nivifugus Singer, 1973d: 52
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Argentina. Buenos Aires: near La
Manaus, 6 April 1977, Singer B 9625 (INPA) Plata, Punta Lara, May 1966, Singer S 734
californicus Singer, 1989a: 85 (BAFC)
HOLOTYPE: USA. Marin County,
California: eitenianus Singer, 1973d: 52
19 November 1981, Calhoun 81-2590 (F) HOLOTYPE: Brazil. Amazonas, Manaus, no
and others 5205 (NY)
date, Eiten
camaragibensis Singer, 1973d: 51
HOLOTYPE: Brazil. Pernambuco: Camaragibe, etenianus van obscurus Singer, 1982e: 105
B 3298 (BAFC)
7 July 1960, Singer HOLOTYPE: Brazil. Amazonas: near Punta Ne-
Rio Negro, 15 January 1978,
gra, left side of
campinaranae van adnatifolius Singer, 1982e:
M. & B 10529 (INPA)
R. Singer [Singer no.]
89
excentricus Singer, 1982e: 49
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ivon,
Caracaraf, 21 March 1980, Singer B 12058 (F)
campinaranae van campinaranae Singer, 1982e:
fimbriatus Singer, 1982e: 81
88 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Brazil. Amazonas: Road Manaus- Manaus, 18 May 1977, Singer B 9661 (INPA)
Caracaraf, 21 March 1980, Singer B 12056 (F) van aporpus Singer, 1982e: 34
fluvialis
cavipes van murinialbus Singer, 1982e: 45 HOLOTYPE: Brazil. Amazonas: Road Manaus-
HOLOTYPE: Brazil. Amazonas: 30 km N of Caracaraf, 5 May 1980, Singer B 12134-
Manaus, 6 July 1977, Singer B 10045 (INPA) 12134a (F)
42 Chapter Three
HYDROPUS
fluvialis var. fluvialis Singer, 1982e: 33 hypopolius Singer, 1982e: 64

HOLOTYPE: Amazonas: Igarap do Tar-
Brazil. HOLOTYPE: Brazil. Amazonas: 30 km N of
umazinho, 19 March 1980, Singer B 12047 Manaus, 1 May 1977, Singer B 9859 (INPA)
1
(F) immutabilis Singer, 1982e: 102

fragilior Singer, 1982e: 67 HOLOTYPE: Brazil. Amazonas: Road Manaus-
HOLOTYPE: Brazil. Amazonas: 30 km N of Caracarai, 20 December 1977, Singer B 10374
Manaus, 18 May 1977, Singer B 9677 (INPA) (INPA)
fraterniger Singer, 1946f: 227 indicus Singer, 1989a: 87
HOLOTYPE: USA. Florida: Highlands County, HOLOTYPE: India. Punjab: near Patiala, no
Highlands Hammock State Park, Singer F 643 date, Saini 19 (F)

(FH) lacuster Singer, 1978h: 36
fuscoalbus Singer, 1982e: 91

HOLOTYPE: Brazil. Amazonas: Road Manaus- May 1973, Singer 7532 (F)
Caracarai, 30 December 1977, Singer B 10379 lacuster var. riparius Singer, 1982e: 79
(INPA) HOLOTYPE: Brazil. Amazonas: Igarap do Tar-
umazinho, 16 April 1980, Singer B 12103 (F)
fuscogriseus Singer, 1989a: 86
HOLOTYPE: Brazil. Amazonas: Road Manaus- levis Singer, 1982e: 72
Caracaraf, km 110, 4 April 1978, Singer B

10925 (INPA) December 1977, Araujo [Singer no.] B 10388
(INPA)
fuscomycelinus Singer, 1982e: 112
HOLOTYPE: Argentina. Jujuy: Lagunas de lipocystis Singer, 1969a: 141

pf National Park, Arroyo Frutillas, 9 April
gerronematoides Singer, 1969a: 143
HOLOTYPE: Chile. Valdivia: 1/4
Hueycolla,
lipocystis var. bisporis Singer, 1969a: 142
May 1967, Singer M 6688 (SCO)
glabripes Singer, 1969a: 144
HOLOTYPE: Chile. Valparaiso: near Quintero,
Huapf National Park, near Santa Maria, 30
27 June 1967, Singer & M 701 l/QUI-5
3705 (BAFC)
Lazo
lutescentipes Singer, 1982e: 35
(SCO)
HOLOTYPE: Brazil. Amazonas: Reserva Ducke,
gomezii Singer, 1982e: 119 25 km N of Manaus, 7 January 1979, Singer
HOLOTYPE: Costa Rica. Guanacaste: Parque
B 11478 (INPA)
12472 (F) maculatipes Singer, 1982e: 82
gracilis Singer, 1982e: 117 Caracaraf, 10 August 1978, Singer B 11362
HOLOTYPE: Brazil. Amazonas: Irarap do Tar-
(INPA)
marasmioides Singer, 1973d: 54
(F)
HOLOTYPE: Colombia. Boyaca: between Ra-
guaporensis Singer, 1982e: 106 miriquf and Rondon, 30 July 1960, Singer B
HOLOTYPE: Brazil. Rondonia: Guajara mirim, 3628 (BAFC)
10 March 1956, Singer B 1786 (F)
martensii Singer, 1989a: 87
heterocystis Singer, 1973d: 53 HOLOTYPE: China. Yunnan: Mt. Yun Shan, no
HOLOTYPE: Argentina. Catamarca, 2 March date, Handel Mazzetii 11.207 (WU)
1952, Singer T 1914 (MICH, LIL)
megalobasidium Singer, 1982e: 83
humilior Singer, 1982e: 83 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Brazil. Amazonas: 30 km N of Manaus, 23 June 1977, Singer B 10013 (F)
Manaus, 31 October 1977, Singer B 10245 (F) mesites Singer, 1982e: 98
hydrophoroides Singer, 1982e: 71 HOLOTYPE: Brazil. Pernambuco: Dois Irmaos,
HOLOTYPE: Colombia. Valle: Cali, Buenaven- 14 April 1964, Singer B 4079a (F)
Calima, 18 April 1968, Singer B 6204
tura,
microcephalus Singer, 1982e: 80
(F) HOLOTYPE: Brazil. Amazonas: Road Manaus-
hygrophilus Singer, 1973d: 54 Caracaraf, 28 February 1978, Singer B 10718
HOLOTYPE: Colombia. Valle: Anchicaya, 26 (INPA)
April 1968, Singer B 6396 (F) microsporus Singer, 1969a: 142
hyperythms Singer, 1982e: 75 HOLOTYPE: Argentina. Neuquen: Lago Cor-
HOLOTYPE: Brazil. Amazonas: 30 km N of rentoso, 16 April 1965, Singer M
5011
Manaus, 18 May 1977, Singer B 9657 (F) (BAFC)

HYDROPUS
montanus Singer, 1982e: 103 sabalis Singer, 1946c: 119

HOLOTYPE: Argentina. Tucumdn: Tafi del Val- Type information: USA. Florida.
le, Quebrada del Mastil, 17 February 1965, semimarginellus Singer, 1973d: 59
Singer T 4082 (F) HOLOTYPE: Brazil. Para: Belem, 10 June 1966,
mutabilis Singer, 1978h: 36 Singer B 4251 (BAFC)
HOLOTYPE: Brazil. Guajard mirim, 10 March stenocystis Singer, 1978h: 38
1956, Singer B 1781 (F) HOLOTYPE: Ecuador. Tungurahua, steep moun-
mycenoides var. minor Singer, 1982e: 100 tainside S of Banos, 29 April 1973, Singer M
Brazil. 7181 (F)
subcartilagineus var. aquosus Singer, 1982e: 31
mycenoides var. pusillus Singer, 1982e: 100 HOLOTYPE: Amazonas: Road Manaus-
Brazil.
HOLOTYPE: Brazil. Amazonas: 30 km N of Caracarai, 20 December 1977, Singer B 10369
(INPA)
(INPA)
subcartilagineus var. bisporiger Singer, 1982e:
nanus Singer, 1982e: 42
29
HOLOTYPE: Argentina. Buenos Aires: Santa
Manaus, 17 May 1977, Singer B 9642 (INPA)
Catalina, 19 March 1966, Singer S 638
nigromarginatus Singer, 1982e: 60
HOLOTYPE: Brazil. Amazonas: Road Manaus- (BAFC)
Caracarai, 21 March 1980, Singer B 12063 (F) subcartilagineus var. napoensis Singer, 1982e:
occidental Singer, 1982e: 100 30
HOLOTYPE: Colombia. Valle: Cali, Buenaven- HOLOTYPE: Ecuador. Napo: Shushufindi, 15
tura, Calima, 18 April 1968, Singer B 6217 May 1973, Singer B 7470 (F)
(F) terraefirmae Singer, 1982e: 90

omphaliniformis Singer, 1973d: 55
HOLOTYPE: Colombia. Valle: Cali, Buenaven- Manaus, Reserva Ducke, 6 October 1978,
tura, 19 April 1968, Singer B 6240 (F) Freire [Singer no.] B 11399 (INPA)
papillatus Singer, 1973d: 56 tucumanus Singer, 1982e: 18

HOLOTYPE: USA. Florida: Alachua County, 20 HOLOTYPE: Argentina. Tucuman: Chicligasta,
July 1943, Singer F 2857 (F) Rio Cochuna, 11 February 1952, Singer T
phyllogenus bahiensis Singer, 1982e: 93
var. 1823 (LIL)
HOLOTYPE: Brazil. Bahia: Itabuna, 5 June veraecrucis Singer, 1973d: 59
1978, Singer B 11085 (INPA) HOLOTYPE: Mexico. Veracruz: near Montepio,
phyllogenus phyllogenus Singer, 1982e: 92
var. 21 June 1969, Singer M
8102 (F)
HOLOTYPE: Brazil. Amazonas: Manaus, 7 Feb-
viscidulus Singer, 1969a: 145
ruary 1978, Singer B 10609 (INPA) HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
pinetorum Singer, 1973d: 57 1967, Singer M 6703 (SGO)
HOLOTYPE: USA. Florida: Miami, 26 October
xanthosarx Singer & Grinling, 1967c: 362
1942, Singer F 1290 (F)
HOLOTYPE: Zaire. Brazzaville: Fl. Djoue,
platensis Singer, 1973d: 58 March 1965, Grinling 50,313 (BAFC)
HOLOTYPE: Argentina. Buenos Aires: Punta
Lara, 1 May 1966, Singer S 741 (BAFC) xerophilus Singer, 1973d: 60
69
HOLOTYPE: Argentina. Salta: Gemes, 7 Feb-
platycystis Singer, 1982e:
HOLOTYPE: Brazil. Amazonas: 30 km N of ruary 1957, Singer T 2971 (BAFC)
Manaus, 20 October 1977, Singer B 10230 xuthophyllus Singer, 1973d: 61
(INPA) HOLOTYPE: Argentina. Tucuman: Burruyacu,
19 February 1957, Singer T 2991 (LIL, BAFC)
pyxidatoides Singer, 1969a: 140
HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
7 May 1965, Singer M 5596 (BAFC)
recedens Singer, 1982e: 116
HYGROASTER Singer, 1955f: 370
HOLOTYPE: Argentina. Tucumn: near Potrer-
illos, 10 February 1952, Singer T 1819 (F) Type species: Hygrophorus nodulisporus Den-
nis
riograndensis Singer, 1953d: 122
HOLOTYPE: Brazil. Rio Grande do Sul: Pareci,
2 November 1951, Singer B 68 (LIL) albellus Singer, 1989a: 6
rionegrensis Singer, 1982e: 70 HOLOTYPE: Brazil. Amazonas: 30 km N of

HOLOTYPE: Amazonas: Manaus,
Brazil. 11 Manaus, 3 April 1978, Singer & Araujo [Sing-
October 1977, Singer B 10202 (F) er no.] B 10874 (INPA)
44 Chapter Three
HYGROCYBE
HYGROCYBE rhodoleuca Singer, 1973d: 6

HOLOTYPE: Brazil. Paraiba: Joao Pessoa, 11
amazoniensis Singer, 1989a: 3 B 3345
July 1960, Singer (LIL, BAFC)
rubroflava Singer, 1973d: 6
Manaus, 13 July 1977, Singer B 10097
HOLOTYPE: Colombia. Valle: Cali, Buenaven-
(INPA)
tura, 22 April 1968, Singer B 6276 (F)
ammophila Singer, 1968a: 129 & Kuthan, 1976g: 7
streptopus Singer
HOLOTYPE: Uruguay. Punta del Este, 5 June HOLOTYPE: Czechoslovakia. Slovakia: Cadca,
1965, Eskuche [Singer no.] B 4138 (BAFC)
Rakova, 27 July 1974, Kuthan & Singer
araucana Singer, 1969a: 11
[Singer no.] C 5652 (F)
striatella Singer, 1969a: 20
Mirador, 5 May 1965, Singer M 5516 (BAFC) HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
batistae Singer, 1965e: 20 M 6732 (SGO)
1967, Singer
HOLOTYPE: Brazil. Pernambuco: Recife, no
da Silva & Batista s.n. (LIL)
subheteromorpha Singer, 1969a: 12
date,
HOLOTYPE: Chile. Concepcion: Boca del Bio-
blestiana Singer, 1969a: 18
HOLOTYPE:
Bio, 17 November 1967, Singer M 7818
Argentina. Rio Negro: Nahuel
(SGO)
Huapi National Park, Brazo Blest, 23 March swanetica Singer, 1931: 522
1963, Singer M
3139 (SCO)
Type information: USSR. Caucasus: KM.
campinaranae Singer, 1989a: 4
Brazil.
transformabilis Singer, 1973d: 5
Caracaraf, 22 May 1978, Singer B 11022
San Javier, 20 December 1958, Singer T 3149
(INPA)
(F)
chilensis Singer, 1969a: 18
HOLOTYPE: Chile. Llanquihue: Puella, 20 veselskyi Singer & Kuthan, 1976g: 9
HOLOTYPE: Czechoslovakia. Moravia: Ostra-
1916 (SGO)
Halde Lucina, 19 July 1974, Kuthan, Sing-
va,
hololeuca Singer, 1989a: 4
er & Veselsky [Singer no.] C 5575 (F)
Manaus, 17 May 1977, Singer B 9639 (INPA)
lactarioides Singer, 1989a: 5 HYGROPHOROPSIS
HOLOTYPE: Bolivia. El Beni: Guayaramerin,
16 March 1956, Singer B 2012 (LIL) aurantiaca var nana Singer, 1946a: 546
lateritiorosella Singer, 1969a: 19
HOLOTYPE: Chile. Colchagua: Highlands Hammock State Park, September
Pumanque,
Lazo PU-54 (SGO) 1942, Singer F 640 (FH)
megistospora Singer, 1989a: 5 panamensis Singer, 1983f: 28

HOLOTYPE: HOLOTYPE: Panama. Chiriqui: near Aguacate,
Aguas Emendadas
Brazil. Goias:
21 January 1978, Singer B 10560 about 7 km NE of El Hato del Volcan, Cerro
Res.,
Punta Road, 4 July 1975, Dumont & Carpen-
(INPA)
ter [Dumont no.] P 2170 (NY)
mexicana Singer, 1958d: 225
HOLOTYPE: Mexico. Oaxaca: Huautla de Jim- tapinia Singer, 1946a: 544
1531 (LIL) Type information: USA. Florida: Dade County,
mutabilis Singer, 1989a: 5 Miami, Simpson Park (FH)
HOLOTYPE: Brazil. Amazonas: Laguna Jan-
uaryauari, 21 December 1978, Singer B 11465 HYGROPHORUS
(INPA)
octayensis Singer, 1969a: 15 quercuum Singer, 1973d: 1
HOLOTYPE: Chile. Osorno: Puerto Octay, 15 HOLOTYPE: Colombia. Boyaca: Bosque de La

September 1967, Singer M
7538 (SGO) Mercedes, 8 April 1968, Singer B 6020 (F)
paraibensis Singer, 1965e: 19

HOLOTYPE: Brazil. Paraiba: Joao Pessoa, 11
HYGROTRAMA Singer, 1958d: 221
July 1960, Singer B 3320 (LIL)
proteus Singer, 1969a: 14 Type species: Hygrotrama dennisianum Singer
Mirador, 30 April 1967, Singer M 6560 (SGO) albipes Singer, 1973d: 4
pseudoamylospora Singer, 1989a: 6 HOLOTYPE: Mexico. Veracruz, 7 km S of
HOLOTYPE: Ecuador. Tungurahua: Rio Topo, Montepio, Estacion Biologica de las Tuxtlas,
30 April 1973, Singer B 7190 (F) 29 July 1969, Singer M 8734 (F)

HYGROTRAMA
araguense Singer, 1989a: 2 dilutecinnamomea Singer, 1954b: 127

HOLOTYPE: Venezuela. Aragua: Parque Na- HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
cional Henry Pittier, 6 July 1971, Dumont, pi National Park, Puerto Manzano, 18 May
Haines & Samuels s.n. (NY) 1952, Singer M 750a (LIL). Cited with anoth-
dennisianum Singer, 1958d: 221 er specimen, type not indicated.
HOLOTYPE: Mexico. Oaxaca: Huautla de Jim- dissocystis Singer, 1953a: 231
nez, 13 July 1957, Singer M 1553 (LIL) HOLOTYPE: Argentina. Tierra del Fuego: Rio
leucopus Singer, 1973d: 3 Grande, Estancia Nueva Argentina, 10 Feb-
HOLOTYPE: Brazil. Paraiba: Joao Pessoa, 11 ruary 1950, Singer M
159 (LIL)
June 1960, Singer B 3344 (BAFC) erythrobasis Singer, 1954b: 127, 128
tetrasporum Singer, 1989a: 3 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
HOLOTYPE: Brazil. Amazonas: Roads Manaus- pf National Park, Puerto Manzano, 18 May
Caracarai, km 45, 28 February 1978, Singer 1952, Singer M 748? (LIL)
& Araujo B 10714 (INPA) eutheles van dollfmii Singer, 1938e: 16
Type information: USSR. (LE?)
HYMENOGASTER fuscata Singer, 1953a: 236
pulcher Singer, 1969a: 372 Grande, Estancia Nueva Argentina, 13 Feb-
HOLOTYPE: Lago Cor-
Argentina. Neuquen:
20 April M5083
249 (LIL)
rentoso, 1965, Singer
fuscocinnamomea Singer, 1953a: 233
(BAFC)
HYPSIZYGUS Singer, 1947b: 77
M
306 (LIL)
ruary 1950, Singer
Type species: Pleurotus tessulatus (Bull.: Fr.) geophyllomorpha Singer, 1953a: 229
Gill. HOLOTYPE: Argentina. Tierra del Fuego: Rio
INOCYBE ruary 1950, Singer M
272 (LIL)
gigacystis Singer, 1953a: 231
amazoniensis Singer & Araujo, 1983f: 182 HOLOTYPE: Argentina. Tierra del Fuego: Rio
HOLOTYPE: Brazil. Amazonas: Road Manaus- Grande, Estancia Nueva Argentina, 10 Feb-
Caracaraf, km 125, 21 May 1978, Singer & M
167 (LIL)
ruary 1950, Singer
Araujo [Singer no.] B 11019 (INPA)
homomorpha Singer, 1931: 536
argentea Singer, 1931: 535 Type information: USSR. Caucasus: Kha.
Type information: USSR. Caucasus: Nakratal.
langei van maior Singer, 1936i: 432
bridgesiana Singer, 1953a: 227 Type information: Spain. Catalonia.
HOLOTYPE: Argentina. Tierra del Fuego: Rio lutescens Singer, 1953a: 223
201 (LIL)
brunnea f.
insignior Singer, 1947c: 245 ruary 1950, Singer M
160 (LIL)
Type information: Spain. Catalonia. Salardu, matrisdei Singer, 196 la: 112, 113
September, 1934. HOLOTYPE: Bolivia. Pando: Manuripi, 27
carelica Singer, 1938e: 16 March 1956, Singer B 2285 (LIL)
Type information: USSR. 26 August 1936,
mixtiliformis Singer, 19545: 129
Freindling & Singer (LE?) HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
catalaunica Singer, 1947c: 245
Type information: Spain. Catalonia, Base de Singer M523 (LIL)
Baricauba, September, 1934 (LE).
neotropicalis Singer, 1983f: 186
caucasica Singer, 1937: 236 HOLOTYPE: Costa Rica. Guanacaste: Parque
Type information: USSR. Western Caucasus: Ya- Nacional Santa Rosa, 25 July 1981, Singer B
tyrgarta, between Carices and Hypna, 30 July 12441 (F)
1936, Vasilieva (LE?)
neuquenensis Singer, 1954b: 130
cerasphora Singer, 1953a: 228 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
HOLOTYPE: Argentina. Tierra del Fuego: Rio pi National Park, Quetrihue, 20 May 1952,
Grande, Estancia Nueva Argentina, 15 Feb- Singer M 781 (LIL)
290 (LIL)
notodryina Singer, 1983f: 175
chilensis Singer, 1965i: 175 HOLOTYPE: Costa Rica. Guanacaste: Parque
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, Nacional Santa Rosa, 25 July 1981, Gomez &
28 March 1963, Singer M 3196 (BAFC) Singer [Singer no.] B 12446 (F)
46 Chapter Three
INOCYBE
obscura var. subpusio Singer, 1947c: 232, 233 rotundifoliae Singer, 1938g: 18
Type information: Spain. Catalonia. Riu de Bon- No type information.
aigua, July 1934.
paucigibba Singer, 1965i: 176 KUEHNEROMYCES Singer & Smith, 1946d:

504
3200 (BAFC)
plocamophora Singer, 1983f: 177
Type species: Pholiota mutabilis Schaeff. : Fr.
castaneiceps Singer, 1969a: 281
Singer [Singer no.] B 12451 (F) HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
pseudohiulca ssp. subduriuscula Singer, 1947c: 1967, Singer M 6715 (SGO)
232 cystidiosus Singer, 1953a: 221
Type information: Spain. Catalonia: Pyrenees HOLOTYPE: Argentina. Tierra del Fuego: Rio
Mtns., Salardu, August, 1934. Grande, Estancia Nueva Argentina, 14 Feb-
retipes Singer, 1953a: 225 ruary 1950, Singer M 276 (LIL)
HOLOTYPE: Argentina. Tierra del Fuego: Lago depauperatus Singer & Smith, 1946d: 513
Fagnano, Cabercera del Lago, 19 February HOLOTYPE: USA. Alachua County,
Florida:
1950, Singer M 380 (LIL) Devil's Millhopper, 25 July 1943, Singer F
squarrosoamthystina Singer, 1938e: 17 2992 (FH)
Type information: USSR. Leningrad, December macrosporus Singer, 1953a: 220
1935, Singer (LE?) HOLOTYPE: Argentina. Tierra del Fuego: Rio
subargentea Singer, 1938e: 16 Grande, Estancia Nueva Argentina, 16 Feb-
Type information: USSR. Leningrad, September ruary 1950, Singer M 310 (LIL)
1936, Singer (LE?) nudus Singer, 1950c: 194
subfibrosoides Singer, 1953a: 234 HOLOTYPE: Argentina. Tucuman: Sierra de
HOLOTYPE: Argentina. Tierra del Fuego: Rio San Javier, Park Aconquija, 12 March 1949,
Grande, Estancia Nueva Argentina, 15 Feb- T 207
Singer (LIL)
ruary 1950, Singer M 291 (LIL)
rostratus Singer & Smith, 1946d: 510
subfuscocinnamomea Singer, 1954b: 131 HOLOTYPE: USA. Michigan: Milford, Smith
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- 15002 (MICH)
vernalis f. amara Singer & Smith, 1946d: 520
Singer M 663 (LIL) HOLOTYPE: USA. Wyoming: Little Brooklyn
subgeophyllomorpha Singer, 1954b: 132
Lake, Arenberg 68 (FH?)
pi National Park, Quetrihue, 19 May 1952,
Singer M 764 (LIL) LACCARIA
subgigacystis Singer, 1954b: 133
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- altaica Singer ex Singer, 1967d: 122
pi National Park, Quetrihue, 16 May 1952, HOLOTYPE: USSR. Altai Mtns., 31 July 1936,
Singer M 676 (LIL) Singer A 439 (LE)
tequendamae Singer, 1963b: 244 bullulifera Singer, 1965i: 149
HOLOTYPE: Colombia. Cundinamarca: Salto HOLOTYPE: Mexico. Puebla: Popocatepetl: Pi-
de Tequendama, 23 July 1960, Singer B 3500 neto, W
of Paso de Cort, 21 July 1957, Singer
(BAFC) M 1581 (MICH). Also see 1957a: 355.
calospora Singer, 1973d: 7
IXOCOMUS HOLOTYPE: USA. Massachusetts: Hampden
jacuticus Singer, 1938c: 40 County, Southwick, 21 June 1945, Waban
Type information: USSR. Siberia: Lake Char- (FH)
czarbyt Yacotia, Benois, August 1925, 26 galerinoides Singer, 1965i: 147
(LE?) HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
sibiricus Singer, 1938c: 46 28 March 1963, Singer M 3212 (BAFC)
Type information: USSR. Altai Mtns.: Lake Te- gomezii Mueller & Singer, 1988g: 224
lezkoje, July-September 1937, Singer & Vasi- HOLOTYPE: Costa Rica. San Jose: La Chonta,
along Interamerican Hwy, km 55, September
lieva (LE?)
1982, Gomez 18443 (F 1051926)

KROMBHOLZIA laccata var. qffinis Singer, 1967d: 110
oxydabilis Singer, 1938g: 18 HOLOTYPE: England. Kent: Bedgebury, 30
No type information. October 1960, Singer C 3118 (BAFC)

LACCARIA
laccata var. anglica Singer, 1967d: 111 tetraspora var. scotica Singer, 1967d: 1 14
HOLOTYPE: England. Kent: Bedgebury, 20 HOLOTYPE: Scotland. Lake Katrin, 1
August
October 1960, Singer C 3119 (BAFC) 1964, Singer C 4002 (BAFC)
laccata var. carbonicola Singer, 1967d: 110 tetraspora var. valdiviensis Singer, 1967d: 113
HOLOTYPE: USA. Massachusetts: Blue Hills, HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
11 November (FH)
1943, Singer s.n. 28 March 1963, Singer M 3191 (BAFC)
laccata var. chilensis Singer, 1967d: 109 tetraspora var. xena Singer, 1967d: 116
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, Type was not indicated.
laccata var. gibba Singer, 1969a: 27
LACHNELLA
2 km S of Mirador, 6 May 1967, Singer M
australis Singer, 1966a: 12
6738 (SGO)
HOLOTYPE: Argentina. Rio Negro, Nahuel
laccata var. intermedia Singer, 1977d: 368
Huapf National Park, between Laguna Frias
HOLOTYPE: Czechoslovakia. Moravia: Jeseni-
and Paso de las Nubes., 24 March 1963, Sing-
ky, Reiviz near Jesenik, 24 July 1974, &
&
M. Singer [Singer no.]
J.
C 5664
J.
er & Moser [Singer no.] M
3185 (BAFC)
Kuthan, R.
cecropiae Singer, 1966a: 13
(F)
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ivon,
laccata var. moelleri Singer, 1973d: 9
3 April 1956, Singer B 24901 (LIL, BAFC)
HOLOTYPE: Czechoslovakia. Bohemia: Chlum
near Trebon, 30 September 1970, Singer C
5227 (F) LACHNOCLADIUM
laccata var. subalpina Singer, 1977d: 365
HOLOTYPE: Czechoslovakia. Slovakia: Hone hericiiforme Singer, 1938e: 4
Tatra, Solisko, 4 September 1974, Singer C HOLOTYPE: USSR. Vassilkov (LE?)
5870 (F)
laccata var. tatrensis Singer, 1977d: 367
LACTARIUS
HOLOTYPE: Czechoslovakia. Slovakia: Hohe
Tatra, Mala studena
dolina, 13 September
allochrous Singer, 1948c: 41
1974, Singer 6001 (F)C
laccata var. vulcanica Singer ex Veselsky & theson Hammock, 16 September 1942, Singer
Singer, 1977d: 362 F 714 (FH)
alni Singer, 1961b: 83
va, Halde Lucina, 30 August 1968, Veselsky
s.n. (F)
montana Singer, 1973d: 8
(LIL)
HOLOTYPE: Switzerland. Borgne de
Valais:
amazonensis Singer, 1983f: 295
Ferpecle, 11 July 1971, Singer 5442 (F)
Brazil.
proximella Singer, 1965i: 146
Caracarai, km20 July 1977, Singer &
125,
March 1963, Singer M 3274 (BAFC) Araujo [Singer no.] B 11315 (INPA)
annulifer Singer, 1983f: 290
tetraspora Singer, 1946e: 689
HOLOTYPE: USA. Florida: Highlands HOLOTYPE: Brazil. Amazonas: Road Manaus-
County,
Caracarai, km 45, 31 January 1980, Webber,
Highlands Hammock State Park, 11 August
1942, Singer F 160 (FH) Bradley & Singer [Singer no.] B 12087
tetraspora aberrans Singer, 1967d: 116
var. (INPA)
HOLOTYPE: Argentina. Neuquen, Nahuel Hua- brasiliensis Singer, 1983f: 308
pi National Park, 14 March 1959, Singer M HOLOTYPE: Brazil. Amazonas: Road Manaus-
1774 (BAFC) Caracarai, km 45, 28 February 1978, St. John,
tetraspora f. major Singer, 1967d: 113 Araujo & Singer [Singer no.] B 10729 (INPA,
Type specimen was not indicated. F)
tetraspora var. peladae Singer, 1967d: 117 campinensis Singer, 1984a: 438
El Mirador, 5 May 1965, Singer M 5515 Caracarai, 25 April 1980, Singer B 12124
(BAFC) (DSTPA)
tetraspora var. peullensis Singer, 1967d: 113 caucae Singer, 1973d: 105
HOLOTYPE: Chile. Llanquihue\ 22 March HOLOTYPE: Colombia. Cauca: Los Araoz, 12
1959, Singer M
1991 (BAFC) April 1968, Singer B 6059 (F)
48 Chapter Three
LACTARIUS
costaricensis Singer, 1983f: 321 veraecrucis Singer, 1973d: 104

HOLOTYPE: Costa Rica. Guanacaste: Parque HOLOTYPE: Mexico. Veracruz: near Montepio,
Nacional Santa Rosa, 25 July 1981, Gomez & 19 June 1969, Singer 8025 (F) M
Singer [Singer no.] B 12437 (F) xanthydrorheus Singer, 1945d: 426
gigasporus Singer, 1983f: 323 HOLOTYPE: USA. Florida: Highlands County,
HOLOTYPE: Brazil. Amazonas: Punta Negra, Highlands Hammock State Park, 10 August
24 December 1978, Singer B 11444 (INPA) 1942, Singer F 134 (FH)
guanacastensis Singer, 1983f: 325

HOLOTYPE: Costa Rica. Guanacaste: Bagaces, LACTOCOLLYBIA Singer in Singer &
7 km NW of town, October 1982, Gomez Digilio, 195 le: 177. [Published in 1952].
18686 (F)
Type species: Colly bia lacrimosa Heim. Also
igapoensis Singer, 1984a: 441 see Schweiz. Zeitschr. f. Pilzk 17: 16, 1939
HOLOTYPE: Brazil. Amazonas: Igarap do Tar-
umazinho, 16 April 1980, Singer B 12099 aequatorialis Singer, 1978h: 11
(INPA) HOLOTYPE: Ecuador. Napo: Shushufmdi, 13
laricinus Singer, 1989a: 132 May 1973, Singer B 7402 (F)
HOLOTYPE: USA. Illinois: McHenry County, angiospermarum Singer, 1948c: 32
26 August 1984, Singer N 5152 (F) HOLOTYPE: USA. Florida: Dade County, Ma-
mamorensis Singer, 1983f: 302 theson Hammock, 1 November 1942, Singer
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 3 km (FH)
below Guayaramerin along the left bank of the aurantiaca Singer in Singer & Digilio, 195 le:
Mamore river, 12 March 1956, Singer B 1882 180
(F, BAFC)
neotropicus Singer, 1952b: 299
HOLOTYPE: Trinidad. Between Maqueripe trail (LIL). [Published in 1952].
and N Post, 30 September 1949, Dennis 97 graminicola Singer, 1989a: 19

HOLOTYPE: Venezuela. Dto Federal: near Rio
(LIL)
Caracas, 22 July 1972, Dumont and others VE
pallidipes Singer, 1983f: 299 5632 (NY)
Caracarai, km 125, 20 July 1978, Singer B
microspora Singer, 1961b: 51
HOLOTYPE: Argentina. Misiones: Iguazii,
11328 (INPA)
Puerto Iguazu, 12 April 1957, Singer 950 M
panuoides Singer, 1952b: 300 (LIL)
HOLOTYPE: Trinidad. Naranja, 2 October
modesta Singer, 1970d: 785
1949, Dennis 109 (LIL) HOLOTYPE: Chile. Colchagua: Pumanque,
paulensis Singer, 1983f: 305 Lazo PU-141 (SGO)
HOLOTYPE: Brazil. Sao Paulo: Serra da Can-
lowest part of Reserva Florestal, 17
tareira,
LAMPTEROMYCES Singer, 1947b: 79
December 1964, Singer B 4093 (F)
purgatorii Singer, 1948c: 42 Type species: Pleurotus japonicus Kawamura

Type information: USA. Massachusetts: Nor-
wood, July 1941 (FH) LECCINUM
quercuum Singer, 1963b: 249 albellum f. epiphaeum Singer, 1947a: 120
HOLOTYPE: Colombia. Boyaca: Tunja, Arca-
buco, Finca del Sr. Luis Antonio Porras, 29 Gainesville.
ty,
July 1960, Singer B 3611 (BAFC)
chalybaeum Singer, 1945i: 799
subpallidipes Singer, 1983f: 298 Type information: USA. Florida: Alachua Coun-
HOLOTYPE: Brazil. Amazonas: Road Manaus- ty, Gainesville.
Caracaraf, km 125, 21 May 1978, Singer B roseoscabrum Singer & Williams, 1992c: 726
11048 (F) HOLOTYPE: USA. Florida: Sarasota Co.,
subreticulatus Singer, 1983f: 314 Myakka River State Park, 13 January 1991,
HOLOTYPE: Brazil. Amazonas: Road Manaus- Williams & Singer, (Williams No.) 394 (=
Caracaraf, km 45, 5 May 1980, Singer B Singer No. F 3908) (F)
12145 (INPA) subglabripes var. corrugatoides Singer, 1945i:
uyedae Singer, 1984a: 436 799
HOLOTYPE: Japan. Shiga: Otsu, Kokubu, 3 Type information: USA. Florida: Alachua Coun-
August- 10 September 1973, Uyeda (F) ty, Gainesville.

LENTINELLUS
LENTINELLUS LEPISTA
collybiiformis Singer, 1989a: 10
calyciformis Singer, 1969a: 379 HOLOTYPE: Brazil. Amazonas: Road Manaus-
HOLOTYPE: Argentina. Rio Negro: Laguna km 45, 3 August Freire
Caracarai, 1978,
Frias, to Paso de las Nubes, 24 March 1963,
Singer M 3187 (BAFC) diemii Singer, 1954b: 103, 104
LENTINUS pi National Park, Quetrihue, May 1952, Sing-
er M 772/M 732? (LIL)
panziensis Singer, 1973d: 1 fibrosissima Singer, 1954b: 104
HOLOTYPE: Zaire. Panzi Kivu, March 1956, HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
Goossens-Fontana 5559 (BR) pf National Park, Quetrihue, May 1952, Sing-
er M 581/M 771? (LIL)
prancei Singer, 198 Ic: 152
HOLOTYPE: Brazil. Amazonas: Ilha Marchan- geotropoides Singer in Singer & Digilio, 195 le:
daria, 1 July 1981, Singer B 12205 (F)

44
& HOLOTYPE: Argentina. Tucuman: Cerro Alto
sayanus Singer in Singer Digilio, 1951e: 139
del Taficillo, 4March 1951, Singer T 1363
HOLOTYPE: Argentina. Tucuman: Rio de las
(LIL) [published in 1952]
Pavas, October, Say [Singer no.] T 633 (LIL)
nuda var. tucumanensis Singer in Singer & Di-
[published in 1952]
gilio, 1951e: 43
LEPIOTA San Javier, Anta Muerta, 16 April 1950, Sing-
er T 960 (LIL) [published in 1952]
amyloidea Singer, 1943a: 171 rickenii Singer, 1948c: 26
Type information: USSR. Altai Mtns., Oirotia: HOLOTYPE: USSR. Rajajoki, 26 September
Lake Telezkoje, September. 1940, Singer 5-823 (LE)
azurea Singer in Singer &
Digilio, 195 le: 278
HOLOTYPE: Argentina. Salta: Rio Piedras, 25 LEPTOGLOSSUM
March 1951, Singer T 1426 (LIL) [published
alnetorum Singer, 1970a: 75
in 1952]
brunneosperma Singer in Singer & Digilio, Yala, 13 February 1966, Singer T 5084 (F)
195 le: 279
subbryophilum Singer, 1931: 521
HOLOTYPE: Argentina. Tucuman: Garden of Type information: USSR. Caucasus: Kha.
the Instituto Lillo, 2 January 1949, Singer &
Digilio [Singer no.] T 76 (LIL) [published in
LEPTONIA
1952]
catalaunica Singer, 1936i: 428
gomezii Singer, 1989a: 99
HOLOTYPE: Costa Rica. Guanacaste: Bagaces, Type information: Spain. Catalonia: Central Pyr-
7 km NW of town, October 1982, Gomez enees, September 1934.
18713 (F)
izonetae Singer, 1989a: 100
LEUCOAGARICUS (Locquin) Singer, 1948c:
Brazil.
35
Manaus, 26 December 1977, Aguiar & Singer Type species: Leucocoprinus macrorhizus Loc-
[Singer no.] B 10409 (INPA) quin
neuquenensis Singer, 1969a: 171
acaciarum Singer, 1973d: 67
HOLOTYPE: Argentina. Corrientes: Curuzii
zano, 22 April 1964, Singer M
3425 (BAFC)
Cuatia, 7 November 1965, M. Singer [Singer
patagonica Singer, 1969a: 172 no.] S 564 (BAFC)
atrofibrillosus Singer, 1969a: 162
Frias, 2 May 1964, Singer M 3847 (BAFC)
subcortinaria Singer, 1969a: 171
zano, Rfo Bonito, 6 April 1963, Singer M
HOLOTYPE: Argentina. Rio Negro: near Santa 3291 (BAFC)
Maria, 19 April 1965, Singer M 5055 (BAFC) exannulatus Singer, 1973d: 68
xanthophylla Singer, 1973d: 67 HOLOTYPE: Argentina. Tucuman: Parque,
HOLOTYPE: Brazil. Para: Belem, 10 June 1966, Aconquija, 12 March 1949, Singer T 211
Singer B 4249 (BAFC) (LIL)
50 Chapter Three
LEUCOAGARICUS
lilaceus Singer in Singer &

Digilio, 195 le: 274 oaxacana Singer, 1957a: 359
HOLOTYPE: Argentina. Tucuman: Garden of Type information: Mexico. Oaxaca, 10-12 July
the Institute Lillo, 9 April 1949, Antonuccio 1957 (MICH). Two specimens were cited: M
& Golbach [Singer no.] T 306 (LIL) [pub- 1504 and M 1522, but no type was designated.
lished in 1952]
tricolor Singer, 1989a: 98

LINDEROMYCES Singer, 1947d: 157
Brazil.

Type species: Paxillus lateritius Fetch
LEUCOCOPRINUS corneri Singer, 1961c: 14

minutulus Singer, 1941b: 86
nabara, Corcovado, 4 December 1948, Corner
Type information: USSR. Krasnodar Krai: Ac- 329 (R)
zipsta, August 1936, Vasilieva (LE?)
LEUCOINOCYBE LULESIA Singer, 1970a: 16

Singer, 1943a: 144
Type species: Mycena lenta Maire Type species: Armillariella densifolia Singer in
Singer & Digilio
LEUCOPAXILLUS
albissimus var. kauffmanii Singer & Smith,
LYOPHYLLUM
1943b: 106
HOLOTYPE: USA. Ann Arbor, 14
langei Singer, 1969a: 24
Michigan: HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
November 1926, Kauffman (MICH)
pi National Park, above Puerto Manzano, 1 1
albissimus var. lentus f. olympianus Singer & April 1963, Singer M
3384 (BAFC)
Smith, 194?b: 109
nodulosporum Singer, 1969a: 26
HOLOTYPE: USA. Washington: Olympic Na- HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
Lake Mills, 26 September 1941,
tional Forest,
1967, Singer M 6708 (SGO)
Smith 17257 or 17333 (MICH?)
semiustum Singer in Singer & Digilio, 195 le:
albissimus var. lentus f.fucatus Singer & Smith, 17
1943b: 110 HOLOTYPE: Argentina. Tucuman: near Tapia,
HOLOTYPE: USA. Washington: Mt. Baker,
17 February 1951, Singer T 1230 (LIL) [pub-
Boulder Creek Trail, Mains & Smith [Smith
lished in 1952]
no.] 16593 (MICH) tucumanense Singer in Singer & Digilio, 195 le:
albissimus var. monticola Singer & Smith,
17
1947f: 730 HOLOTYPE: Argentina. Tucuman: Rio de los
HOLOTYPE: USA. Idaho: Payette Lakes, Au-
Sosas, 27 January 1951, Singer T 1146 (LIL)
gust 1943, Gruber P-27 (MICH). [Published
[published in 1952]
in 1948].
gracillimus Singer &

Smith, 1943b: 131
HOLOTYPE: Rio Grande do Sul (FH)
Brazil.
MACOWANITES
patagonicus Singer, 1954b: 111, 112 &
acris Signer Smith, 1960h: 86
HOLOTYPE: USA. Idaho: near Stanley, 20 July
pi National Park, Isla Victoria, 22 May 1952,
58967 (MICH)
1958, Smith
Singer (LIL)
sainii Singer, 1989a: 19 albidigleba Singer & Smith, 1960h: 85
HOLOTYPE: USA. Idaho: near McCall, 30 July
HOLOTYPE: India. Himachal Pradesh: Narkan-
59225 (MICH)
1958, Smith
da, 9 August 1979, Atri PUN 328 (F)
americanus Singer & Smith, 1960h: 88
septentrionalis Singer & Smith, 1947f: 726
HOLOTYPE: USA. Idaho: Payette National For-
HOLOTYPE: USA. Oregon: Mt. Hood National
est, 16 July 1954, Smith & Bigelow [Smith
Forest, Bear Springs, Gruber [Smith no.]
no.] 45032 (MICH)
24982 (MICH). [Published in 1948].
citrinus Singer & Smith, 1960h: 82
HOLOTYPE: USA. Idaho: near Stanley, 19 July
LIMACELLA 58942 (MICH)
1958, Smith
grisea Singer, 1989a: 92 fulvescens Singer &

Smith, 1960h: 84
HOLOTYPE: Spain. Catalonia: Prat de Llobre- HOLOTYPE: USA. Idaho: near McCall, 5 Au-
gat, 23 October 1979, Singer C 9595 (F) gust 1958, Smith 59584 (MICH)

MACOWANITES
fuscoviolaceus Singer & Smith, 1960h: 87 alneus Singer, 1973a: 44, 322
HOLOTYPE: USA. Idaho: Payette National For- HOLOTYPE: Mexico. Oaxaca: San Andres Hi-
est, 30 July 1958, Westerdale [Smith no.] dalgo, 9 July 1969, Singer 8378 (F) M
59261 (MICH) alvaradoi Singer, 1973a: 277, 322
mollis Singer & Smith, 1960h: 79 HOLOTYPE: Argentina. Jujuy: 15 km NE of
HOLOTYPE: USA. Washington: Mt. Rainier Posta de Lozano, 20 February 1966, Singer T
National Park, 23 September 1948, Sturtz 5311 (F)
[Smith no.] 31683 (MICH) alvaradoi \ar.filicinus Singer, 1975c: 887
setchellianus Singer & Smith, 1960h: 92 HOLOTYPE: Ecuador. Napo: Shushufindi, 13
HOLOTYPE: USA. California: Contra Costa
County, no date, Parks 1440 (NY) amazoniensis Singer, 1989a: 26
HOLOTYPE: Brazil. Amazonas, 30 km N of
MACROCYSTIDIA Manaus, 18 May 1977, Singer B 9643 (INPA).
The type information of Cystolepiota amazon-
africana Singer, 1973d: 15 ica Singer (1989a: identical to this.
99) is
HOLOTYPE: Zaire, near Panzi Kivu, February
Based on Singer's unpublished notebook,
1956, Goossens-Fontana 5548 (BR)
however, the type of this species might be
incarnata Singer, 1973d: 16
Singer B 9640 from the same location. See
HOLOTYPE: Zaire, near Binga, 30 May 1928,
note under C. amazonica Singer.
Goossens-Fontana (BR)s.n.
& ambiguus Singer, 1965e: 25

indica Saini, Atri Singer, 1982c: 236
HOLOTYPE: Brazil. Amazonas: 30 km from
HOLOTYPE: India. Himachal Pradesh: Simla,
Summer Hill, 21 August 1979, Atri PUN 275 Manaus, 19 July 1960, Singer B 3442 (BAFC)
(PUN). Isotype: F amphicystis Singer, 1975c: 880

occidentalis Singer in Singer & HOLOTYPE: Colombia. Valle: Cali, Buenaven-
Digilio, 1951e:
174 tura, Calima, 18 April 1968, Singer B 6289
HOLOTYPE: (F)
Argentina. Tucuman: Marcos Paz,
26 March 1949, Singer T 281 (LIL) [pub- androsaceiformis Singer, 1973a: 127, 322
lished in 1952] HOLOTYPE: Mexico. Oaxaca: San Andres Hi-
dalgo, Sierra Mazateca, 9 July 1969, Singer M
8387
MACROLEPIOTA Singer, 1946g: 141.
(F)
[Published in 1948]. angustispermus Singer, 1973a: 54, 323

Type species: Lepiota procera Scop. : Fr. Montepio, Estacion Biologica de las Tuxtlas,

8046 (F)
MACROMETRULA Donk & Singer, 1948a:
antarcticus Singer, 1969a: 73
264 HOLOTYPE: Chile. Osorno: Pilmayquen, 13
May 1967, Singer M 6860 (SGO)
Type species: Agaricus rubriceps Cooke & aporposeptus Singer, 1973a: 201, 323
Mass. HOLOTYPE: Brazil. Para: Belem, 10 June 1966,
Singer B 4261 (F)
MANURIPIA Singer, 1960f: 273, 279 aquilus Singer, 1973a: 148, 323
HOLOTYPE: Bolivia. El Beni: Vaca Diez,
Type species: Manuripia bifida Singer Guayaramerin, 7 March 1956, Singer B 1635
(F)
bifida Singer, 1960f: 273
HOLOTYPE: Bolivia. Pando: aurantiorufescens Singer, 1973a: 66, 323
Manuripi, Con-
26 March 1956, Singer B 2272 (LIL) HOLOTYPE: Mexico. Chiapas: San Cristobal de
quista,
las Casas, 24 km SW, 3 August 1969, Singer
MARASMIELLUS M 8929 (F)

baeosporoides Singer, 1989a: 26
albifolius Singer, 1973a: 155, 322 HOLOTYPE: Brazil. Amazonas: Igarape do Tar-
HOLOTYPE: Pernambuco: Camaragibe,
Brazil. umazinho, 12 March 1980, Singer B 12025
6 July 1960, Singer B 3132 (F) (INPA)
albobrunnescem Singer, 1973a: 121, 322 baeosporus Singer, 1973a: 69, 323
HOLOTYPE: Mexico. Veracruz: 7 km S of HOLOTYPE: Colombia. Valle: Cali, Buenaven-
Montepio, Estacion Biol6gica de las Tuxtlas, tura, Calima, 25 April 1968, Singer B 6366
8087 (F) (F)
52 Chapter Three
MARASMIELLUS
bauhiniae Singer, 1989a: 27 couleu Singer, 1954b: 118, 119

HOLOTYPE: Brazil. Amazonas: Manaus, 24 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
April 1977, Singer B 9801 (INPA) pf National Park, Puerto Manzano, 18 May
berkeleyi Singer, 1973a: 98, 324 1952, Singer M

747 (LIL)
Type information: Cuba. Wright 150 (FH). No crinipelloides Singer, 1973a: 76
site data or date given. HOLOTYPE: Mexico. Veracruz: 3 km S of
bisporiger Singer, 1973a: 189, 324 Montepfo, 22 June 1969, Singer M 8153 (F).
HOLOTYPE: Colombia. Valle: Cali, Buenaven- No Latin diagnosis.
tura, Calima, Campamento, 24 April 1968, cystidiosus Singer, 1973a: 177, 326
Singer B 6353 (F) HOLOTYPE: Argentina. Buenos Aires: Santa
bolivarianus Singer, 1973a: 257, 324 Catalina, 19 April 1963, Singer S 404 (F)
HOLOTYPE: Venezuela. Bolivar: Chimanta daguae Singer, 1973a: 146, 326
Massif, Toronto-tepui, 23 February 1955, HOLOTYPE: Colombia. Valle: Corr. El Vergel,
Steyermark & Wurdack 117 (NY) Municip. Dagua, Finca La Araucana, 27 April
brunneomarginatus Singer, 1973a: 180, 324 1968, Singer B 6428 (F)
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ar- defibulatus Singer, 1955a: 772
royo Florida, 22 km E of Riberalta, 7 April HOLOTYPE: 28 Sep-
Brazil. Parana: Paranagua,
1956, Singer B 255 (LIL, BAFC) tember 1952, Singer B

401 (F)
californicus Singer, 1989a: 28 defibulatus var. insignis Singer, 1973a: 195, 326
HOLOTYPE: USA. California: San Mateo HOLOTYPE: Brazil. Pernambuco: Mata de Sao
County, 16 July 1975, Singer N 7531 (F) Joao 16 July 1960, Singer B 3420 (F)
,
caracassensis Dennis ex Singer, 1973a: 100, defibulatus var. strictior Singer, 1973a: 192, 326
324 HOLOTYPE: Colombia. Valle: Cali, Buenaven-
HOLOTYPE: Venezuela. Dto. Federal: Caracas San Joaqufn, 22 April 1968, Singer B
tura,
Botanic Garden, 3 July 1958, Dennis 1134 (K) 6281 (F)

cinereus Singer, 1973a: 291, 325 dendroegrus Singer, 1973a: 59, 326
HOLOTYPE: Mexico. Veracruz: 3 km S of HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri-
Montepfo, 22 June 1969, Singer 8143 (F) M beralta, 31 March 1956, Singer B 2384 (LIL)
clitocybe Singer, 1955c: 148 devenulatus Singer, 1961b: 57

HOLOTYPE: Argentina. Tucuman: Quebrada, HOLOTYPE: Bolivia. El Beni: Vaca Diez, 1
25 January 1955, Singer T 2082 (LIL). The April 1956, Singer B 2417 (LIL)
type information for this species is identical distoriformis Singer in Singer & Digilio, 195 le:
to that of Neoclitocybe Singer latispora 155
(1973d: 22), except for the herbarium of de- HOLOTYPE: Argentina. Tucuman: Rfo de los
posit. Sosas, 13 December 1949, Singer T 761 (LIL)
cnacopolius Singer, 1973a: 266, 325 [published in 1952]
HOLOTYPE: Colombia. Cauca: Los Araoz, dryogeton Singer, 1973a: 64, 326
road to Cali-Popayan, 12 April 1968, Singer HOLOTYPE: Mexico. Oaxaca: Sierra Mazateca,
B 6041 (F) San Augustin, 10 July 1969, Singer 8439A M
cocosensis Singer, 1989a: 27 (F)
HOLOTYPE: Costa Rica. Cocos Island, trail be- echinocephalus Singer, 1955a: 773
tween Chatham and Wafer Bay, December HOLOTYPE: Panama. Zona Canalis: Barro Col-
1981, Gomez 17974 (F) orado Island, Martin 4082 (F, IA)
columbianus Singer, 1973a: 204, 325 edite Singer, 1969a: 75
HOLOTYPE: Colombia. Cauca: El Guayabo, HOLOTYPE: Chile. Llanquihue, 20 March
road to Cali and Popayan, 29 April 1968, Id- 1959, Singer M 1882 (BAFC)
robo [Singer no.] B 6797 (F) elongatisporus Singer, 1973a: 85, 326
confertifolius Singer, 1973a: 114, 325 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Cer-
HOLOTYPE: Peru. Loreto: Nanay, 1 March ro Uchumachi, 25 February 1956, Singer B
1968, Vigo s. n. (F) 1452 (F)

corticum Singer, 1973a: 297, 325 enodis Singer, 1973a: 61, 327
HOLOTYPE: USA. Florida: Highlands County, HOLOTYPE: Brazil. Pernambuco: Dois Irmaos,
20 August 1942, Singer F 325 (F) 14 April 1964, Singer B 4074 (F)
corynophloeus Singer, 1973a: 52, 325 epitrichialis Singer, 1989a: 28

HOLOTYPE: Mexico. Veracruz: 7 km S of HOLOTYPE: Brazil. Amazonas, Road Manaus-
Montepfo, Estacion Biologica de las Tuxtlas, Caracaraf, km 45, 28 February 1978, Singer
29 July 1969, Singer 8745 (F) M B 1073 la (INPA)

MARASMIELLUS
eugeniae Singer, 1989a: 29 illinorum Singer, 1989a: 31

HOLOTYPE: Brazil. Amazonas, 30 km N of HOLOTYPE: USA. Illinois: Lake County, High-
Manaus, 23 June 1977, Singer10010 (INPA) B land Park, 6 August 1972, Singer N 4152 (F)
foliicola Singer in Singer &
Digilio, 1951e: 148 incarnatipallens Singer, 1973a: 137, 328
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Colombia. Valle: Corn El Vergel,
San Javier, 24 April 1951, Singer T 1523
Municip. Dagua, Finca La Aracuana, 27 April
(LIL) [published in 1952]. This taxon was B 6431
1968, Singer (F)
renamed as Mycena neotropicalis Singer in incrustatus Singer in Singer & Digilio, 1951e:
Singer, 1989a: 82.
153
gigantosporus Singer, 1975c: 881 HOLOTYPE: Brazil. Parecf: Estado Rio Grande
HOLOTYPE: Lagunas de
Argentina. Jujuy:
do Sul, 2 November 1951, Singer B 60 (LIL)
[published in 1952]
gilvus var. bambusearum Singer, 1973a: 287,
inodermatoides Singer, 1973a: 241, 328
327
atas de San Juan, 28 January 1956, Singer B Guayaramerin, 9 March 1956, Singer B 1720
643 (LIL) (F)
1973a: 288, juniperinus var. gilvoides Singer, 1975c: 890

gilvus var. chusquearum Singer,
327 HOLOTYPE: Ecuador. Tungurahua: Rio Topo,
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- 30 April 1973, Singer B 7213 (F)
tapata, 28 January 1956, Singer B 655 (LIL) juniperinus var. radicicola Singer, 1973a: 272,
gilvus var. cundinamarcae Singer, 1975c: 888 328

HOLOTYPE: Cundinamarca: near
Colombia. HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Cer-
Nazareth, 24 July 1974, Dumont & Haines ro Uchumachi, 25 February 1956, Singer B
[Dumont no.] CO 1917 (F). Isotype: NY 1445 (F)
gilvus var. minusculus Singer, 1973a: 289, 327 latispermus Singer, 1989a: 31
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, HOLOTYPE: Peru. Cuzco, Machu Picchu, 20
Charobamba, 30 January 1956, Singer B 745a B 3026 (BAFC)
May 1958, Singer
(LIL)
laurifoliae Singer, 1973a: 124, 328
gomezpompae Singer, 1973a: 161, 327 HOLOTYPE: USA. Florida: Highlands County,
6 August 1942, Singer F 94 (F)
Montepio, 22 June 1969, Singer M 8125 (F)
griseolobrunneolus Singer in Singer & lecythidacearum Singer, 1989a: 31
Digilio,
1951e: 165
HOLOTYPE: Argentina. Tucuman: Sierra de Manaus, 10 October 1977, Singer B 10197

San Javier, Anta Muerte, 16 April 1950, Sing- (INPA)
er T 965 (LIL) [published in 1952] leucophyllus Singer, 1989a: 32
guzmanii Singer, 1973a: 218, 327 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Mexico. Morelos, NW
of Tepoz- Manaus, 17 May 1977, Singer B 9645 (INPA)
tlan, 27 June 1969, Singer M
8210 (F) lysochlorus Singer, 1989a: 33
helminthocystis Singer, 1989a: 29 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Amazonas: Ponta Negra, 8
Brazil. Manaus, 21 April 1977, Singer B 9772 (INPA)
April 1978, Singer B 10963 (INPA)
mesosporus Singer, 1973c: 469
hypolissus Singer, 1989a: 30 HOLOTYPE: USA. North Carolina: Hatteras Is-
HOLOTYPE: Brazil. Amazonas, 30 km N of December 1970, Lucas s.n.
land, 6 (F)
Manaus, 18 January 1978, Singer B 10540
minutalis Singer, 1989a: 33
(INPA)
icterinus Singer, 1948c: 30
Manaus, 29 June 1977, Singer B 10022
(INPA). Deposited as "M. brasiliensis var.
theson Hammock, 30 October 1942, Singer
minutalis"
1350 g (FH)
idroboi Singer, 1973a: 266, 327 minutus Singer, 1973a: 43, 328
HOLOTYPE: Colombia. Valle: Cali, Buenaven- HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Cer-
tura, Calima, 23 April 1968, Singer B 6333
ro Uchumachi, 25 February 1956, Singer B
(F) 1446 (F)
iguazuensis Singer, 1973a: 149, 328 misionensis Singer, 1973a: 277, 329
HOLOTYPE: Argentina. Misiones: Refugio Ya- HOLOTYPE: Argentina. Misiones: Puerto Igua-
guarete", 15 April 1957, Singer M
1024 (F) zii, 14 May 1957, Singer M 981 (F)
54 Chapter Three
MARASMIELLUS
musacearum Singer, 1973a: 239, 329 parlatorei Singer in Singer & Digilio, 1951e:
Guayaramerin, 16 March 1956, Singer B 2022 HOLOTYPE: Argentina. Tucuman: Cerro Alto
(F) del Taficillo, 6 January 1950, Singer T 835
in
napoensis Singer, 1975c: 877 (LIL) [published 1952]
HOLOTYPE: Ecuador. Napo: Shushufindi, 15 paurosporus Singer, 1958d: 228
May 1973, Singer B 7445 (F) HOLOTYPE: Mexico. Oaxaca: Huautla de Jim-
nothofagineus Singer, 1969a: 80

1533 (LIL)
HOLOTYPE: Argentina. Neuquen: Lago Cor- perangustispermus Singer, 1973a: 330

rentoso, 30 April 1958, Pujols & Gamundi
Gamundi 328 (BAFC, LIL) tura,Quinta La Flora between Cisneros and
Cordoba, 23 April 1968, Singer B 6323 (F)
nubigenus Singer, 1973a: 197, 329
HOLOTYPE: Colombia. Valle: Corr. El Vergel., pernambucensis Singer, 1973a: 233, 330
HOLOTYPE: Brazil. Pernambuco: Caramagibe,
Municip. Dagua, Finca La Araucana, 14 April 14 July 1960, Singer B 3408 (F)
1968, Singer B 6133 (F)
petiolicola Singer, 1975c: 891
oblongisporus Singer, 1973a: 254, 329 HOLOTYPE: Ecuador. Tungurahua: Rio Topo, 8
HOLOTYPE: Mexico. Oaxaca: Rancho del May 1973, Singer B 7292 (F)
Cura, below Huautla de Jimenez, 7 July 1969,
peullensis Singer, 1969a: 77
Singer M
8287 (F) HOLOTYPE: Chile. Llanquihue: Puella, 20
omphalina Singer in Singer & Digilio, 195 le: March 1959, Singer M
1924a (BAFC)
167 platyhyphes Singer, 1973a: 275, 330
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Colombia. Valle: Cali, Buenaven-
San Anta Muerte, 27 March 1949,
Javier, tura, Calima, 25 April 1968, Singer B 6360
Singer T 295 (LIL) [published in 1952] (F)
omphalophorus Singer, 1969a: 79 polyphyllm Singer, 1973a: 182, 330

HOLOTYPE: Chile. Llanquihue: Puella, 20 HOLOTYPE: Mexico. Veracruz: 7 km S of
March 1959, Singer M 1897 (SCO) Montepio, Estacion Biologica de las Tuxtlas,
orinocensis Singer, 1973a: 280, 329

8061 (F)
HOLOTYPE: Venezuela. Amazonas: Tama- potamogeton Singer, 1989a: 34

HOLOTYPE: Brazil. Amazonas: Parana do Jan-
Tama, Rio Orinoco, 21 June 1959, Wurdack
auaca, 23 May 1980, Singer B 12178 (F)
&Adderley 43097 (NY)
potassiovirens Singer, 1989a: 35
osornensis Singer, 1969a: 83
HOLOTYPE: Chile. Llanquihue: Volcan Osor- Manaus, 21 April 1977, Singer B 9782 (INPA)
no, 10 May 1967, Singer M 6846 (SGO)
pseudoparaphysatus Singer, 1973a: 195, 331
panamensis Singer, 1973a: 251, 329 HOLOTYPE: Colombia. Valle: Cali, Buenaven-
HOLOTYPE: Panama. Zona Canalis: Barro Col- tura,Quinta La Flora between Cisneros and
orado Island, 6 July 1952, Martin & Welden Cordoba, 23 April 1968, Singer B 6313 (F)
7468 (F) quercinus Singer, 1973a: 55, 331
pandoensis Singer, 1973a: 71, 329 HOLOTYPE: Colombia. Cundinamarca: Bojaca,
HOLOTYPE: Bolivia. Pando: Manuripi, 23 Bosque de La Mercedes, 8 April 1968, Singer
March 1956, Singer B 2160 (LIL) B 6010 (F)
ramealis var. consimilis Singer, 1973a: 174, 331
pantholocystis Singer, 1989a: 34
HOLOTYPE: Indonesia. Pulo-Penang, no date, HOLOTYPE: Argentina. Tucuman: Sierra de
Wawra 1407 (W) San Javier, Ciudad Universitaria, 16 January
papillatomarginatus Singer, 1973a: 183, 330
ramealis var. foliicola Singer, 1969a: 75
HOLOTYPE: Colombia. Valle: Corr. El Vergel.,
Municip. Dagua, Finca La Araucana, 14 April November M 7827 (SGO)
1967, Singer
1968, Singer B 6136 (F)
ramealis var. multifolius Singer, 1969a: 74
papillifer Singer, 1969a: 79 HOLOTYPE: Chile. Chiloe: Pudeto, May 1967,
HOLOTYPE: Chile. Llanquihue: 15 km NE of M 6793 (SGO)
Singer
Puerto Pargua, 9 May 1967, Singer M 6801
ramealis var. secretanii Singer, 1973a: 311
(SGO) HOLOTYPE: Switzerland. Vaud: NW of Bel-
paraensis Singer, 1973a: 45, 330 mont, 1
August 1970, Singer C 5035 (F). No
HOLOTYPE: Brazil. Para: Belem, 10 June 1966, Latin diagnosis cites other publication with
Singer B 4258 (BAFC) Latin

MARASMIELLUS
ramealis var. subtropicalis Singer, 1973a: 173, stenosporus Singer, 1973a: 111, 332
331 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
HOLOTYPE: Argentina. Tucumdn: Tafi del Val- Guayaramerin, 7 March 1956, Singer B 2080
le, 15 January 1950, Singer T 872 (LIL) (F)
ramealis var. tucumanensis Singer, 1973a: 175, stratosus Singer, 1975c: 893
331 HOLOTYPE: Ecuador. Tungurahua: Rio Topo,
HOLOTYPE: 30 April 1973, Singer B 7194 (F)

Argentina. Tucumdn: Sierra de
San Javier, Ciudad Universitaria, 28 January subcoracinus ssp. montanus Singer, 1973a: 208
1957, Singer T 2919 (F) HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Cer-
ramorum Singer in Singer & Digilio, 1951e: Uchumachi, 25 February 1956, Singer
ro B
1473 (BAFC). No Latin diagnosis.
163
HOLOTYPE: Tucuman: Sierra de subhirtellus Singer, 1989a: 36
Argentina.
San Javier, 1 March 1949, Singer T 174 (LIL) HOLOTYPE: Brazil. Para: Belem, 10 June 1966,
[published in 1952] Singer B 4255 (BAFC)
reductm subinodermatoides Singer, 1989a: 36
Singer, 1973a: 305, 331
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car- HOLOTYPE: Brazil. Amazonas: 30 km N of
men Pampa, 19 February 1956, Singer B 1266 Manaus, 2 January 1978, Singer B 10470 b-c
(F) (INPA)
rhodophyllus Singer, 1973a: 212, 331 subolivaceomelleus Singer, 1973a: 68, 332
HOLOTYPE: Colombia. Cauca: above Totora", HOLOTYPE: Colombia. Valle: Cali, Buenaven-
Pdramo Gabriel Lopez, San Pedro, 4 May tura, San Joaqum, 19 April 1968, Singer B
1968, Singer B 6990 d (F) 6233 (F)

riberaltensis Singer, 1973a: 281, 331 subramealis Singer in Singer & Digilio, 195 le:
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri- 157
beralta, 29 March 1956, Singer B 2324 (F) HOLOTYPE: Argentina. Tucuman: Sierra de
rosascentifolius Singer, 1989a: 35 San Javier, Parque Aconquita, 24 April 1951,
HOLOTYPE: Brazil. Amazonas: 30 km N of Singer T 1521 (LIL) [published in 1952]
Manaus, 3 April 1978, Singer B 10905 (INPA) subramealis var. catamarcensis Singer, 1973a:
sacchari Singer in Singer &
Digilio, 1951e: 170 166, 332
HOLOTYPE: Argentina. Tucuman: Marcos Paz, HOLOTYPE: Argentina. Catamarca: Cuesta del
26 March 1949, Singer T 280 (LIL) [pub- Totoral, 26 May 1959, Singer T 3368 (F)
lished in 1952] subramealis
var. Julius Singer, 1973a: 171
sanctaemarthae Singer, 1973a: 226, 332 HOLOTYPE: Brazil. Pernambuco: Camaragibe,
HOLOTYPE: Colombia. Magdalena: Sierra Ne- 7 July 1960, Singer B 3293 (F). No Latin di-
vada de Santa marta, Hacienda Cincinnati, 18 agnosis.
August 1935, Martin 3493 (F) subramealis var. platani Singer, 1973a: 168,
segregabilis Singer, 1973a: 246, 332 333
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- HOLOTYPE: Mexico. Oaxaca: below Huautla
roico, 15 February 1956, Singer B 1163 de Jimnez, 7 July 1969, Singer M
8321 (F)
(MICH) tener Singer, 1973a: 139, 333
semiustus var. abortivus Singer, 1973a: 312, HOLOTYPE: Mexico. Oaxaca: San Andres Hi-
332 dalgo, 9 July 1969, Singer M 8348 (F)
HOLOTYPE: Philippines. Manila, 13-14 No- thaxteri Singer, 1973a: 248, 333
vember 1947, Davis s. n. (FH) Type information: USA. Florida: Dade County,
septicoides Singer, 1955a: 773 Coconut Grove, Thaxter (F). No date or col-
HOLOTYPE: Brazil. Rio de Janeiro, Angra dos lection number given.
Reis, 1 October 1952, Singer B 427 (F) trichodermialis Singer, 1975c: 883
sphaerospoms Singer, 1973a: 214, 332 HOLOTYPE: Venezuela. Aragua: along road
HOLOTYPE: Argentina. Corrientes: Depto. from El Junquito, to Colonia Tovar, 44-45 km
Cosme, Paso de la Patria, 5 April 1957, Singer from Caracas, 22 June 1971, Dumont VE-440
M 903 (LIL) (NY)
stenocystis Singer, 1973a: 151, 332 tricolorf. americana Singer, 1946g: 128
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Type information: USA. Massachusetts: Arling-

Charobamba, 30 January 1956, Singer B 732 ton, Singer (FH). [Published in 1948].
(F) umbilicatus Singer, 1973a: 129, 333
stenophylloides f.
bispora Singer, 1978h: 17 HOLOTYPE: Colombia. Valle: Cali, Buenaven-
HOLOTYPE: Ecuador. Napo: Lago Agrio, 14 tura, Juanchaco, 21 April 1968, Singer B 6270
May 1973, Singer B 7439 (F) (F)
56 Chapter Three
MARASMIELLUS
umbonifer Singer, 1973a: 150, 333 alpinus Singer, 1937: 231

HOLOTYPE: Argentina. Misiones: General Bel- HOLOTYPE: USSR. Western Caucasus: Monte
grano, near San Antonio, 19 April 1957, Sing- Jatyrgvarta, July 1936, Vasilieva (LE?)
er M1080 (F) apatelius Singer, 1964f: 332
varzeae Singer, 1989a: 36 HOLOTYPE: Zaire. Bas-Zaire: Kisantu, no date,
HOLOTYPE: Brazil. Amazonas: Una da mar- Vanderyst 695 (BR)
chantaria, 1 July 1981, Singer B 12215 apogonus Singer, 1978h: 24
(INPA) HOLOTYPE: Ecuador. El Oro, 17 km. from Pi-
vinosus Singer in Singer &
Digilio, 1953b: 10 nas towards Machala on the Arenillas-Loja
HOLOTYPE: Argentina. Misiones: Cataratas del road, 29 July 1975, Dumont EC 1683 (NY)
Iguazu, 26 November 1949, Singer & Digilio aporpohyphes Singer, 1965c: 131, 333
[Singer no.] M 84 (LIL). The type information HOLOTYPE: Chile. Llanquihue: Lago Todos los
is Marasmius neosessilis
identical to that of Santos, Puella, 22 March 1959, Singer M
(1958a: 103) and Marasmius iguazuensis 1996 (LIL)
(1951e:187). Based on Singer's unpublished aporpus Singer, 1969a: 97
field notebook, however, the description of M. HOLOTYPE: Chile. Valdivia: near Hueycolla, 4
vinosus matches Singer's collection no. M 82 May 1967, Singer M 6705 (SGO)
from the same location.
araucariae Singer, 1965c: 319, 333
volvatus Singer ex. Singer, 1989a: 37 HOLOTYPE: Argentina. Misiones: Frontera,
HOLOTYPE: Brazil. Amazonas: Road Manaus- General Manuel Belgrano, 20 April 1957,
Caracarai, km 125, 20 July 1978, Singer B Singer M 1147 (LIL)
1 1 31 7 (INPA)
asemiformis Singer, 1989a: 59
xerophyticus Singer, 1973a: 57, 333 HOLOTYPE: Brazil. Amazonas: Manaus, 20
HOLOTYPE: Argentina. Tucuman: 2 km of W March 1983, Singer B 12457 (F)
Tapia, 6 January 1952, Singer T 1706 (MICH) asemus 60
Singer, 1989a:
yalae Singer, 1973a: 141, 333 HOLOTYPE: Amazonas: 30 km N of
Brazil.
HOLOTYPE: Argentina. Jujuy: Lagunas de Manaus, 18 January 1978, Singer B 10545
Yala, 13 February 1966, Singer T 5087 (F) (INPA)
aspilocephalus Singer, 1965c: 176, 334
MARASMIUS HOLOTYPE: Bolivia. Pando: Conquista, 25
March 1956, Singer B 2214 (LIL)
adisianus Singer, 1989a: 59 ater Singer, 1976a: 182
HOLOTYPE: Brazil. Amazonas: Lago Janauari, HOLOTYPE: Venezuela. Dto. Federal: El Avila,
18 March 1983, Singer and others [Singer

24 July 1972, Dumont and others VE 5980
no.] B 12454 (INPA) (NY)

atlanticus Singer, 1976a: 77
aequatorialis Singer, 1976a: 112
HOLOTYPE: Ecuador. Pichincha: Rio Machan- HOLOTYPE: USA. Florida: Highlands County,
Highlands Hammock State Park, 21 August
garo, 20 May 1973, Singer B 7538 (F)
1942, Singer F 347 (F)
aimara Singer, 1965c: 219, 333
atroincrustatus var. atroincrustatus Singer,
Charobamba, 13 1956, Singer B 1976a: 76
February
1134 (LIL) HOLOTYPE: Mexico. Oaxaca: Rancho del
albertianus Singer, 1964f: 351 Cura, 7 July 1969, Singer M

8320 (F)
HOLOTYPE: Zaire. Des Lacs Edouard- atroincrustatus var. inodorosus Singer, 1976a:
Ganta,Kivu, Pare National Albert, Kalonge S. 76

Butahu, no date, Fredericq in de Witte 8969 HOLOTYPE: Mexico. Chiapas: San Cristobal de
(BR) las Casas, 15 km W, 2 August 1969, Singer
albidomurinus Singer, 1989a: 45 M 8816 (F)
HOLOTYPE: Brazil. Amazonas: 30 km N of atrorubens var. cystidifer Singer, 1976a: 242
Manaus, 20 February 1978, Singer B 10670 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
(INPA) Guayaramerin, 8 March 1956, Singer B 1700
allocystis Singer, 1976a: 206 (LIL)
HOLOTYPE: Bolivia. El Beni: Vaca Diez, atrorubens dumontii Singer, 1976a: 243
var.
Guayaramerin, 3 km below March

village, 12 HOLOTYPE: Venezuela. Sucre: trail from Los
1956, Singer B 1875 (LIL). This species was Pocitos, to Rio Grande Arriba, 13 July 1972,
invalidly published in Singer, 1965c: 292. Dumont and others VE 4877 (NY)

MARASMIUS
austrorotula Singer, 1969a: 103 calhouniae Singer, 1989a: 63

HOLOTYPE: Argentina. Neuquen: Nahuel Hua- HOLOTYPE: USA. California: Marin County,
pi National Park, Puerto Blest, Los Cantaros, 11 November 1981, Calhoun 81-2606 (F)
10 November 1966, Singer M
6045 (BAFC) caliensis Singer, 1976a: 91
aztecus Singer, 1976a: 235 HOLOTYPE: Colombia. Valle: Cali, Saladito,
HOLOTYPE: Mexico. Mexico near Pena Blan- 15 April 1968, Singer B 6149 (F)
SE of Valle de Bravo, 26 September
ca, 1967, campinaranae Singer, 1989a: 57
Guzman 6148 (ENCB) HOLOTYPE: Brazil. Amazonas: Road Manaus-
Caracarai, km 45, 28 February 1979, Singer
bactrosporus Singer, 1965c: 127, 334
HOLOTYPE: Tucuman: Sierra de B 10724 (INPA)
Argentina.
San of Ciudad Univ-
Javier, ravine northwest
carcharm Singer, 1964f: 361
ersitaria, 9 January 1960, Singer T 3521 (LIL)
HOLOTYPE: Zaire. Forestier Central: Binga, no
Goossens-Fontana 4064 (BR)
date,
baeocephalus Singer, 1976a: 121
HOLOTYPE: carminis Singer, 1976a: 108
Ecuador. Napo, 8 May 1973, Sing-
er B 7311 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car-
(F)
bambusiniformis Singer, 1976a: 167
(LIL). This species was invalidly published in
Singer, 1965c: 156.
May 1973, Singer B 7480 (F) carneotinctus Singer, 1976a: 107
bathomphalus Singer, 1989a: 45 HOLOTYPE: Colombia. Valle: Cali, along road
to Buenaventura, 26 July 1964, Guzman 4580
(ENCB)
(INPA)
carpenterianus Singer, 1958e: 70, 71
batistae Singer, 1965c: 352
HOLOTYPE: Peru. Huanuco: Tingo Marca,
HOLOTYPE: Brazil. Pernambuco: Mata do Sao Cueva de los Pavos, 3 September 1949, Car-
Joao 16 July 1960, Singer B 3436 (BAFC)
penter 136 (LIL)
,
beniensis Singer, 1965c: 261, 334

carpophilus Singer, 1937: 231
HOLOTYPE: Bolivia. El Beni: Guayaramerin,
Type information: USSR. Western Caucasus: re-
14 March 1956, Singer B 2000 (LIL)
gion Czernoreczie, August 1936, Vasilieva
bezerrae var.bezarrae Singer, 1976a: 185 (LE?)
HOLOTYPE: Pernambuco: Caramagibe,
Brazil. castellanoi Singer, 1976a: 99
14 July 1960, Singer & Bezerra B 3409 (F) HOLOTYPE: Brazil. Pard: Estancia Pirelli, 7
bezerrae var. chitepecensis Singer, 1976a: 185 June 1966, Singer B 4199 (BAFC)
HOLOTYPE: Mexico. Chitepec, 30 October caucasicus Singer, 1937: 230
Guzman 2800 B (F)
1960, Type information: USSR. Western Caucasus: Mt.
bingaensis Singer, 1964f: 382 Caucasus, August 1936, Vasilieva (LE?)
HOLOTYPE: Zaire. Forestier Central: Binga, no cenospilus Singer, 1982d: 35
Goossens-Fontana 791 (BR)
date, HOLOTYPE: Costa Rica. Guanacaste: Parque
boliviae Singer, 1965c: 196, 334 Nacional Santa Rosa, 25 July 1981, Gomez &
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- Singer B 12468 (F)
roico, 12 February 1956, Singer B 1098 (LIL) centroamericanus Singer, 1989a: 60

HOLOTYPE: Costa Rica. Alajuela: near Canas-
brachysporus Singer, 1989a: 45
HOLOTYPE: Brazil. Amazonas: 30 km N of Upala, October 1982, Gomez 18649 (F)
Manaus, 2 January 1978, Singer B 10465 chiapasensis Singer, 1976a: 73

HOLOTYPE: Mexico. Chiapas: between Finca
(INPA)
Sospiro, and El Pozo, 4 August 1969, Singer
bromeliacearum Singer in Singer & Digilio,
M 8969 (F)
1951e: 212
chrysoblepharis Singer, 1976a: 237
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Mexico. Veracruz: Tlilapa, banks
San 30 March 1951, Singer T 1457
Javier,
of Rio Aserradero, 24 June 1969, Singer M
(LIL) [published in 1952] 8201 (F)
brunneocinctus Singer, 1976a: 156
chrysocephalus Singer, 1976a: 144
HOLOTYPE: Mexico. Chiapas: between Finca HOLOTYPE: Bolivia. El Beni: Vaca Diez, be-
Sospiro, and El Pozo, 4 August 1969, Singer low Guayaramerin, 12 March 1956, Singer B
M
8982 (F) 1936 (LIL)
buzungolo Singer, 1964f: 371 cibarius Singer, 1976a: 274
HOLOTYPE: Zaire. Bas-Zaire, near Leopold- HOLOTYPE: Peru. Cuzco: Canal, El Descanso,
ville, no date, Dubois 1501 (BR) 5 February 1951, Vargas 10050 (LIL)
58 Chapter Three
MARASMIUS
cladophyllus var. glaberripes Singer, 1976a: cundinamarcae Singer, 1976a: 102

198 HOLOTYPE: Colombia. Cundinamarca: Bojaca,
HOLOTYPE: Ecuador. Tungurahua: Rio Verde, Bosque de la Mercedes, 8 April 1968, Singer
5 of Rio Pastaza, 28 April 1973, Singer B B 6009 (F)
7160 (F) cyatheae Singer, 1936g: 157, 158
cladophyllus var. intermedius Singer, 1976a: Type information: USSR. Leningrad: Botanical
199 Gardens, February 1936, Tranzschel & Singer
HOLOTYPE: Ecuador. Napo: Sacha 4, 10 May LE?
1973, Singer B 7343 (F) dauliporpus Singer, 1989a: 63
cohaerens americanus Singer, 1976a: 247
var. HOLOTYPE: Costa Rica. San Jose": El Empal-
HOLOTYPE: USA. Michigan: Cheboygan me, 23 July 1981, Singer B 12410 (F)
County, 21 July 1953, Singer N 700 (F) dennisii Singer, 1965c: 306, 335
cohortalis var. arenicolor Singer, 1958e: 92 HOLOTYPE: Trinidad. St. Joseph, 27 Septem-
HOLOTYPE: Argentina. Tucuman: Parque Ave- ber 1949, Dennis 42 A (K). No material stud-
llaneda, 26 March 1951, Singer T 1445 (LIL). ied. The description was adapted from Dennis
No Latin diagnosis.
by the author.
cohortalis var. pseudocollinus Singer & Digilio digilioi Singer in Singer &
ex Singer, 1958e: 91 HOLOTYPE: Argentina. Tucuman: Sierra de
HOLOTYPE: Argentina. Tucuman: Anta Muer- San Javier, 24 April 1949, Digilio & Singer
ta, 26 December 1951, Singer T 1667 (LIL). [Singer no.] T 488 (LIL) [published in 1952]
No Latin diagnosis
ditopotrama Singer, 1965c: 224, 335
conquistensis Singer, 1965c: 192, 334 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
HOLOTYPE: Bolivia. Pando: Manuripi, Con- Guayaramerin, 17 March 1956, Singer B 2068
quista, 25 March 1956, Singer B 2190 (LIL) (LIL)
convoluticeps Singer, 1976a: 173 dodecaphyllus Singer, 1965c: 159, 335
HOLOTYPE: Brazil. Amazonas: Manaus, Ita- HOLOTYPE: Bolivia. Pando: Manuripi, Santa
coatiara road, 30 January 1963, G. & L. Eiten
Rosa on Rio Madre de Dios, 22 March 1956,
6 Felipe 5254 (F)
Singer B 2151 (LIL)
corrugatiformis Singer, 1964f: 374
20 km dryinus Singer, 1990e: 28
HOLOTYPE: Zaire. Forestier Central,
HOLOTYPE: Mexico. Morelos: of Tepoz- NW
NE de Yambao, no date, Louis 15 275 (BR) 27 June 1969, Singer 8251 (F) M
tlan,
corrugatus var. lacustris Singer, 1976a: 194
dysodes Singer, 1965c: 130, 335
Type information: Ecuador. Napo: Lago Agrio
(F?). Three collections were listed, but no type
Guayaramerin, 7 March 1956, Singer B 1690
were designated: 7 May 1973, Singer B 7262,
B 7422; B 7366. (LIL)
11 May 1973,
var. portonovensis Singer, 1976a: echinosphaerus Singer, 1964f: 325
corrugatus
HOLOTYPE: Zaire. Haul-Zaire: Kipushi, Kipo-
195
HOLOTYPE: Brazil. Santa Catarina: Porto po, Schmitz-Levecq 169 (BR)
no date,
Novo, Rick 576 (FH). No date given. elaeocephalus Singer, 1964f: 384
costaricensis Singer, 1989a: 61
HOLOTYPE: Zaire. Forestier Central: Yangam-
bi, Louis 14 875 (BR)
HOLOTYPE: Costa Rica. Heredia: near Puerto
Viejo, 22 December 1979, Lodge 284 (F) ellipsoidosporus Kumar, Singer, Lakhanpal &
crinis-equi var. monocotyledonum Singer, 1976a: Step, 1991b: 227
148 HOLOTYPE: India. Himachal Pradesh: Simla,
Charawara, 8 August 1983, Kumar 254 (F
Type information: Zaire. Bamania: Eala, Staner
C 325 (BR). In mat. stud. Lebrun 3009 (BR) 1084722)
was indicated as type. eorotula Singer, 1976a: 77
cryptotrichus Singer, 1976a: 72

HOLOTYPE: Mexico. Morelos: of Tepoz- NW tura, Campamento Pulpapel, 25 April 1968,
tlan, 27 June 1969, Singer 8250 (F) M Singer B 6380 (F)
cuatrecasasii Singer, 1976a: 191 epelaeus Singer, 1965c: 266, 335

HOLOTYPE: Colombia. Valle: Rio Pance near HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ar-
Cali, 1
May 1968, Singer B 6517 (F) royo Florida, 22 km W
of Riberalta, 7 April
cuimisedus Singer in Singer &
Digilio, 195 le: 1956, Singer B 3047 (LIL)
193 episemus Singer, 1964f: 365

HOLOTYPE: Argentina. Tucuman: San Pablo, HOLOTYPE: Zaire. Panzi Kivu, Des Lacs
29 March 1949, Singer T 313 (LIL) [pub- Edouard et Kivu, no date, Goossens-Fontana
lished in 1952] 5091 (BR)

MARASMIUS
equicrinis var. dicotyledonem Singer, 1965c: graminum var. subalpinus Singer, 1965c: 204,
198, 336 336
HOLOTYPE: Argentina. Catamarca: Andalgald, HOLOTYPE: Argentina. Tucumdn: Tafi del Val-
Rio Potrero, 10 February 1952, Singer T 1854 le, Quebradita del Mastil, 7 February 1959,
(LIL) Singer S 159 (LIL)
equicrinis var. rhizomorphogeton Singer ad int.,
grandisetulosus Singer, 1964f: 379
1965c: 197, 335 HOLOTYPE: Zaire. Panzi-Kivu, Des Lacs
HOLOTYPE: Bolivia. Pando: Madre de Dios, Edouard et Kivu, no date, Goossens-Fontana
Las Piedras, 5 April 1956, Singer B 2534 5076 (BR)
(LIL) var. tenuisetulosus Singer, 1964f:
grandisetulosus
equicrinis var. rhizomorphogeton f. arrhizomor- 380
phicus Singer, 1965c: 198, 336 HOLOTYPE: Zaire. Kasai: Kikwit, Vanderyst
HOLOTYPE: Bolivia. Pando: Las Piedras, 5
4015 (BR)
April 1956, Singer B 2518 (LIL)
griseofuscescens Singer, 1976a: 104
eucladopus Singer, 1965c: 169, 338 HOLOTYPE: Ecuador. Napo: Lago Agrio, 11
men Pampa, 26 February 1956, Singer B 1478 May 1973, Singer B 7361 (F)
(LIL) guanacastensis Singer, 1989a: 64
euosmus HOLOTYPE: Costa Rica. Guanacaste: Parque
Singer, 1958e: 69
HOLOTYPE: Panama. Zona Canalis: Barro Col- Nacional Santa Rosa, 25 June 1981, Singer B
orado Island, 24 August 1952, Martin & Wei- 12470 (F)

den 8702 (IA) guyanensis var. ery throe ephalus Singer, 1965c:
exiguus Singer, 1969a: 100 186, 336
May HOLOTYPE: Bolivia. El Beni: Vaca Diez, 3 km
1967, Singer M 6577 (SCO) below Guayaramerin, 12 March 1956, Singer
ferrugineus var. gardneri Singer, 1976a: 223 B 1876 (LIL)
Type information: Brazil. Minas Gerais, Gard- guyanensis var. guayanensis Singer, 1965c:
ner (K: Hooker Herbarium)
187, 336
flavus Singer, 1964f: 344 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
HOLOTYPE: Zaire. Forestier Central: Eala, no B 2014
Guayaramerin, 16 March 1956, Singer
Goosens-Fontana 151 (BR)
date,
(LIL)
flotowiophilus Singer, 1965c: 125, 336
guzmanianus Singer, 1976a: 218
HOLOTYPE: Mexico. Mexico: La Marquesa,
Huapf National Park, Arroyo Blest, 14 March
M
1790 (LIL) Parque Nacional, September 1960, Guzman
1959, Singer
2601 (ENCB). Isotype: F.
foliicola Singer, 1976a: 158
HOLOTYPE: Argentina. Tucuman: S. M. Tu- haediniformis Singer, 1964f: 363
cuman, Garden of the Institute
HOLOTYPE: Zaire. Des Lacs Edouard et Kivu,
Miguel Lillo,
T 2147 Pare National Albert, environs d'Hoysha, no
10 February 1955, Singer (LIL). See
date, Fredericq in de Witte 9432 (BR)
Sydowia 12:205, 1965 for "ad interim"
fuligineorotula Singer, 1965c: 174, 336 haematocephalus var. atroviolaceus Singer,
HOLOTYPE: La Paz: Nor-Yungas, Rio
Bolivia. 1965c: 300, 337
Yariza, 16 February 1956, Singer B 1193 HOLOTYPE: Bolivia. Pando: Manuripi, Con-
(LIL) quista, 25 March 1956, Singer 2189 (LIL)
fusicystis Singer in Singer & Digilio, 195 le:
haematocephalus var. leucophyllus Singer,
187 1976a: 214
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Bolivia. La Paz: Charobamba, 30
January 1956, Singer B 738 (LIL)
(LIL) [published in 1952]
haematocephalus var. macrocephalus Singer,
1965c: 299, 337
HOLOTYPE: Costa Rica. Alajuela: Alto Berlin,
26 July 1981, Singer & Gomez [Singer no.] HOLOTYPE: Bolivia. El Beni: Vaca Diez,
B 12490 (F) Guayaramerin, 16 March 1956, Singer B 2045
(LIL)
graminum var. schini Singer ex Singer, 1965c:
204, 336 haematocephalus var. oenechinus Singer, 1958e:
HOLOTYPE: Argentina. Catamarca: Santa 120
Rosa, Suncho, 20 January 1952, Singer T HOLOTYPE: Cuba. Cienfuegos: Soledad, no
1788 (LIL) date, White (FH)
60 Chapter Three
MARASMIUS
haematocephalus var. pseudotageticolor Singer, hylaeae var. hypsizygus Singer, 1976a: 231
1958e: 120 HOLOTYPE: Ecuador. Napo: Sacha 4 (Norte),
HOLOTYPE: Panama. Rio Sardinilla, no date, 10 May 1973, Singer B 7340 (F)
Martin & Bouch 7549 (F, IA) hylaeicola Singer, 1976a: 114
haematocephalus var. purpureomarginatus Sing- HOLOTYPE: Ecuador. Napo: Lago Agrio, 8
er, 1965c: 298, 337 May 1973, Singer B 7298 (F)
HOLOTYPE: Bolivia. El Beni: Vaca Diez, idroboi Singer, 1976a: 115
Guayaramerin, 14 March 1956, Singer B 1967 HOLOTYPE: Colombia. Valle: Cali, Cerro Hor-
(LIL) queta, 2 May 1968, Singer B 6925 (F)
haematocephalus var. rubicundus Singer, 1965c: iguazuensis Singer in Singer & Digilio, 195 le:
299, 337 189
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, HOLOTYPE: Argentina. Misiones: Cataratas del
Charobamba, 30 January 1956, Singer B 743 Iguazii, 26 November 1949, Singer & Digilio
(LIL) [Singer no.] M
84 (LIL). The type information
var. transiens Singer, 1965c: is identical to that of M. neosessilis (1958a:
haematocephalus
300, 337 103) and that of Marasmiellus vinosus
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- (1953b: 10). See notes under the latter two
27 January 1956, Singer B 592 (LIL)
roico, species, [published in 1952]
helvoloides Singer, 1976a: 175 ilicis Singer, 1953d: 142
HOLOTYPE: Colombia. Valle: Cali, Buenaven- HOLOTYPE: Brazil. Rio Grande do Sul: Taim-
tura, Calima, 22 April 1968, Singer B 6298a besinho, 5 November 1951, Singer B 88/fB
(F) 88a (LIL)
hemimycena Singer, 1965c: 348 inundabilis Singer, 1989a: 47

HOLOTYPE: Chile. Valdivia: Cordillera Pelada, HOLOTYPE: Brazil. Amazonas: Ilha Janauari,
28 March 1963, Singer M 3193 (BAFC). Also 18 March 1983, Singer and others [Singer
published in Singer et al., 1965i: 154. no.] B 12441 (F)
hendecaphyllus Singer, 1989a: 46 iodactylus Singer, 1989a: 48

HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Brazil. Amazonas: 30 km N of
Manaus, 21 April 1977, Singer B 9783 (INPA) Manaus, 29 June 1977, Singer B 10032 (INPA)
heterocheilus Singer, 1965c: 328, 337 izonetas Singer, 1989a: 48
HOLOTYPE: Bolivia. El Beni: Vaca Diez, HOLOTYPE: Brazil. Amazonas: 30 km N of
Guayaramerin, 17 March 1956, Singer B 2076 Manaus, 8 September 1977, Aguiar & Singer
(LIL) [Singer no.] B 10138 (INPA)
hexaphyllus Singer, 1989a: 46 janauariensis Singer, 1989a: 49
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Brazil. Amazonas: Ilha Janauari,
Manaus, 10 January 1978, Singer B 105 13a 18 March 1983, Singer B 12446 (F)
(INPA) katangensis Singer, 1964f: 375

hiorami var. austroamericanus Singer, 1976a: HOLOTYPE: Zaire. Haut-Zaire: Kipushi, Kipo-
140 315 (BR)

po, Schmitz-Levecq
HOLOTYPE: Bolivia. Pando: Madre de Dios, kisangensis Singer, 1964f: 324
Las Piedras, 5 April 1956, Singer B 2499 HOLOTYPE: Zaire. Haut-Zaire: Keyberg, Kis-
(LIL) anga, no date, Soyer 127 (BR)

horridulus Singer, 1986a: 367 latiusculospermus Singer, 1976a: 267
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Mexico. Mexico: Quertaro, 5 km
Manaus, 12 July 1978, Sousa & Aguiar [Sing- Wof Lobo, 26 June 1969, Guzman 1582
er no.] B 11272 (INPA). Also see Singer, (ENCB)
1989a: 47. leguminosarum Singer, 1965c: 348
hyalinotrichus Singer, 1965c: 126, 337 HOLOTYPE: Brazil. Pernambuco: Dois Irmaos,
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 13 July 1960, Singer M 3394 (BAFC)
Guayaramerin, 9 March 1956, Singer B 1779 leoninus var. aberrans Singer, 1976a: 189
(LIL) HOLOTYPE: Mexico. Veracruz: 3 km Fortm W
hydropodoides Singer, 1989a: 64 de las Flores, 24 January 1969, Singer 8188 M
HOLOTYPE: Brazil. Amazonas: Road Manaus- (F)
Caracarai, 15 September 1978, Silva [Singer leoninus var. griseoferrugineus Singer, 1965c:
no.] B 11 390 (INPA) 277, 337
hylaeae var. hylaeae Singer, 1976a: 230 HOLOTYPE: Bolivia. El Beni: Vaca Diez, 3 km
HOLOTYPE: Ecuador. Napo: Sacha 4 (Sacha below Guayaramerin in virgin forest, 12
Norte), 10 May 1973, Singer B 7342 (F) March 1956, Singer B 1919 (LIL)

MARASMIUS
leoninus var. olivascenticeps Singer, 1958e: 105 magnoliae Singer, 1945d: 435
HOLOTYPE: USA. Florida: Highlands County, Type information: USA. Florida: Alachua Coun-
Highlands Hammock State Park, 5 September ty, Gainesville, Singer (FH)
1942, Singer F 596 (F). No Latin diagnosis manuripiensis Singer, 1965c: 154, 338
leptus Singer, 1964f: 370 HOLOTYPE: Bolivia. Pando: Manuripi, Con-
HOLOTYPE: Zaire. Forestier Central: Yangam- 25 March 1956, Singer B 2227 (LIL)
quista,
bi, no date, Louis 14 969 (BR) marthae Singer, 1958e: 137
leucorotalis Singer, 1965c: 156, 337 HOLOTYPE: Argentina. Tucuman: Garden of
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio the Instituto Lillo, 14 January 1952, M. Singer
Yariza, 23 February 1956, Singer B 1414 [Singer no.] T 1730 (LIL)
(LIL) marthae var. paucilamellatus Singer, 1958e:
leucorotalis var. discopallescens Singer, 1976a: 138
103 HOLOTYPE: Argentina. Tucuman, 20 February
HOLOTYPE: 1955, Singer T 2190 (LIL)
Argentina. Tucuman: Rio Cochu-
27 February 1952, Singer T 3772 (LIL)
na, matrisdei Singer, 1965c: 259, 338
leucozonites Singer, 1965c: 163, 337

HOLOTYPE: Bolivia. Pando: Madre de Dios,
Las Piedras, 5 April 1956, Singer B 2523
Type information: Argentina. Tucuman: Sierra
de San Javier, Ciudad Universitaria, 15 Janu- (LIL)
ary 1955, Singer T 2018 (LIL). The specimen

mazatecus Singer, 1976a: 168
was also designated as the type of Marasmius HOLOTYPE: Mexico. Oaxaca: San Andres Hi-
pluvialis Singer (1965c: 163). dalgo, Sierra Mazateca, 8 July 1969, Singer M
8368 (F)
leucozonitiformis Singer, 1976a: 106
HOLOTYPE: Colombia. Valle: Dagua, Corr. El megalospermus Singer, 1965c: 183, 338
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 3 km
Vergel, Finca La Araucana, 14 April 1968,
B 6105 A below Guayaramerin, 12 March 1956, Singer
Singer (F)
B 1927 (LIL)
lilacinoalbus var. albus Singer, 1964f: 359
HOLOTYPE: no date,
Zaire. Bas-Zaire: Binga, megistosporus Singer, 1965c: 281, 338
Dubois 1533 (BR)
lilacinoalbus var. lilacinocarmineus Singer, Guayaramerin, 14 March 1956, Singer B 1971
(LIL)
1964f: 359
HOLOTYPE: melinocephalus Singer, 1965c: 257, 338
Zaire. Forestier Central: Binga, no
HOLOTYPE: Argentina. Tucuman: 5 km S of
date, Goossens-Fontana 4066 (BR)
Burruyacii, 19 February 1957, Singer 2997
linden Singer, 1948c: 33
(LIL). SeeSydowia 12: 115.
HOLOTYPE: Liberia. Memeh Town, 27 August
mesocephalus Singer, 1989a: 49
1926, Under 438 (FH)
liquidambari Singer, 1976a: 70 Manaus, 26 December 1977, Singer B 10411
HOLOTYPE: Mexico. Oaxaca: San Augustin, (INPA)
10 July 1969, Singer M
8418/8420 (F)
mesosporus Singer, 1964f: 349
lomatiae Singer, 1969a: 96 HOLOTYPE: Zaire. Forestier Central: Binga,
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- August, 1947, Goossens-Fontana 4065 (BR).
pf National Park, Puerto Blest, Los Cantaros,
mesosporus Singer, 1973c: 469
6044 (BAFC) HOLOTYPE: USA. North Carolina: Hatteras Is-
louisii Singer, 1964f: 332 land, 6 December 1970, Lucas s.n. (F)
HOLOTYPE: Zaire. Forestier Central: Yangam- microdendron Singer, 1965c: 349
bi,Reserve Isalowe, no date, Louis 14931 HOLOTYPE: Brazil. Amazonas: 30 km N of
(BR) Manaus, 19 July 1960, Singer B 3448 (BAFC)
maasii Singer, 1989a: 65 microhaedinus Singer, 1965c: 260, 338
HOLOTYPE: Brazil. Amazonas: Road Manaus- HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri-
Itacoatiara, km 65, Maas B 10179 (INPA) beralta, 8 April 1956, Singer B 2578 (LIL)
macrolobieti Singer, 1964f: 362 microrotalis Singer, 1989a: 49
HOLOTYPE: Zaire. Forestier Central, 20 km HOLOTYPE: Brazil. Amazonas: 30 km N of
NE of Yambao, no date, Louis 15 243 (BR) Manaus, 13 July 1977, Singer B 10104 (F)
magnisetulosus Singer, 1976a: 129 minusculus Singer, 1989a: 50
HOLOTYPE: Venezuela. Dto. Federal: Parque HOLOTYPE: Brazil. Amazonas: 30 km N of
Nacional El Avila, 27 February 1972, Dumont Manaus, 9 February 1978, Singer B 10649
and others VE 6206 (NY) (INPA)
62 Chapter Three
MARASMIUS
misionensis Singer, 1965c: 190, 338 nogalesii Singer, 1965c: 305, 339
HOLOTYPE: Argentina. Misiones: Iguazii, Re- HOLOTYPE: Bolivia. El Beni: Vaca Diez, 7
fugio Yaguarete, 16 April 1957, Singer M April 1956, Singer B 2558 (LIL)
1047 (LIL) nothomyrciae Singer, 1959c: 383, 384
montagnei Singer, 1965c: 283, 339 HOLOTYPE: Chile. Juan Fernandez Island:
HOLOTYPE: Bolivia. Vaca Diez,
El Beni: Masa Tierra, path up to El Camote, 1 8 March
Guayaramerin, 17 March 1956, Singer B 2089 1955, Sparre F 56 (S)
(LIL). Also published as M. montagneanus oaxacanus Singer, 1958d: 232
Singer in 1976a: 219 based on the same type HOLOTYPE: Mexico. Oaxaca: Huautla de Jim-
specimen. nez, 10 July 1957, Singer M 1519 (LIL)
mundulus Singer, 1989a: 50 ochropus Singer, 1964f: 369
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Zaire. Bas-Zaire: Kisanga, no
Manaus, 13 July 1977, Singer B 10092 date, Vanderyst s.n. (BR)
(INPA)
oleiger Singer, 1965c: 293, 339
munyozii Singer, 1969a: 100 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio
HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16 Llolosa, 31 January 1956, Singer B 776 (LIL)
6517 (SGO)
oligocystis Singer, 1958e: 73, 75
myocephalus Singer, 1976a: 271 HOLOTYPE: Brazil. Parana: near Paranagua,
HOLOTYPE: Colombia. Valle: Municipalidad
September 1952, Singer B 403 (F)
Cali, Saladito, 15 April 1968, Singer B 6152
olivascenticeps Singer, 1976a: 180
(F) HOLOTYPE: USA. Florida: Highlands County,
nanorotalis Singer, 1989a: 51
1942, Singer F 596 (F). See Sydowia 12: 105,
Manaus, 2 January 1978, Singer B 10459 1958 for "ad interim"
(INPA) onoticus Singer, 1976a: 169
napoensis Singer, 1976a: 186 HOLOTYPE: Argentina. Tucuman: Burruyacu,
5 km S of town, 19 February 1957, Singer T
May 1973, Singer B 7519 (F) 2995 was
(LIL). This species invalidly pub-
nebularum Singer, 1965c: 350 lished in Singer, 1965c: 252.
HOLOTYPE: Colombia. Boyaca: between Ra-
pacificus Singer, 1969a: 95
miriqui and Rondon, 30 July 1960, Singer B HOLOTYPE: 4
Chile. Valdivia: Hueycolla, May
3637 (BAFC) M 6707 (SGO)
1967, Singer
neglectus Singer, 1976a: 175
pallenticeps Singer, 1976a: 127
HOLOTYPE: Colombia. Valle: Cali, Buenaven- HOLOTYPE: Argentina. Misiones: Frontera,
tura, along Juanchaco road, 25 April 1968,
Manuel Belgrano near San Antonio, 19 April
Singer B 6381 (F) 1957, Singer M 11 23 (LIL)
neorotula Singer, 1989a: 51
pallidocinctus var. latisporus Singer, 1976a:
157
Manaus, 20 October 1977, Singer B 10208
(INPA)
29 January 1956, Singer B 535 (LIL)
roico,
neosessilis Singer, 1958a: 103
HOLOTYPE: Argentina. Misiones: Cataratas del pallidocinctus var. pallidocinctus Singer, 1976a:
26 November 1949, Singer & Digilio 156
Iguazii,
M
84 (LIL). This species was
HOLOTYPE: Venezuela. Dto. Federal: Mt. La
[Singer no.]
also published in Singer, 1958e: 83. The type Naiguata, above La Planta, 25 June 1971, Du-
information is identical to that of M. igua- mont and others VE 701 (NY)
zuensis (1951e: 187) and Maramiellus vinosus panamensis Singer, 1976a: 96
(1953b: 10). Based on Singer's unpublished HOLOTYPE: Panama. Zona Canalis: Barro Col-
notebook, however, the description of M. neo- orado Island, 15-16 August 1952, Martin &
matches Singer's collection no.
sessilis 57 M Welden 8733 (F)
from Iguazu, collected in 24 November 1949. pandoanus Singer, 1965c: 153, 339
neosessilis var. montepiensis Singer, 1976a: 262 HOLOTYPE: Bolivia. Pando: Manuripi, Santa
HOLOTYPE: Mexico. Veracruz: 3 km from Rosa on Rio Madre de Dios, 22 March 1956,
Montepio, 30 July 1969, Singer M 8757 (F) SingerB 2145 (LIL)
nephelogenes Singer, 1990e: 29 panerythrus Singer, 1965c: 303, 339
HOLOTYPE: Costa Rica. Puntarenas: Bosque HOLOTYPE: Bolivia. El Beni: Vaca Diez,
Nuboso, Reserva Biologica near Monteverde, Guayaramerin, 6 March 1956, Singer B 1609
25 July 1986, Singer B 14453 (F) (LIL)

MARASMIUS
pararotula Singer, 1965c: 140, 340 polychaetopus Singer, 1965c: 129, 341
HOLOTYPE: Bolivia. El Beni: Vaca Diez, HOLOTYPE: Chile. Llanquihue: Lago Todos los
Guayaramerin, 12 February 1956, Singer B Santos, Puella, 20 March 1959, Singer M

1886 (LIL) 1883 (LIL)
paulensis Singer, 1976a: 255 polycladoides Singer, 1989a: 52

HOLOTYPE: Brazil. Sao Paulo: Reserva Flores- HOLOTYPE: Panama. San Bias, Puerto Obaldia
tal, 10 km N of Sao Paulo, 17 December Darien, 22 June 1975, Dumont PA 914 (NY)
1964, Singer B 4099 (BAFC) polycystis Singer, 1976a: 254
perlongispermus Singer, 1976a: 264
HOLOTYPE: Colombia. Cauca: road to sulphur February 1956, Singer B 1499 (F)
mine at Purace", 30 April 1968, Singer B 6489 polylepidis \ax.jujuyensis Singer, 1989a: 58
(F)
perrarus Singer, 1989a: 51
HOLOTYPE: Brazil. Amazonas: 30 km N of poromycenoides Singer, 1965c: 226, 342
Manaus, 1 1 May 1977, Singer B 9856 (INPA) HOLOTYPE: Peru. Loreto: Rio Nanya, 31 Oc-
tober 1958, Lowy 365 (LIL)
perreductus Singer, 1978h: 22
HOLOTYPE: Ecuador. El Oro, 14 km from Pi- praeandinus Singer, 1965c: 353
nas towards Machala on the Arenillas-Loja HOLOTYPE: Argentina. Tucuman: Sierra de
San 8 March 1961, Singer T 3668
road, Dumont and others EC 1636 (NY, F)
Javier,
(LIL)
phaeocystis Singer, 1976a: 221
HOLOTYPE: Argentina. Misiones: Arroyo Piray
praecox Singer, 1958e: 140
Guazii, 24 February 1960, Gerrero 61 (LIL)
HOLOTYPE: Argentina. Tucuman: Yerba Buena
Tafi and Viejo, 20 January 1955, Singer T
pichinchensis Singer, 1989a: 55
2033 (LIL)
HOLOTYPE: Ecuador. Pichincha: 24 km from
praecox f. aurantiicolor Singer, 1965c: 201,
section of Quito-Machachi road, 7 August
Dumont EC 2368 340
1975, (NY)
pilgerodendri Singer, 1965c: 124, 341
HOLOTYPE: Argentina. Rio Negro: Nahuel le, Parque Aconquija, 17 February 1955, Sing-
er T 2180 (LIL)
Huapf National Park, near Puerto Blest, 15
rubroumbonatus Singer, 1965c: 202
1812 (LIL) praecox
HOLOTYPE:
f.
Argentina. Tucuman: Tafi del Val-

platyspermus Singer in Singer & Digilio, 1951e:
le, Parque Aconquija, 12 March 1949, Singer
215 T 208 (LIL)
praegrandispermus Singer, 1989a: 52
San Javier, 12 March 1949, Singer T 203 HOLOTYPE: Brazil. Amazonas: 30 km N of
(LIL) [published in 1952] Manaus, 23 June 1977, Singer B 9993 (INPA)
platyspermus f. scandens Singer, 1965c: 161,
pseudocorrugatus Singer, 1965c: 311, 340
341 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
Argentina. Guayaramerin, 16 March 1956, Singer B 2049
San Javier, Ciudad Universitaria, 24 March (LIL)
1960, Singer T 3591 (LIL). Also see 1976a: pseudocupressiformis Singer, 1989a: 53
111 HOLOTYPE: Brazil. Amazonas: 30 km N of
pleuracanthus Singer, 1965c: 313, 341 Manaus, 2 March 1978, Singer B 10743
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 3 km (INPA)
below Guayaramerin in virgin forest, 12 pseudocupressiformis var. alboflavescentulus
March 1956, Singer B 1920 (LIL) Singer, 1989a: 53
pleurotelloides Singer, 1976a: 263 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Brazil. Par<i: Estancia Pirelli, 7 Manaus, 9 February 1978, Singer B 10620
June 1966, Singer B 4240 (BAFC) (INPA)
pluvialis Singer, 1965c: 163, 341 pseudocupressiformis var. atripes Singer, 1989a:
HOLOTYPE: Argentina. Tucumdn: Sierra de 54
San Javier, Ciudad Universitaria, 15 January HOLOTYPE: Brazil. Amazonas: 30 km N of
1955, Singer T 2018 (LIL). The specimen was Manaus, 20 October 1977, Singer B 10223
also designated as the type of Marasmius leu- (INPA)
cozonites Singer (1965c: 163). pseudocupressiformis var. conspicuus Singer,
podocarpi Singer in Sing & Digilio,
1951e: 188 1989a: 53
HOLOTYPE: Argentina. Tucuman: Cerro Alto HOLOTYPE: Brazil. Amazonas: 30 km N of
del Taficillo, 4 March 1951, Singer T 1345 Manaus, 10 January 1978, Singer 10514
(LIL) [published in 1952] (INPA)
64 Chapter Three
MARASMIUS
pseudocupressiformis var. hendecaphyllophorus rotaliformis Singer, 1982d: 43, 44

Singer, 1989a: 53
HOLOTYPE: Costa Rica. Puntarenas: Monte-
HOLOTYPE: Brazil. Amazonas: 30 km N of verde, Monteverde Cloud Forest Reserve,
Manaus, 29 February 1978, Singer B 10665 Chomogo trail, December 1979, Lodge 305
(INPA) (F)
ruber Singer, 1965c: 263, 342
pseudocupressiformis var. intermedius Singer,
1989a: 53 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
HOLOTYPE: Brazil. Amazonas: 30 km N of Guayaramerin, 7 March 1956, Singer B 1638
Manaus, 2 March 1978, B 10764 (LIL)
Singer
rubromarginatus var. albilamellatus Singer,
(INPA)
1989a: 54
pseudominutus Singer, 1989a: 56
HOLOTYPE: Panama. Panama: 2-3 km N of El HOLOTYPE: Brazil. Amazonas: 30 km N of
Llano-Carti Rd., 29 June 1975, Dumont P Manaus, 18 May B 9663 (INPA)
1977, Singer
1488 (NY) rufomarginatus Singer, 1958e: 134

HOLOTYPE: USA. Florida: Dade County,
pseudoniveus Singer, 1965c: 328, 340
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 7 Simpson Park, October 1942, Singer F 1219
March 1956, Singer 1648 (LIL) (FH)
var. amylocystis Singer, 1976a: ruforotula Singer, 1948c: 34
pseudoniveus
245
theson Hammock, 3 November 1942, Singer
F 1456 (FH)
sanctixaverii Singer, 1965c: 139, 342
(F)
Argentina.
psychotriophilus Singer, 1965c: 350 San Javier, Ciudad Universitaria, 20 Decem-
HOLOTYPE: Brazil. Rio de Janeiro: Serra dos
ber 1958, Singer T 3156 (LIL)
Orgaos, 24 October 1961, Singer B 4029
sanguirotalis Singer in Singer & Digilio, 1951e:
(BAFC)
192
pusio var.graminivorus Singer, 1976a: 201 HOLOTYPE: Argentina. Tucuman: Quebrada de
Lules, 18 February 1951, Singer T 1240 (LIL)
[published in 1952]
pusio poliobasis Singer, 1965c: 280, 342
var.
schultesii Singer, 1976a: 131
HOLOTYPE: Colombia. Amazonas: Soratama,
Charobamba, 30 January 1956, Singer B 722
Rio Apaporis, between Rio Pacoa & Rio Kan-
(LIL)
anari, 3 July 1951, Schultes & Cabrera 12869
puttemansii var. oligocladus Singer, 1976a: 142 (F)
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri-
scleronematis Singer, 1989a: 54
beralta, 29 March 1956, Singer B 2330 (LIL).
Invalidly published as Marasmius oligocladus
Singer in 1965c: 190 based on the same spec-
(INPA)
imen.
scotophysinus Singer, 1965c: 135, 342
puttemansii var. subtrichorrhizus Singer, 1965c: HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Cer-
352 ro Uchumachi, 25 February 1956, Singer B
HOLOTYPE: Brazil. Pernambuco: Camaragibe, 1444 (LIL)
7 July 1960, Singer B 3251b (BAFC)
scototephrodes Singer, 1976a: 105
radicellicola Singer, 1976a: 74 HOLOTYPE: Mexico. Oaxaca: Rancho del
HOLOTYPE: Brazil. Pernambuco: Dois Irmaos, Cura, 7 July 1969, Singer 8299 (F) M
14 April 1964, Singer B 4078
sejunctus Singer, 1976a: 260
(F)
rhododendri Singer, 1947c: 224 HOLOTYPE: Bolivia. La Paz: Nor-Yungas,
Type information: Spain. Catalonia: Port of Bon- gorge near San Jeronimo, 7 February 1956,
aigua, June 1934. The species was invalidly Singer B 980 (F)
published in Singer, 1936d: 77. sessiliaffinis Singer, 1958e: 81, 82
riparius Singer in Singer & Digilio, 195 le: 198 HOLOTYPE: Panama. Zona Canalis: Barro Col-
HOLOTYPE: Argentina. Tucuman: Quebrada de orado Island, 11 July 1952, Martin & Welden
Lules, 18 February 1951, Singer T 1244 (LIL) 7606 (IA, F)
[published in 1952] setulosifolius Singer, 1965c: 312, 343
robertsonii Singer, 1965c: 199, 342 HOLOTYPE: Bolivia. El Beni: Vaca Diez,
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri- Guayaramerfn, 7 March 1956, Singer B 1649
beralta, 30 March 1956, Singer B 2349 (LIL) (LIL)

MARASMIUS
shideiae Singer, 1989a: 61 trichorhizus var. pallenticeps Singer, 1965c:

HOLOTYPE: Japan. Kyoto: Yamashina-ku, 24 178
August 1983, Shidei M
645 (F) HOLOTYPE: Argentina. Misiones: Frontera,
sibiricus Singer, 1943a: 127 Manuel Belgrano near San Antonio, 19 April
Type information: USSR. Siberia through Altai 1957, Singer M
1123 (LIL). nom. nud. No
Mtns., September. Latin diagnosis.
silvicola Singer in Singer &

trinitatis var. immarginatus Singer, 1965c: 266,
HOLOTYPE: Argentina. Tucuman: Sierra de 344
San Javier, 14 April 1951, Singer T 1479 HOLOTYPE: Bolivia. El Beni: Vaca Diez, 3 km
(LIL) [published in 1952] below Guayaramerin, 12 March 1956, Singer

B 1883 (LIL)
sphaerodermatoides Singer, 1989a: 56
HOLOTYPE: Argentina. Buenos Aires: Parque truncorum Singer, 1989a: 58
Pereyra, 20 June 1962, Horak & Singer [Sing-
HOLOTYPE: Brazil. Amazonas: Ponta Negra, 2
er no.] S 201 (BAFC) B 11267 (INPA)
July 1978, Singer
truncorum var. ramorum Singer, 1989a: 59
spiculosus Singer, 1965c: 314, 343
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio HOLOTYPE: Costa Rica. San Jose: El Empal-
Yariza, 16 February 1956, Singer B 1206 me, 25 July 1981, Singer B 12416 (F)
(LIL) tucumanus Singer in Singer & Digilio, 195 le:
spinosissimus Singer, 1950c: 193

206
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Argentina. Tucuman: Sierra de
San Javier, Anta Muerta, 27 March 1949,
San Javier, 14 April 1951, Singer T 1460
Singer T 298 (LIL, MICH) (LIL) [published in 1952]

subarborescens Singer, 1964f: 364 ustilago Singer, 1965c: 215, 344
HOLOTYPE: Zaire. Forestier Central: Eala, no HOLOTYPE: Bolivia. El Beni: Vaca Diez, 3 km
Goossens-Fontana 113 (BR) below Guayaramerin, 12 March 1956, Singer

date,
B 1870 (LIL)
submarginatus Singer, 1989a: 62
HOLOTYPE: USA. Illinois: White Pine State
valdivianus Singer, 1969a: 101
HOLOTYPE: Chile. Valdivia: Cuesta Santa El-
Park, 1
September 1975, Singer N 7539 (F)
vira, 14 May 1967, Singer M 6897 (SGO)
subruforotula Singer, 1964f: 339
variabiliceps Singer, 1965c: 165, 344
Type information: Zaire. Forestier Central: Eala.
tanyspermus Singer, 1965c: 157, 343 men Pampa, 26 February B 1531
1956, Singer
(LIL)
variabiliceps var. derubricans Singer, 1965c:
(LIL)
168, 344
tasmaniensis Singer, 1989a: 58
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio
HOLOTYPE: Australia. Tasmania: near Floren-
Yariza, 23 February 1956, Singer B 1395
tina River,27 January 1977, Engel 13629 (F)
(LIL)
tenuiparietalis Singer, 1969a: 99
variabiliceps var. heptachroinus Singer, 1965c:
167, 344
pi National Park, above Puerto Manzano, 26
June 1964, Singer M
3563 (BAFC)
tephromelanus Singer, 1989a: 62 (LIL)
variabiliceps var. mesites Singer, 1965c: 166,
Manaus, 3 June 1977, Singer B 9913 (INPA)
344
tereticeps Singer, 1965c: 170, 343 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio
HOLOTYPE: Bolivia. Pando:Madre de Dios, Yariza, 16 February 1956, Singer B 1210
Santa Teresa, 28 March 1956, Singer B 2299
(LIL)
(LIL) 1965c:
variabiliceps var. nothotereticeps Singer,
terraefirmae Singer, 1989a: 54 168, 344
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio
Manaus, 20 February 1978, Singer B 10690 Yariza, 23 February 1956, Singer B 1411
(INPA) (LIL)
tetrachrous Singer, 1965c: 171, 343 variabiliceps var. separatus Singer, 1965c: 167,
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car- 344
men Pampa, 26 February 1956, Singer B 1528 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car-
(LIL). The species eiphet was also spelled men Pampa, 26 February 1956, Singer B 1547
"tetrachroinus." (LIL)
66 Chapter Three
MARASMIUS
variabiliceps var. tucwnanensis Singer, 1965c: brunnescens Singer & Smith, 1960h: 37
166, 344 HOLOTYPE: USA. Oregon: Linn County, 31
HOLOTYPE: Argentina. Tucuman: Chicligasta, May 1939, Doty s.n. (NY)
Rio Cochuna, 20 February 1951, Singer T californica Singer & Smith, 1960h: 33
1290 (LIL) HOLOTYPE: USA. California: Guadeloupe, 9
ventalloi Singer, 1947c: 225 April 1919, Parks 524 (NY)
Type Information: Spain. Catalonia: Prat de Llo- fallax Singer & Smith, 1960h: 34
bregat, 1934. HOLOTYPE: USA. California: Guadeloupe
ventallonii Singer, 1936d: 77 Mines, no date, Parks Z-253 (NY)
Type information: Spain. Catalonia, October
foetens Singer & Smith, 1960h: 36
1934. No Latin diagnosis. HOLOTYPE: USA. Idaho: Idaho County, 10
vergeliensis Singer, 1976a: 98 August 1958, Smith 59864 (MICH)
HOLOTYPE: Colombia. Valle: Dagua, Com El idahoensis Singer &
Smith, 1960h: 43
Vergel., 27 April 1968, Singer B 6426 (F) HOLOTYPE: USA. Idaho: Idaho County, Au-
vigintifolius Singer, 1965c: 164, 345 gust 1958, Smith 60161(MICH)
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Co- maculata Singer & Smith, 1960h: 28
roico, 4 February 1956, Singer B 871 (LIL)
HOLOTYPE: USA. Oregon: Comstock County,
violeorotalis Singer, 1976a: 107 6 November 1937, Zeller 8349 (NY)
HOLOTYPE: Colombia. Valle: near Dagua road
occidentals Singer & Smith, 1960h: 42
above km 25, 28 April 1968, Singer B 6450
HOLOTYPE: USA. California: Saratoga, 23
(F)
March 1919, McMurphy 272 (NY)
witteanus Singer, 1964f: 354
HOLOTYPE: Zaire. Des Lacs Edouard etKivu,
parksii Singer & Smith, 1960h: 32
Katavu-
HOLOTYPE: USA. Oregon, Comstock County,
Environs de Kalonge, pres de la riv.
6 November 1937, Zeller 8362 (NY)
leko, no date, Fredericq in de Witte 8934
(BR) subfulva Singer & Smith, 1960h: 35
HOLOTYPE: USA. Oregon: Clackamas County,
xerampelinus Singer, 1976a: 138
16 June 1939, Gilkeys s.n. (NY)
HOLOTYPE: Mexico. Chiapas: between Finca
thaxteri Singer, 1969a: 357
Sospiro, and El Pozo, 4 March 1969, Singer
M 8961 (F) HOLOTYPE: Argentina. Rio Negro: Laguna
Frias and Paso de las Nubes, 24 March 1963,
xerophyticus Singer, 1965c: 184, 345
HOLOTYPE: Argentina. Salta: Gemes, lower Singer M 3183 (BAFC)
part of Quebrada de Yaquiasme, 7 February variabilispora Singer & Smith, 1960h: 27
1957, Singer T 2948 (LIL) HOLOTYPE: USA. Oregon: Benton County, 4
xestocephalus Singer, 1964f: 367 October 1948, Smith s.n. (NY)
HOLOTYPE: Zaire. Forestier Central: Yangam- vesiculosa Singer &
Smith, 1960h: 29
bi, no date, Louis 14 938 (BR) HOLOTYPE: USA. Oregon: Comstock County,
xylodendron Singer, 1989a: 55 20 October 1937, Zeller 8323 (NY)
(INPA)
MELANOLEUCA
yalae Singer, 1976a: 65
HOLOTYPE: Argentina. Jujuy: Lagunas de brevispora Singer, 1954b: 115, 116
pi, Cerro Otto, 12 May 1952, Singer
491 M
yangambiensis Singer, 1964f: 335
(LIL)
HOLOTYPE: Zaire. Forestier Central: He Esali,
catalaunica Singer, 1935a: 124
opposite of Yangambi, no date, Louis 7091
(BR) Type infromation: Spain. Catalonia: Encantats,
upper Pyrenees Mtns., July.
yarizae Singer, 1965c: 308, 345
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio crassotunicata Singer, 1943a: 56
Yariza, 23 February 1956, Signer B 1369 Type information: USSR-China. Mt. Tian Shan.
(LIL) cremeospora Singer, 1989a: 20
MARTELLIA San Javier, 9 March 1961, Singer T 3684 (F)
albella Singer & Smith, 1960h: 38 longispora Singer, 1952c: 206
Huapi National Park, Puerto Blest, 17 Febru- Grande, Estancia Nueva Argentina, 15 Feb-
ary 1960, Singer M
3024 (LIL) ruary 1950, Singer 281 (LIL) M

MELANOLEUCA
luteolosperma van styptica Singer, 1969a: 67 inocyboides Singer, 197 Id: 461
HOLOTYPE: Argentina. Neuquen: Puerto Man- HOLOTYPE: Chile. Valdivia: 25 km S toward
zano, 5 April 1963, Singer 3270 (BAFC) M Uni6n, 12 September 1967, Singer M 7768
mackleri Singer, 1989a: 20 (SGO)
HOLOTYPE: USA. California: Placerito Can- leucocephala Singer, 1989a: 119
yon State Park, 14 February 1982, Singer N HOLOTYPE: Costa Rica. San Jose: San Gerardo
5018 (F) de Dota, 3 August 1981, Singer B 12656 (F)
melanosarx Singer, 1954b: 114, 115 mazatecorum Singer, 1973d: 94
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- HOLOTYPE: Mexico. Oaxaca: Sierra Mazateca,
pf National Park, Quetrihue, 15 May 1952, 9 July 1969, Singer M
8385 (F)
Singer M 646 (LIL) pacifica Singer, 197 Id: 463
myceniformis Singer, 1969a: 67 HOLOTYPE: Chile. Llanquihue: 10 km N of
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, Colaco, 9 May 1967, Singer M 6807 (SGO)
Chiveria, 10 May 1965, Singer M 5703
platyphylla Singer, 197 Id: 454
(BAFC) HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
sparrei Singer, 1953b: 5 1967, Singer M 6727 (SGO)
HOLOTYPE: Argentina. Catamarca, between universitaria Singer, 197 Id: 452
Minas Capillitas and Cerro Negro, 3 March HOLOTYPE: Brazil. Pernambuco: Dois Irmaos,
1952, Sleumer & Sparre [Singer no.] T 1923 13 July 1960, Singer B 3387 (BAFC)
(LIL)
vernifera Singer, 197 Id: 449
tucumanensis Singer in Singer & Digilio, HOLOTYPE: Chile. Llanquihue: Osorno, Lago
1951e: 100
Puyehue, 14 September 1967, Singer 7523 M
HOLOTYPE: Argentina. Tucuman: Tafi del Val- (SGO)
le, 10-18 February 1950, Singer T 844 (LIL) viscosa Singer, 197 Id: 447
[published in 1952] HOLOTYPE: Chile. Llanquihue: Volcan Osor-
xavierii Singer, 1989a: 21
no, west slope, 12 May 1967, Singer M 6857
HOLOTYPE: Argentina.Tucuman: Sierra de
(SGO)
San Javier, 9 March 1961, Singer T 3682 (F)
MELANOTUS
MELANOMPHALIA
brevisporus Singer, 1978h: 57
argipoda Singer, 1975b: 242
HOLOTYPE: HOLOTYPE: Ecuador. Tungurahua: Puyo, 16
Ecuador. Napo: Lago Agrio, 11
B 7365 April 1973, Singer B 7130 (F)
May 1973, Singer (F)
decapitatus Singer, 1989a: 109
baeospora Singer, 1973d: 93
HOLOTYPE: Venezuela. Miranda: La Silla, 18
June 1971, Dumont VE 135 (NY)
columbiana Singer, 197 Id: 456 dumontii Singer, 1989a: 110
HOLOTYPE: Colombia. Boyaca: Arcabuco, 29
HOLOTYPE: Colombia. Cundinamarca: be-
tween Fomenque and Calera, 10 July 1974,
July 1960, Singer B 3587 (SGO)
Dumont and others [Dumont no.] CO 1008
cortinarioides Singer, 197 Id: 450
HOLOTYPE: (F)
Argentina. Neuquen: Nahuel Hua-
heteroloma Singer, 1989a: 110
pi National Park, Puerto Blest, Los Cantaros,
6054 (BAFC) HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 4
dwyeri Singer, 197 Id: 452 February 1956, Singer B 867 (BAFC)
HOLOTYPE: New Guinea. Admin. Australia. matrisdei Singer, 1989a: 111
Rio Wahgi Valley, no date, Dwyer 1726 (K) HOLOTYPE: Bolivia. Amazonas: Pando, 28
March 1956, Singer B 2313 (BAFC)
emarginata Singer, 197 Id: 459
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, musacearum Singer, 1989a: 111
6 May 1967, Singer M 6746 (SGO) HOLOTYPE: Brazil. Bahia: Urucuca, 8 June
globulifera Singer, Ovrebo & Lewis, 1990g: 48 1978, Singer B 11144 (INPA)
HOLOTYPE: USA. Mississippi: Perry County, patagonicus Singer, 1969a: 258
Black Creek Area, 22 July 1988, Ovrebo 2682 HOLOTYPE: Argentina. Neuquen: Puerto Man-
(F) March 1963, Singer
zano, 17 M 3060 (BAFC)
hymenocephala Singer, 1973d: 93 poliocephalus Singer, 1978h: 58
HOLOTYPE: Mexico. Veracruz: near Montepio, HOLOTYPE: Ecuador. Napo: Shushufindi, 13
20 June 1969, Singer 8092 (F) M May 1973, Singer B 7391 (F)
68 Chapter Three
MELANOTUS
polylepidis Singer, 1973d: 84 aconquijensis Singer, 1959b: 382

HOLOTYPE: Argentina. Jujuy: Lagunas de HOLOTYPE: Argentina. Tucuman: Taficillo, 23
Yala, 16 February 1966, Singer T 5215 (F) February 1955, Singer T 2205 (LIL)
adonis var. variegata Singer, 1947c: 226
MICROMPHALE Type information: Spain. Catalonia: Salardu,

1934.
austrochilense Singer, 1969a: 84 aequatorialis Singer, 1978h: 27

May HOLOTYPE: Ecuador. Napo: Lago Agrio, 14
1967, Singer M 6687 (SCO) May 1973, Singer B 7418 (F)
griseum Singer, 1973d: 22 agloea Singer, 1989a: 69

HOLOTYPE: Mexico. Veracruz: near Montepio, HOLOTYPE: Brazil. Amazonas: Road Manaus-
30 July 1969, Singer 8752 (F) M Caracarai, km 45, 25 July 1977, Singer B
latisporum Singer, 1973d: 23 10124 (INPA)
HOLOTYPE: Colombia. Cauca: above Totora, algeriensis var. hypoxera Singer, 1969a: 139
Paramo Gabriel Lopez, San Pedro, 5 May HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
1968, Singer B 6983 (F) 7 May 1965, Singer M 5593 (BAFC)
macrosporum Singer, 1973d: 24 algeriensis var. perqffinis Singer, 1969a: 139

HOLOTYPE: Brazil. Acre: Municipalidad Mad- HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
ureira, 7 October 1968, Prance and others E slope, 7 May 1967, Singer M 6781 (SCO)
7896 (F, NY) amazonica Singer, 1983c: 112
occidentale Singer, 1973d: 24 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Colombia. Valle: western Cordil- Manaus, 23 March 1978, Singer B 10826
lera, 2 May 1968, Singer B 6902 (F) (INPA)
separatum Singer, 1973d: 25 aosma Singer, 1973d: 37
HOLOTYPE: Mexico. Chiapas: between Sospi- HOLOTYPE: Brazil. Para: Belem, 10 June 1966,
ro and El Pozo, 4 August 1969, Singer M Singer B 4265 (BAFC)
8943a (F) araujae Singer, 1983c: 114
tomentosum Singer, 1965e: 35 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Brazil. Pernambuco: Sao Louren- Manaus, 26 December 1977, Singer B 10414
50, 16 July 1960, Singer B 3411 (BAFC) (INPA)
asterophora Singer, 1983c: 114
MICROPSALLIOTA HOLOTYPE: Ecuador. Napo: Shushufindi, 15
cinnamomeopallida Singer, 1982d: 37 atroincrustata Singer, 1969a: 132

HOLOTYPE: Costa Rica. Guanacaste: Parque HOLOTYPE: Argentina. Neuquen: Lago Cor-
Nacional Santa Rosa, 25 July 1981, Gomez & rentoso, 25 April 1965, Singer 3519 a M
Singer B 12460 (F) (BAFC)
purpurea Singer, 1978H: 48 austroalcalina Singer, 1959b: 384
HOLOTYPE: Ecuador. Napo: Sacha Norte, Sa- HOLOTYPE: Argentina. Tucuman: Rio de los
cha 4, 10 May 1973, Singer B 7354 (F) Sosas, 4 February 1955, Singer T 2122 (LIL)
austroavenacea Singer, 1969a: 115
MNIOPETALUM HOLOTYPE: Argentina. Rio Negro: Laguna
March 1963, Singer
Frias, 21 M
3 125 (BAFC)
bisporum Singer ex Singer, 1966a: 10 austrocapillaris Singer, 1959c: 391

HOLOTYPE: Argentina. Tucuman: Tafi del Val- HOLOTYPE: Chile. Juan Fernandez Island:
le, Quebradita del Mastil, 25 April 1959, Masa Tierra, Quebrada Piedra Agujereada, 12
Guerrero, Singer T 3335 (LIL, BAFC) March 1955, Sparre F 36 (S). Isotype: LIL?
megalosporum Singer, 1969a: 87 austrororida Singer, 1959c: 393, 394
HOLOTYPE: Chile. Valdivia, Fundo Universi- HOLOTYPE: Chile. Juan Fernandez Island:
tario, 13 September 1967, Singer M 7404 Masa Tierra, Lower Quebrada Pangal, 24
(SCO) March 1955, Sparre F 66 (S)
biornata Singer, 1973d: 38
MYCENA HOLOTYPE: Brazil. Para: Belem, 10 July 1966,
Singer B 4267 (BAFC)
abieticola Singer, 1973d: 37 biornata var. manausensis Singer, 1983c: 114
HOLOTYPE: Mexico. Dto. Federal, near San HOLOTYPE: Brazil. Amazonas: 30 km N of
Pedro, 16 July 1969, Singer MA 33 (F) Manaus, 21 July 1977, Singer B 9785 (INPA)

MYCENA
blechnophila Singer, 19651: 156 desfontaineae Singer, 1965i: 157

HOLOTYPE: Chile. Valdivia: Cordillera Pelada, HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
3208 (BAFC) 28 March 1963, Singer M 3211 (BAFC)
caesiocana Singer, 1969a: 110 dinae Singer, 1973d: 39
HOLOTYPE: Chile. Santiago: near Ranque, 6 HOLOTYPE: Argentina. Jujuy: near Posta de
July 1967, Singer M 7156 (SGO) Lozano, 20 February 1966, Singer & Sejzer T
carminis Singer, 1973d: 39 5310 (F)
discogena Singer, 1959c: 388, 389
February 1956, Singer s.n. (LIL) HOLOTYPE: Chile. Juan Fernandez Island:
castaneomarginata Singer, 1989a: 70 Masa Tierra, along path to El Camote, 18

HOLOTYPE: Brazil. Amazonas: Road Manaus- March 1955, Sparre F 58a (S). Isotype: LIL?
Caracaraf, km 45, 6 November 1977, Freire dryopteridis Singer, 1959c: 387, 388
[Singer no.] B 10268 (INPA) HOLOTYPE: Chile. Juan Fernandez Island:
castaneostipitata Singer, 1989a: 70 Masa Tierra, Quebrada de la Damajuana, 3
HOLOTYPE: Costa Rica. Alajuela: Alto Berlin, March 1955, Sparre F 15 (S). Isotype: LIL?
26 July 1981, Singer B 12493 (F) dumontii Singer, 1989a: 72
chlorinosma Singer, 1937: 232 HOLOTYPE: Colombia. Cundinamarca: be-
HOLOTYPE: USSR. Leningrad: Botanical Gar- tween Zipaquira and Pacho, 9 June 1976, Du-
dens, May 1937 (LE?) mont CO
4476 (NY)
chloroxantha Singer, 1983c: 114
endoglobulosa Singer, 1978H: 29
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Ecuador. Napo: Lago Agrio, 11
Manaus, 23 March 1978, Singer B 10836 7362 (F) B
May 1973, Singer
(INPA) 1969a: 111
eucryphiarum Singer,
chrysocorypha Singer, 1938a: 197 HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
HOLOTYPE: USSR. Oirotia: Kuraiski Khrebet, 7 May 1965, Singer M 5578 (BAFC)
19 August 1937, Singer (LE?)
februaria Singer, 1959b: 386
chusqueophila Singer, 1965i: 159 HOLOTYPE: Argentina. Tucuman: Taff del Val-
Parque Aconquija, 17 February 1955, Sing-
28 March 1963, Singer M 3202 (BAFC) le,
er T 2184 (LIL)
cnaphocystis Singer, 1989a: 70
fernandeziana Singer, 1959c: 392
Manaus, i5 December 1977, Singer B 10317
Masa Tierra, Lower Quebrada Pangal, 18
(INPA)
March 1955, Sparre F 69 (S). Isotype: LIL?
costaricensis Singer, 1982d: 41
HOLOTYPE: Costa Rica. Guanacaste: Parque fernandeziana var. osornoana Singer, 1969a:
139
HOLOTYPE: Chile. Osorno: Lago Puyehue, 14
Singer B 12479 a (F)
cyanella Singer, 1969a: 110
October 1967, Singer M
7526 (SGO)
HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May fuscocystidiata Singer, 1989a: 72

1967, Singer M 6706 (SGO) HOLOTYPE: Colombia. Valle : Cali, Buenaven-
cyanocephala Singer, 1969a: 112
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, (F)
10 May 1965, Singer M 5732 (BAFC) gloeoloma Singer, 1989a: 72

cyanosyringea Singer, 1982d: 44
HOLOTYPE: Costa Rica. Puntarenas: Monte- Manaus, 26 December 1977, Singer B 10407
verde, Monteverde Cloud Forest Reserve, 29 (INPA)
December 1979, Lodge 306 (F) glutinocothurnata Singer, 1989a: 73
debiliformis Singer, 1978H: 28
HOLOTYPE: Ecuador. Tungurahua, along Am- Manaus, 9 February 1978, Singer B 10634
bato-Puyo road ca 73 km from Ambato, Du- (INPA)
mont EC 1418 (NY) gomezii Singer, 1989a: 73
delica Singer, 1989a: 71 HOLOTYPE: Costa Rica. San Jos<: El Empal-
HOLOTYPE: Brazil. Bahia: Itabuna, 7 June me, 23 July 1981, Singer B 12387 (F)
1978, Singer B
11121 (INPA) griseoradiata Singer, 1989a: 74
depilata Singer, 1989a: 72 HOLOTYPE: Costa Rica. Heredia: Puerto Viejo,
HOLOTYPE: Brazil. Amazonas: 30 km N of OTS Station "La Selva", 23 December 1979,
Manaus, 18 May 1977, Singer B 9634 (INPA) Lodge 274 (F)
70 Chapter Three
MYCENA
griseoradiatella Singer, 1989a: 74 ixoleuca Singer, 1973d: 41

HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Bolivia. El Beni: Vaca Diez, 7
Manaus, 20 February 1978, Singer B 10674 March 1956, Singer s.n. (BAFC)
(INPA) ixoxantha Singer, 1973d: 42
griseorete Singer, 1961b: 64 HOLOTYPE: Colombia. Cauca: near Vulcan Pu-
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 27 race, 30 April 1968, Singer B 6476 (F)
January 1956, Singer B 566 (LIL) juaniicola Singer, 1959c: 390, 391
haematopoda var. chilensis Singer, 1969a: 107 HOLOTYPE: Chile. Juan Fernandez Island:
HOLOTYPE: Chile. Valdivia: Hueycolla, 7 May Masa Tierra, El Yunque, 6 March 1955, Kun-
1965, Singer M 5592 (BAFC) kel, Charpentier [Sparre no.] F 23 (S). Iso-
helminthobasis Singer, 1969a: 137 type: LIL?

HOLOTYPE: Chile. Valdivia: Cordillera Pelada, kermesina Singer, 1986a: 406
E slope, 7 May 1967, Singer M 6760 (SCO) HOLOTYPE: Brazil. Amazonas: 30 km N of
hemitrichialis Singer, 1989a: 74 Manaus, 31 October 1977, Araujo Aguiar
HOLOTYPE: Brazil. Amazonas: 30 km N of [Singer no.] B 10252 (INPA). For species de-
Manaus, 23 March 1978, Singer B 10851 scription see 1989a:77.
(INPA) kuhneri Singer, 1943a: 138
herbarum Singer, 1943a: 138 Type information: USSR. Altai Mtns., August.
Type information: USSR. Altai Mtns., Oirotia: kuthanii Singer, 1989a: 78

near lake Telez. Rensem, August. HOLOTYPE: Czechoslovakia. Moravia: Morav-
heroica Singer, 1959b: 387 sko-Slovanskie Beskidy, 20 April 1974, Sing-
HOLOTYPE: Argentina. Tucuman: Taficillo, 22 er & Kuthan [Singer no.] C 5605 (F)
February 1955, Singer T 2235 (LIL) lacrimans Singer, 1989a: 78

heterotrama Singer, 1973d: 40 HOLOTYPE: Brazil. Amazonas: Reserva Ducke,
HOLOTYPE: Chile. Santiago: 2 km from 4 December 1978, Singer B 11429 (INPA)

Ranque, 6 July 1967, Singer M 7153 (SCO) lazoi Singer, 1969a: 124
hyalinotricha Singer, 1969a: 124 HOLOTYPE: Chile. Santiago: Cerro Robles, 17
HOLOTYPE: Chile. Llanquihue: 10 km N of June 1967, Lazo & Singer [Singer no.] M
Colaco, 9 May 1967, Singer M 6810 (SCO) 6973 (SGO)
hydropodoides Singer, 1989a: 75 lecythidacearum Singer, 1989a: 79
Manaus, 26 December 1977, Singer B 10421 Manaus, 3 June 1977, Singer B 9920 (INPA)
(INPA) leucoxantha Singer, 1973d: 42
hypsizyga Singer, 1969a: 122 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 30
HOLOTYPE: Argentina. Buenos Aires: Sierra January 1956, Singer B 742 (LIL, BAFC)
de la Ventana, 14 November 1962, Singer S lohwagii Singer, 1929b: 93
231 (BAFC) Type information: USSR. Caucasus.
icterinoides Singer, 1989a: 75 longicrinita Singer, 1989a: 79
HOLOTYPE: Brazil. Amazonas: Road Manaus- HOLOTYPE: Brazil. Amazonas: 30 km N of
Caracaraf, km 45, 11 July 1977, Singer B Manaus, 13 July 1977, Singer B 10106
10087 (F) (INPA)
idroboi Singer, 1973d: 41 macrocystidiata Singer, 1936i: 430
HOLOTYPE: Colombia. Valle: Cali, 16 April No type information.
1968, Singer & Idrobo [Singer no.] B 6157 melandeta Singer, 1973d: 43
(F) HOLOTYPE: Mexico. Dto. Federal, between
igapoensis Singer, 1989a: 76 San Pedro de Nexapa &, Paso de Cortes, near
HOLOTYPE: Brazil. Amazonas: Igarape do Tar- Barranca de Ameyalco, 21 July 1969, Singer
umazinho, 19 March 1980, Aguiar & Singer MA 051 (F)
[Singer no.] B 12035 (INPA) melinocephala Singer, 1973d: 43
indica Singer, 1989a: 76 HOLOTYPE: Argentina. Jujuy: Lagunas de
HOLOTYPE: India. Himachal Pradesh: Simla, Yala, 11 February 1966, Singer T 5032
Tutu, 20 August 1979, Atri 12 (F) (BAFC)
intervenosa Singer, 1989a: 77 metuloidifera Singer, 1969a: 116
HOLOTYPE: Brazil. Amazonas: 30 km N of HOLOTYPE: Chile. Valdivia: Hueycolla, 1
May
Manaus, 6 April 1977, Singer B 9615 (INPA) 1967, Singer M 6572 (SGO)
ionocephala Singer, 1989a: 77 microjonia Singer, 1989a: 80
Manaus, 29 June 1977, Singer B 10025 Manaus, 29 August 1977, Aguiar & Singer
(INPA) [Singer no.] B 10143 (INPA)

MYCENA
microleuca Singer, 1965i: 157 neotropicalis Singer, 1989a: 82

HOLOTYPE: Argentina. Rio Negro: Huapf HOLOTYPE: Tucuman: Sierra de
Argentina.
Frias, 21 March 1963, Singer M 3096/1 San 24 April 1951, Singer T 1523
Javier,
(BAFC) (LIL). This is a new name for Marasmiellus

microleucoides Singer, 1989a: 80 foliicola Singer (195 le: 148).
HOLOTYPE: Argentina. Jujuy: Lagunas de nothofagetorum Singer, 1969a: 114
Yala, 9 February 1965, Singer T 3898 (F) HOLOTYPE: Argentina. Rio Negro: Laguna
micromelaena Singer, 1989a: 81 Frias, 21 March 1963, Singer M 3115 (BAFC)
HOLOTYPE: Brazil. Amazonas: 30 km N of nothomyriciae Singer, 1959c: 389, 390
Manaus, 10 November 1977, Singer B 10276
(INPA) Masa Tierra, path up to ElCamote, 1 8 March

& 1955, Sparre F 57 (S). Isotype: LIL?
micromphale Singer in Singer Digilio, 195 le:
236 olivaceoalcalina Singer, 1938a: 198
HOLOTYPE: Tucuman: Las Len- Type information: USSR. Oirotia, Singer & Vas-
Argentina.
ilieva (LE?)
guas, 29 February 1951, Singer T 1268 (LIL)
omniumsanctorum Singer, 1969a: 137
[published in 1952]
HOLOTYPE: Chile. Llanquihue: Puella, 20
micropolia Singer, 1989a: 81
March 1959, Singer M 1924 (SGO)
Manaus, 8 August 1977, Singer B 10133 omphalophora Singer, 1978h: 30
HOLOTYPE: Ecuador. Tungurahua: Rio Verde,
(INPA)
28 June 1973, Singer B 7168 (F)
microstena Singer, 1989a: 81
oreadeoides Singer, 1973d: 45
km B HOLOTYPE: Zaire. Kivu: Panzi-kivu, Novem-
Caracarai, 45, 10 August 1978, Singer
ber 1948, Goossens-Fontana 5068 (BR)
11357 (INPA)
palmivora Singer, 1961b: 64
microtephra Singer, 1989a: 82
HOLOTYPE: USA. Florida, Highlands Ham-
mock State Park, 11 August 1942, Singer F
158 (F)
microtrichialis Singer, 1989a: 82
HOLOTYPE: Brazil. Amazonas: 30 km N of pantopolia Singer, 1989a: 82
Manaus, 10 January 1978, Singer B 10506a
(INPA)
(INPA)
microxantha Singer, 1989a: 82
papilligera Singer, 1978h: 31
HOLOTYPE: Brazil. Amazonas: Igarape do Tar-
umazinho, 8 September 1978, Singer B 11388
(INPA)
paraboliciformis Singer, 1969a: 134
minutuliaffinis Singer, 1969a: 131 HOLOTYPE: Argentina. Rio Negro: Nahuel
HOLOTYPE: Chile. Valdivia: Los Conales, 14
Huapi National Park, Brazo Tristeza, 7 May
May 1967, Singer M 6909 (SCO) 1964, Singer M 3992 (BAFC)
mostnyae Singer, 1969a: 108
patagonica Singer, 1969a: 121
HOLOTYPE: Chile. Osorno: Puyehue, 13 May HOLOTYPE: Rio Negro: Nahuel
Argentina.
1967, Singer M 6867 (SGO) Huapi National Park, between Laguna Frias
multicaudata Singer, 1973d: 44 and Paso de las Nubes, 24 March 1963, Singer
HOLOTYPE: Colombia. Cundinamarca: Salto M3164 (BAFC)
de Tequendama, 24 July 1960, Singer B 3501
pearsoniana Dennis in Singer, 1958d: 233
(BAFC) HOLOTYPE: Mexico. Puebla: East slopes of
munyozii Singer, 1969a: 126 Popocatpetl, 21 July 1957, Singer 1606 M
HOLOTYPE: Chile. Valparaiso: Quintero, 5 June
(LIL)
1967, Villagrdn [Singer no.] 6951 (SGO) M piguicola Singer, 1978h: 32
nanoclavula Singer, 1989a: 82 HOLOTYPE: Ecuador. Tungurahua: Puyo, 26
HOLOTYPE: Brazil. Amazonas: 30 km N of April 1973, Singer B 7131 (F)
Manaus, 23 March 1978, Singer B 10829
piterbargii Singer, 1969a: 116
(INPA) HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
neospeirea Singer, 1973d: 44 pi National Park, above Puerto Manzano, 26
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 16 April 1964, Piterbarg [Singer no.] M 3557
February 1956, Singer B 1023 (BAFC) (BAFC)
72 Chapter Three
MYCENA
pluvialis Singer, 1989a: 83 subtenerrima Singer, 1989a: 85

Manaus, 13 July 1977, Singer B 10093 Manaus, 14 June 1977, Singer B 9942 (INPA)
(INPA) subulifera Singer, 1969a: 136
poecila Singer, 1989a: 83 HOLOTYPE: Chile. Valdivia: 10 km N of Co-
HOLOTYPE: Brazil. Amazonas: 30 km N of laco, 9 May 1967, Singer M 6809 (SGO)
Manaus, 23 March 1978, Singer B 10849 sudorella Singer, 1943a: 142

(INPA) Type information: USSR. Altai Mtns.: Lake Te-
poliocephala Singer, 1989a: 84 lezkoje, August.
HOLOTYPE: Brazil. Amazonas: 30 km N of tetrasphaerophora Singer, 1959c: 392, 393

Manaus, 9 February 1978, Singer B 10625 HOLOTYPE: Chile. Juan Fernandez Island:
Masa Tierra, Quebrada de la Damajuana, 3
(INPA)
March 1955, Sparre F 9 (S)
porphyrocephala Singer, 1989a: 84
HOLOTYPE: Brazil. Amazonas: 30 km N of trichocephala Singer, 1973d: 47
B 10500 HOLOTYPE: Brazil. Para: Estanicia Pirelli, 7
Manaus, 10 January 1978, Singer
June 1966, Singer B 4232 (BAFC)
(INPA)
triplotricha Singer, 1959c: 386, 387
pratensis Singer, 1973d: 46
HOLOTYPE: Uruguay. Maldonado, 27 May Masa Tierra, path up to El Camote, 18 March
1966, Singer B 4191 (BAFC)
1955, Sparre B 54 (S)
pseudoalnicola Singer, 1969a: 133 130
turficola Singer, 1969a:
HOLOTYPE: Argentina. Rio Negro: near Santa
HOLOTYPE: Chile. Valdivia: "Turbera Hernan-
Maria, 8 April 1963, Singer M 3321 (BAFC)
dez", 6 May 1967, Singer M 6745 (SGO)
pseudostylotabes Singer, 1973d: 46 valdiviana Singer, 1969a: 118
HOLOTYPE: Mexico. Dto. Federal, Barranca HOLOTYPE: Chile. Valdivia: Hueycolla, 7 May
Ameyalco, 14 km viae Paso de Cortes, 16 1967, Singer M 5466 (BAFC)
June 1969, Singer MA 042 (F)
vasilievae Singer, 1943a: 141
ribesina Singer, 1969a: 113
Type information: USSR. Altai Mtns.: Lake Te-
HOLOTYPE: Argentina. Neuquen: Lago Cor- lezkoje, August.
rentoso, 11 March 1963, Singer M
3500
xanthocephala Singer, 1973d: 48
(BAFC) HOLOTYPE: Colombia. Valle: Cali, 15 April
rosaceifolia Singer, 1973d: 47 1968, Singer B 6148 (F)
yalensis Singer, 1973d: 48
February 1956, Singer T 1102 (BAFC) HOLOTYPE: Argentina. Jujuy: Lagunas de
rubrimontana Singer, 1969a: 118 Yala, 9 February 1965, Singer T 3909 a (F)
HOLOTYPE: Argentina. Neuquen: Cerro Colo-
March 1959, Singer
rado, 10 M 1734 (LIL) MYCENELLA Singer, 1938e: 9
saxegothaeae Singer, 1969a: 123
HOLOTYPE: Argentina. Rio Negro: Santa Mar- Type species: Mycena (Mycenella) margaritis-
ia, 19 April 1965, Singer M
5059 (BAFC) pora Lange
semipilosa Singer, 1989a: 84
aristoteliae Singer, 1959c: 380, 381
B 10821a HOLOTYPE: Chile. Juan Fernandez Island:
Manaus, 23 March 1978, Singer
Masa Tierra, lower part of Quebrada Pangal,
(INPA)
24 March 1955, Sparre F 67 (S). Isotype:
sosarum Singer, 1959b: 389
LIL?
cyatheae Singer, 1938e: 9
Sosas, 4 February 1955, Singer T 2128 (LIL)
Type information: USSR. Leningrad, 1936,
sotae Singer, 1961b: 62 Tranzschel & Singer (LE?)
fuliginosa Singer, 1964d: 166
February 1956, Singer & Sola [Singer no.] B HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
1073 (LIL)
pf National Park, Brazo Blest, path to "Los
squamulosa Singer, 1961b: 63 Cantaros", 23 March 1963, Singer 3143
HOLOTYPE: M. Tu-
Argentina. Tucuman: S. (BAFC)
cuman, 4 January 1959, Singer T 3175 (LIL) funebris Singer, 1952c: 213
subinamyloidea Singer, 1943a: 139 HOLOTYPE: Argentina. Tierra del Fuego: Rio
Type information: USSR. Altai Mtns. Central, Grande, Estancia Nueva Argentina, 15 Feb-
August. ruary 1950, Singer M
286 (LIL)

MYCENELLA
minima Singer, 1969a: 94 pallidissima Singer in Singer & Digilio, 1951e:

HOLOTYPE: Chile. Osorno: Puerto Octay, 15 366
September 1967, Singer 7533 (SGO) M HOLOTYPE: Argentina. Tucuman: Tafi del Val-
polylepidis Singer, 1989a: 44 le, Parque Aconquija, 24 November 1950,
HOLOTYPE: Argentina. Jujuy: Lagunas de Singer T 1034 (LIL) [published in 1952]
Yala, 16 February 1966, Singer T 5203 pantelaeodes Singer, 1954b: 138
(BAFC) HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
subtropicalis Singer, 1953b: 7 pi National Park, Quetrihue, 16 May 1952,
HOLOTYPE: Argentina. Tucuman: Rio Cochu- Singer M 682 (LIL)
na, 1 1
February 1952, Singer T 1840 (LIL) polytropa Singer, 1953a: 237
MYCOALVIMIA Singer, 198 la: 504 Grande, Estancia Nueva Argentina, 17 Feb-
337 (LIL)
Type species: Mycoalvimia theobromicola Singer praeandina Singer, 1950a: 149
theobromicola Singer, 198 la: 504
Lules, 2 April 1949, Singer T 373 (LIL)
HOLOTYPE: Brazil. Bahia: Itabuna, 6 June
subvelutina Singer, 1941e: 95
1978, Singer B 11102 (INPA)
Type information: USSR. Altai Mtns.: Jaila,
Singer &
Vasilieva (LE). Also published in
MYXOCOLLYBIA Singer, 1950h: 488 without Latin diagnosis.
subvelutina Singer, 1950a: 149
platycystis Singer, 1943a: 118
Type information: USSR. Altai Mtns.: near Ar-
Type information: USSR. Altai Mtns.: Artubash
tubash, 1 September 1937, Singer A 298 (LE).
Oirotiae, aimak Turoczak, 9 January 1937.
Also published in Singer, 1941e: 95. This may
be a later homonym.
NAEMATOLOMA
amazonicum Singer, 1973d: 79 NEOCLITOCYBE Singer, 1961b: 55
pore, Guajara mirim, 10 March 1956, Singer Type species: Omphalia byssiseda Bres.
B 1802 (BAFC)
aberrans Singer, 1969a: 72
castilloi Singer, 1969a: 236
HOLOTYPE: HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
Argentina. Rio Negro: Laguna
10 May 1965, Singer M 5731 (SGO)
Frias, 24 March 1963, Singer 3133 a M
ciliata Singer, 1989a: 25
(BAFC)
nudum Singer, 1973d: 80 HOLOTYPE: Brazil. Amazonas, Road Manaus-
HOLOTYPE: Brazil. Para: Estancia Pirelli, 7 Itacoatjara, km 65, 8 October 1977, Singer B
March 1966, Singer 10183 (INPA)
s.n. (BAFC)
popperianum Singer, 1973d: 81 latispora Singer, 1973d: 22
HOLOTYPE: USA. California: San Francisco, HOLOTYPE: Argentina. Tucuman: Quebrada de
20 February 1969, Singer N 2041 la Higuera, 25 January 1955, Singer T 2082
(F)
(BAFC). The type information for this species
is Marasmiellus clitocybe
identical to that of
NAUCORIA
Singer (1955c: 148), except for the herbarium
of deposit.
curvipes Singer, 19545: 135, 136
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- lifotama Singer, 1967c: 359
HOLOTYPE: Zaire. Eala, September 1923,
Singer M 744 (LIL) Goossens-Fontana 292 (BR)
macrospora Singer, 1941e: 94 membranacea Singer &

Grinling, 1967c: 358
Type information: USSR. Tatar, Vasilieva (LE?) HOLOTYPE: Zaire. Brazzaville, February 1966,
olivaceiceps Singer, 1954b: 137 Grinling 60,209 (BAFC)

HOLOTYPE: Argentina. Neuquen: Nahuel Hua- microspora Singer, 1969a: 71
pf National Park, Quetrihue, 20 May 1952, HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
Singer M
791 (LIL) Chiveria, 10 May 1965, Singer M 5738
olivaceonana Singer, 1954b: 136, 137 (BAFC)
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- myceliosa Singer, 1961b: 56
pi National Park, Isla Victoria, 17 May 1952, HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 16
Singer M 727 (LIL) February 1956, Singer B 1211 (LIL)
74 Chapter Three
NEOCLITOCYBE
portentosa Singer, 1961b: 56

amarescens Singer, 1973d: 20
HOLOTYPE: Vaca Diez, Ri-
Bolivia. El Beni: HOLOTYPE: Brazil. Pernambuco: Dois Irmaos
beralta, 31 March 1956, Singer B 2368 (LIL) near Recife, 21 July 1964, Singer B 5502
sanctaerosae Singer, 1989a: 26 (BAFC)

HOLOTYPE: Bolivia. Pando: Manuripi, 22 tropicalis Singer, 1973d: 19
March 1956, Singer B 2150 (LIL) HOLOTYPE: Bolivia. El Beni: Vaca Diez, 17
sublateralis Singer, 1965e: 37 March 1956, Singer B 2081 (LIL, BAFC)

HOLOTYPE: Brazil. Pernambuco: Mata de Sao
Joao, 16 July 1960, Singer B 3437 (BAFC) OCTAVIANINA
macrospora Singer & Smith, 1960h: 9
NEOHYGROPHORUS Singer ex Singer, HOLOTYPE: USA. Oregon: Clackamas County,
1961b: 46 9 August 1937, Rogers s.n. (NY)
papyracea Singer & Smith, 1960h: 13
Type species: Hygrophorm angelesianus Smith HOLOTYPE: USA. California, Trinidad, 21 No-
& Hesler vember 1939, Parks 6347 (NY)
rogersii Singer & Smith, 1960h: 12
NEOPAXILLUS Singer, 1948a: 262 HOLOTYPE: USA. Oregon: Benton County, 24
May 1939, Rogers s.n. (NY)
Type species: Neopaxillus echinosporus Singer
OMPHALIA
echinosporus Singer, 1948a: 262
HOLOTYPE: Brazil, near Couto, 1936, Rick swanetica Singer, 1931: 530
(FH) Type information: USSR. Caucasus: KM.
plumbem Singer & Lodge, 1988i: 207, 208
HOLOTYPE: USA. Puerto Rico: Luguillo OMPHALINA
Mtns., Caribbean National Forest, El Verde, 5
September 1986, Lodge & Prieto PR 38 (F) antarctica Singer, 1956b: 16
HOLOTYPE: Antarctica. Shetlands Meridion-
ales: Decepcion, Behn & Sparre [Sparre no.]
NEOSECOTIUM Singer & Smith, 1960e: 154
T 2009 (LIL)
Type species: Secotium macrosporum Lloyd baeospora Singer, 1977c: 120
va, 19 July 1974, Veselsky,Kuthan & Singer
NIVATOGASTRIUM Singer & Smith, 1959g: C 5577 (F)
[Singer no.]
224
columbiana Singer, 1970a: 20
HOLOTYPE: Colombia. Cundinamarca: Paramo
Type species: Secotium nubigenum Harkness
above Pacho, near road to Bogota, 26 July
1960, Singer B 3531 (BAFC)
NOLANEA
defibulata Singer, 1952c: 190
subcaelestina Singer, 1949b: 91
Type information: Finland. Karelia: Kivacz, 24 M 355 (LIL)
ruary 1950, Singer
August 1936 (LE?)
infumata Singer, 1989a: 13
HOLOTYPE: Czechoslovakia. Slovakia, be-
NOTHOCLAVULINA Singer, 1970a: 18 tween Kezmarsko & Kezmarske & Zlaby, 7
September 1974, Singer & Kuthan [Singer
Type species: Nothoclavulina ditopa Singer C 5921 (F)
no.]
ditopa Singer, 1970a: 18 mostnyae Singer, 1973d: 15

HOLOTYPE: Chile. Santiago: Quebrada Diego,
ta, 10 April 1949, Singer & Digilio [Singer 6 July 1967, Singer M 7130 (SGO)
no.] T 403 (LIL). The type information is oreades Singer, 1946g: 123
identical to that of Armillariella ditopa Singer HOLOTYPE: USA. New Hampshire: White
(1950c: 191). Mtns. (FH?) .
pyxidata var. pluviarum Singer in Singer & Di-
NOTHOPANUS Singer, 1944a: 364 gilio, 1951e: 67

HOLOTYPE: Argentina. Tucuman: Tapia, 15
Type species: Agaricus (Pleurotus) eugrammus April 1951, Singer T 1497 (LIL) [published in
Mont. 1952]

OMPHALINA
subobscura Singer, 1970a: 21 sanmartiniana Singer, 1969a: 367

HOLOTYPE: Argentina. Tucuman: San Pedro HOLOTYPE: Argentina. Mendoza: Las Heras,
de Colalao, 23 January 1955, Singer T 2057 Parque General San Martin, 14 February
(LIL) 1939, Leal 6591 (BAFC)
subumbratilis Singer, 1970a: 23
HOLOTYPE: Argentina. Tucuman: upper part of PANAEOLUS
La Angostura, near Mollar, 29 January 1962,
convexulus Singer, 1965i: 161
Singer T 3766 (BAFC) HOLOTYPE: Chile. Moleros, 25 March 1959,
umbratilis var. depauperata Singer & Clemen-
Singer M 2025 (BAFC)
900, 1972b: 318, 319 moellerianus Singer, 1960J: 123
HOLOTYPE: Switzerland. Valais: River Borgne
HOLOTYPE: Australia. Macquarie Island: Half
de Ferpecle, 11 July 1971, Singer C 5460 (F) Moon Bay, 28 March 1950 (MICH)
OUDEMANSIELLA PANELLUS
echinosperma Singer, 1945d: 439 cystidiatus Singer in Singer & Digilio, 195 le:
HOLOTYPE: Brazil. Rio Grande do Sul: Sao 122
Leopoldo, Rick (FH). Deposited as Colly bia
HOLOTYPE: Argentina. Tucuman: Rio Cochu-
napipes. na, 20 February 1951, Singer s.n. (LIL) [pub-
lished in 1952]
glutinosa Singer, 1989a: 43
HOLOTYPE: Colombia. Valle: Cali, Buenaven- dumontii Singer, 1978h: 21
HOLOTYPE: Ecuador. Pichincha, 36 km from
hygrophoroides Singer & Clemencon, 1971b: Quito on the Quito-Tandayapa Rd., 6 August
124 1975, Dumont and others EC 2338 (NY)
HOLOTYPE: Switzerland. Vaud: Bois d' Apples, mirabilis Singer, 1955a: 770
10 July 1971, Clemen?on 710710 (LAU)
linicolor Grinling ex Singer, 1989a: 43 Highlands Hammock State Park, 12 August
HOLOTYPE: Zaire. Brazzaville: Fort de Man- 1942, Singer F 188 (F)
diel, 1968, Bouquet [Grinling no.] 71201 (F) nubigenus Singer, 1969a: 88
macracantha Singer, 1961b: 59 HOLOTYPE: Colombia. Boyaca: between Ra-
HOLOTYPE: Bolivia. Pando: Madre de Dios, 5 miriqui and Rondon, 30 July 1960, Singer B
3650 (BAFC)
April 1956, Singer B 2525 (LIL)
nubigenus ssp. australis Singer, 1969a: 89
subnigra Singer, 1989a: 44
HOLOTYPE: Brazil. Amazonas: Ponta Negra, HOLOTYPE: Argentina. Neuquen: Lago Cor-
18 July 1977, Singer B 10350 (INPA) rentoso, 25 April 1964, Singer 3529 M
(BAFC)
pteridophytorum Singer, 1973d: 31
PACHYLEPYRIUM Singer, 1957h: 321 HOLOTYPE: Mexico. Oaxaca: San Augustfn,
[published in 1958] 10 July 1969, Singer M 8437 (F)
Phaeomarasmius fulvidula Singer stenocystis Singer, 1973d: 32
Type species:
HOLOTYPE:
in Singer & Argentina. Jujuy: Lagunas de
Digilio T 4055
squarrulosum Singer, 1969a: 281

HOLOTYPE: Chile. Valdivia: Cordillera Pelada, PANUS
Mirador, 29 April 1967, Singer M 6550 (SGO) semirudis Singer, 1936f: 142
Type information: USSR.
PALAEOCEPHALA Singer, 1961b: 60
PAXILLUS
Type species: Marasmius cymatelloides Dennis
& Reid amazonicus Singer in Singer & Lodge, 1988i:
210
HOLOTYPE: Peru. Loreto: Rio Yarapa, SE of
PANAEOLOPSIS Singer, 1969a: 367 March 1986, Gerry Miller s. n. (F)
Iquitos,
Type species: Panaelopsis sanmartiniana Singer

boletinoides Singer in Singer & Digilio, 195 le:
431
brasiliensis Singer, 1976J: 62 HOLOTYPE: Argentina. Tierra del Fuego: Es-
HOLOTYPE: Rio de Janeiro: Cabo Frio,
Brazil. tancia Nueva Argentina, 14 February 1950,
near Canal, 16-17 October 1966, Jezek 28 (F) Singer M 264 (LIL) [published in 1952]
76 Chapter Three
PAXILLUS
boletinoides var. leucopus Singer, 1964c: 104 megalospora var.tetraspora Singer, 1987b: 259
HOLOTYPE: Argentina. Rio Negro: Nahuel HOLOTYPE: Amazonas: 30 km to Ma-
Brazil.
Huapf National Park, Rio Frias, 15 March naus, Embrapa, 14 June 1977, Singer B 9975
1959, Singer M
1808 (LIL) (INPA: 100937).
defibulatus Singer in Singer & Digilio, 195 le: oligoporpa Singer, 1987b: 250
434 HOLOTYPE: Costa Rica. San Jose: La Chonta,
HOLOTYPE: Argentina. Tierra del Fuego: Rio along Interamerican Highway, km 55, 1 1
July
Grande, Estancia Nueva Argentina, 11 Feb- 1982, Gomez 18196 (F)
ruary 1950, Singer M 181 (LIL) [published in quercetorum Singer, 1987b: 254
1952] HOLOTYPE: Costa Rica. San Jose: El Empal-
defibulatus var. unicolor Singer in Singer & Di- me, 23 July 1981, Singer B 12399 (F)
195 le: 436 subarduennensis Singer, 1987b: 252
gilio,
HOLOTYPE: Costa Rica. San Jose: Cerro de la
Grande, Estancia Nueva Argentina, 18 Feb- Muerte, 2 August 1981, Gomez & Singer
341 (LIL) [published in [Singer no.] B 12583a (F)
1952] subattenuata Singer, 1961b: 78

guttatus Singer, 1961H: 15
HOLOTYPE: Bolivia. Pando: Madre de Dios, 5
HOLOTYPE: Brazil. Pernambuco: Dois Irmaos, April 1957, Singer B 2510 (LIL)
5 July 1960, Singer B 3057 (LIL)
polychrous Singer, 1937: 226 PHAEODEPAS
HOLOTYPE: USSR. Western Caucasus (LE?)
nutans Singer, 1973d: 28
HOLOTYPE: Colombia. Valle: Mun. Cali, 15
PEGLEROMYCES Singer, 198 la: 500
April 1968, Singer B 6155 (F)
Type species: Pegleromyces collybioides Singer

PHAEOGYROPORUS Singer, 1944a: 360
collybioides Singer, 198 la: 500
HOLOTYPE: Brazil. Amazonas: 30 km N of Type species: Boletus braunii Bres.
Manaus, 2 January 1978, Singer B 10469 (F) beniensis Singer & Digilio, 1960i: 150
PHAEOCOLLYBIA Guayaramerfn, 6 March 1956, Singer B 1613
(LIL)
amazonica Singer, 1961b: 77
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 31 PHAEOMARASMIUS
attenuata ssp. mexicana Singer, 1957a: 367 ciliatus Singer, 1953a: 250
HOLOTYPE: Mexico. Dto. Federal, between HOLOTYPE: Argentina. Tierra del Fuego: Rio
San Pedro de Nexapa and Paso Cortes, 21 July Grande, Estancia Nueva Argentina, 15 Feb-
1957, Singer M
1586 (MICH) ruary 1950, Singer M 288 (LIL)
brasiliensis Araujo ex Singer, 1987b: 257 confragosuliformis Singer, 1969a: 299
HOLOTYPE: Brazil. Amazonas: Reserva Ducke, HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
7 January 1979, Singer B 11473 (INPA) Chiveria, 7 May 1967, Singer M 6773 (SCO)
(Type: 102712) confragosulus Singer, 1953b: 20
columbiana Singer, 1970a: 6 HOLOTYPE: Argentina. Tucuman: Rio Cochu-
HOLOTYPE: Colombia. Valle: Cali, Buenaven- na, 11 February 1952, Singer T 1826 (LIL)
tura, San Joaquin, 22 April 1968, Singer B fulvidula Singer in Singer & Digilio, 1951e:
6274 (F) 390
elaeophylla Singer, 1987b: 257 HOLOTYPE: Argentina. Tucuman: Tapia, 15
HOLOTYPE: Brazil. Amazonas: Reserva Ducke, April 1951, Singer T 1495 (LIL) [published in
26 May 1978, Singer B 11063 (INPA) 1952]
flava Araujo ex Singer, 1987b: 258 fuscomarginatus Singer, 1969a: 294

HOLOTYPE: Brazil. Amazonas: 30 km to Ma- HOLOTYPE: Chile. Colchagua: Pumanque, 14
naus, Embrapa, 14 June 1977, Singer B 9976 June 1967, Lazo PU-99 (SCO)
(INPA) gregarius Singer, 1959n: 586
megalospora Araujo ex Singer, 1987b: 259 HOLOTYPE: USA. Virginia: Giles County, Mt.
HOLOTYPE: Brazil. Amazonas: Road Manaus- Lake Biological Station, Singer V 302 (FH).
Caracaraf, km 45, 11 July 1977, Singer & Ar- Also see Schweiz. Zeitschr. Pilzk. 34 (4): 58,
aujo [Singer no.] B 10082 (INPA: 102698) 1956.

PHAEOMARASMIUS
limulatellus Singer, 1953a: 254 PHELLINTTES Singer & Archangelsky, 1957e: 41

Grande, Estancia Nueva Argentina, 22 Feb- Type species: Phellinites degiustoi Singer &
438 (LIL) Archangelsky
littoralis Singer, 1989a: 113
degiustoi Singer &
Archang., 1957e: 41
Type information: Argentina. Santa Cruz, 66
7 km NW of town, October 1982, Gomez
(LIL)
18682 (F)
malvacearum Singer in Singer & Digilio, 1951e: PHLEBOPUS
389
HOLOTYPE: Argentina. Tucuman: Sierra de brasiliensis Singer, 1983f: 43
San Javier, 31 July 1949, Singer T 604 (LIL) HOLOTYPE: Brazil. Amazonas, 14 April 1980,
[published in 1952] Araujo &
Singer [Singer no.] B 12098 (INPA)
mercedis Singer, 1989a: 114 viperinus Singer, 1947a: 5
HOLOTYPE: Colombia. Cundinamarca: Bojaca, Type information: Type in FH.
mucronosporus Singer, 1969a: 293 PHOLIOTA

Huapi National Park, between Laguna Frias agrocybiformis Singer, 1969a: 276
and Paso de las Nubes, 4 May 1964, Singer HOLOTYPE: Chile. Llanquihue: between En-
M 3955 (BAFC) senada and Rain, 18 May 1967, Singer M
myrceugenellae Singer, 1969a: 298 7598 (SGO)
May alnea Singer in Singer &
Digilio, 1951e: 336
1967, Singer M 6581
(SGO) HOLOTYPE: Argentina. Tucuman: Cerro Alto
nebularum Singer, 1961b: 75 del Taficillo, 4March 1951, Singer T 1398
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 28 (LIL) [published in 1952]
January 1956, Singer B 637 (LIL) aurantioalbida Singer, 1969a: 259
oligosporus Singer, 1989a: 114 HOLOTYPE: Argentina. Rio Negro: Laguna
HOLOTYPE: Brazil. Sao Paulo: Serra de Can- Frias, 21 March 1963, Singer M 3727 (BAFC)
December 1964, Singer B 4122
tareira, 17 (F) baeosperma Singer, 1953a: 217
pascuus Singer, 1989a: 115 HOLOTYPE: Argentina. Tierra del Fuego: Lago
HOLOTYPE: Uruguay. Maldonado, 27 May Fagnano, Cabercera del Lago, 19 February
1966, Singer B 4181 (BAFC) 1950, Singer M
368 (LIL)
rostratus Singer, 1953a: 252 blechni Singer, 1965i: 170
HOLOTYPE: Argentina. Tierra del Fuego: Rio HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
Grande, Estancia Nueva Argentina, 17 Feb- 28 March 1963, Singer M 3217 (BAFC)
318 (LIL) calendulina Singer in Singer & Digilio, 195 le:
spadiceospinulosus Singer, 1973d: 88 337
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 9 HOLOTYPE: Argentina. Tucuman: Santa Rosa,
February 1956, Singer B 1047 (BAFC) 17 December 1950, Singer T 1057 (LIL) [pub-
speireoides van chilensis Singer, 1969a: 295 lished in 1952]
HOLOTYPE: Chile. Llanquihue: Puella, 22 glutinigera Singer, 1959n: 585
March 1959, Singer M 2003 (BAFC) HOLOTYPE: Argentina. Tucuman: Taficillo, 23
subincarnatus van gilvus Singer, 1969a: 292 February 1955, Singer T 2217 (LIL)
HOLOTYPE: Argentina. Rio Negro: Laguna intermedia Singer, 1929b: 107
Frias, and Paso de las Nubes, 24 March 1963,
Type information: USSR. Caucasus: Kodor, Sak-
Singer M
3 170 (BAFC) en, September.
terrigenus Singer, 1973d: 88 irazuensis Singer, 1989a: 112
HOLOTYPE: Mexico. Veracruz: Jalapa, Tlilapa, HOLOTYPE: Costa Rica. Cartago: Reventado
8198 (F). Also pub- October 1982, Gomez 18580 (F)
Irazu,
lished in Singer, 1989a: 115.
kubickae Singer &
Clemen9on, 1971b: 126
HOLOTYPE: Czechoslovakia. Bohemia: near
PHAEOMYCENA Trgbon in Siidbohmen, 30 September 1970,
macrospora Singer in Singer & Digilio, 195 le: Kubicka & Singer [Singer no.] C 5228 (F)
176 maackiae Singer, 1946g: 142
HOLOTYPE: Argentina. Tucuman: Rio de los Type information: USSR. Khaborovsk Krai, 15
Sosas, 27 January 1951, Singer T 1141 (LIL) September 1939, Lyubarski (LE). [Published
[published in 1952] in 1948].
78 Chapter Three
PHOLIOTA
tnajalis Singer, 1969a: 269 altoandina Singer, 1989a: 105

HOLOTYPE: Chile. Valdivia: Cordillera Pelada, HOLOTYPE: Colombia. Risaralda: Valle de la
9 May 1965, Singer M 5535 (BAFC) Quebrada Betania, 25 January 1980, Boekhout
marthae Singer, 1969a: 261 361 (F)
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, australis Singer, 1969a: 214
Mirador, 6 May 1967, R. & M. Singer M 6734 HOLOTYPE: Argentina. Rio Negro: Laguna
(SCO) Frias, path to the Paso de las Nubes, 22 March
megalosperma Singer, 1953a: 216 1963, Singer M 3137 (BAFC)

HOLOTYPE: Argentina. Tierra del Fuego: Rio austrofilaris Singer, 1969a: 217
Grande, Estancia Nueva Argentina, 17 Feb- HOLOTYPE: Argentina. Neuquen: Rincon, 13
ruary 1950, Singer M 336 (LIL) March 1963, Singer M
3537 (BAFC)
microcarpa Singer, 1969a: 277 caricicola Singer, 1989a: 106
HOLOTYPE: Argentina. Rio Negro: Laguna HOLOTYPE: Colombia. Risaralda: Santa Rosa
Frias, path to Paso de las Nubes, 24 March de Cabal, 26 January 1980, Boekhout 370 (F)
1963, Singer M 3169 (BAFC) filaris var. ochracea Singer, 1950h: 429
montana Singer, 1965i: 169 Type information: USSR. Altai Mtns. (LE). No
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, Latin diagnosis.
28 March 1963, Singer M 3214 (BAFC) filaris var. recedens f. subochracea Singer, 1950H:
myxacioides Singer, 1969a: 268 430
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, Type information: USSR. Altai Mtns. (LE). No
Chiveria, 10 May 1965, Singer M 5707 (BAFC) Latin diagnosis.
naucorioides Singer, 1955a: 774 maireiaffinis Singer, 1989a: 106
HOLOTYPE: Brazil. Parana: Paranagua, 27 Sep- HOLOTYPE: USA. Illinois: Ogle County, 9 Oc-
tember 1952, Singer B 410 (F) tober 1976, Singer N 7591 (F)
novembris Singer, 1969a: 267 microspora Singer, 1989a: 107
HOLOTYPE: Argentina. Rio Negro: Laguna HOLOTYPE: USA. Illinois: Cook County, 16
Frias, path to the Paso de las Nubes, 19 No- October 1983, Singer N 5147 (F)
vember 1954, Singer M 4054 (BAFC) procera Singer, 1941e: 98
oedipus var. chilensis Singer, 1970d: 787 Type information: USSR. Altai Mtns.: Artubash,
HOLOTYPE: Chile. Colchagua: Pumanque, Singer &

Vasilieva (LE?) Also published in
1967, Lazo PU-90 (SCO) Singer, 1950a: 142, with more type informa-
tion: USSR. Altai Mtns.: Oirotia, Artubash, 17
psathyrelloides Singer, 1969a: 276
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- September 1937, Singer & Vasilieva A 1148
(LE).
pi National Park, Puerto Blest, Los Cantaros,
March 1959, Singer
15 1827 (BAFC)M ruizlealii Singer, 1969a: 216
HOLOTYPE: Argentina. Mendoza: Las Heras,
septembris Singer, 1969a: 275
HOLOTYPE: Chile. Llanquihue: Salto de Petro-
Sanchon de los Ciruelos, Leal 4.614 (BAFC).
hu, 17 September 1967, Singer M 7588 (SCO) Also in Ruiz Leal's herbarium.
septentrionalis ssp. vasilievae Singer, 1941e: 98
spumosa var. chrysocystidiophora Singer, 1969a:
274 Type information: USSR. Western Caucasus,
1936, Vasilieva (LE?)
HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
tucumana Singer, 1973d: 77
1967, Singer M 6716b (SCO)
spumosa var. crassitunica Singer, 1969a: 272
le, May 1949, Singer T 1199 (LIL). Depos-
1
May ited under P.

1967, Singer M 6712 (SCO) filaris.
verrucispora Singer, 1969a: 213

varzeae Signer, 1989a: 113
HOLOTYPE: Brazil. Amazonas: Parana do Jan-
zano, 18 April 1965, Singer 5048 (BAFC) M
auaca, 23 May 1980, Singer B 12177 (F)
PHYLLOBOLETELLUS Singer in Singer &

PHOLIOTINA Digilio, 1951e: 438 [published in 1952]
altaica Singer, 1941e: 98 Type species: Phylloboletellus chloephorus Singer
Type information: USSR. Altai Mtns., Singer & chloephorus Singer in Singer & Digilio, 1951e:
Vasilieva (LE). Also published in Singer, 438
1950a: 140, with more information: USSR. HOLOTYPE: Argentina. Tucuman, Rio de los
Altai Mtns.: Kurai, Singer & Vasilieva A 728, Sosas, 27 January 1951, T 1 126 (LIL) [pub-
(LE). lished in 1952]

PHYLLOBOLITES
PHYLLOBOLITES Singer, 1942b: 59 apoda Singer, 1955a: 769

HOLOTYPE: Panama. Zona Canalis: Rio Sar-
Type species: Paxillus miniatus Rick dinilla, 10 July 1952, Martin & Welden 7518
(F, IA)
PHYLLOPORUS cibaria Singer, 1963h: 207
HOLOTYPE: Peru. Cuzco: Urubamba, Chicon,
caballeroi Singer, 1973d: 101
no date, Zamalloa 201 (BAFC)
Yala, 14 February 1966, Singer T 5150 paradoxa Singer, 1969a: 55
(F)
HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
centroamericanus Singer & Gomez,
1984g: 169 M 6713 (SGO)
1967, Singer
HOLOTYPE: Costa Rica. San Jose: La Chonta,
pulcherrima Singer, 1978h: 12
September 1983, Gomez 18413 (F)
HOLOTYPE: Colombia. Cundinamarca: Haci-
fibulatus Singer, Ovrebo & Hailing, 1990c: 452
enda La Soche Granada, 23 July 1974, Du-
HOLOTYPE: Colombia. Antioquia, Along road
mont CO 1903 (NY)
from Santa Rosa de Osos to San Jose de la
Montana, 25 November 1986, Ovrebo 2546
(HUA). Isotype: NY. PLEUROCYBELLA Singer, 1947b: 81
gymnocystis Singer, 1989a: 124

Type species: Pleurotus porrigens Pers.: Fr.
Caracarai,km 125, 20 July 1973, Singer B
11310 (INPA) PLEUROFLAMMULA Singer, 1946d: 521
lariceti Singer, 1938d: 170
Type species: Crepidotus dussii Pat.
HOLOTYPE: USSR. Altai Mtns.: Lake Telez-
koje, September, Vasilieva & Singer (LE?) chocoruensis Singer, 1947g: 88
leucomycelinus Singer ex. Singer, 1978a: 426 HOLOTYPE: USA. New Hampshire: Mt. Cho-
HOLOTYPE: USA. Michigan: Warren Woods, corua, 21 September 1905, Krieger and Far-
23 July 1973, Ponce de Leon & Singer [Sing- low (FH). [Published in 1948].
er no.] N 7674 (F). Also see 1984g: 176 fluminensis Singer, 1973d: 86
manausensis Singer, 1978a: 424 HOLOTYPE: Brazil. Rio Grande do Sul: Gua-
HOLOTYPE: Amazonas: Manaus, Au-
Brazil. nabara, Corcovado, February-March, 1924,
gust 1977, Singer B 9790 (F) Murrill 246 (NY)
phaeoxanthus Singer & Gomez, 1984g: 171 majuscula Singer, 1969a: 282
HOLOTYPE: Costa Rica. San Jose: Jardin de HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May
Dota, 6 km W of Empalme, July 1983, Gomez 1967, Singer M 6729 (SGO)
& Alfaro 20583 (F) squarrulosa Singer, 1973d: 86
phaeoxanthus var. simplex Singer & Gomez, HOLOTYPE: Colombia. Valle: Mun. Cali, 2
1984g: 173 May 1968, Singer B 6915 (F)
HOLOTYPE: Costa Rica. San Jose: Empalme,
1.1 km E of "La Perla" restaurant., July 1983, PLEUROMYCENULA Singer, 1973d: 27
Gomez & Alfaro 20623 (F)
purpurellus Singer, 1973d: 100 Type species: Pleuromycenula ellipsoidea Sing-
HOLOTYPE: Colombia. Cauca: near Cali Po- er.
payan, 29 June 1968, Singer B 6775 (F)

circularis Singer, 1973d: 28
HOLOTYPE: Colombia. Valle: Rio Pance, 3
PHYSOCYSTIDIUM Singer, 1962b: 410
Type species: Collybia cinnamomea Dennis ellipsoidea Singer, 1973d: 27
HOLOTYPE: Mexico. Mexico: near Barranca de
PISTILLARIA Ameyalco, 16 July 1969, Singer MA 0007 (F)
flava Singer, 1978h: 42
saxegothaeae Singer, 1969a: 387 HOLOTYPE: Colombia. Valle: Cali, Buenaven-
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, tura, Calima, 18 April 1968, Singer B 6208
W slope, 30 March 1963, Singer M 3256 (F)
(BAFC)
PLEUROTELLUS
PLEUROCOLLYBIA Singer, 1947b: 80
fibulatus Singer, 1955a: 776
Type species: Collybia (Gymnopus) praemulti- HOLOTYPE: USA. Oregon: Benton County,
folia Murr. Mt. Alsea, 4 November 1939, Doty 723a (F)
80 Chapter Three
PLEUROTUS
PLEUROTUS aporpus Singer, 1954b: 122, 123

araucariicola Singer, 1953d: 141 pf National Park, Quetrihue, 16 May 1952,
HOLOTYPE: Brazil. Rio Grande do Sul: Taim- Singer M 675a (LIL)
besinho, 5 November 1951, Singer B 89 (LIL) aporpus f. porpophorus Singer, 196 la: 115, 130
citrinopileatus Singer, 1943a: 149 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
Type information: USSR. Khaborovsk Krai: pi National Park, 6 March 1959, Singer M
Khaborovsk, July-October. 1676 (LIL)
favoloides Singer, 1989a: 2 aquosus Singer, 1956c: 148
Brazil. HOLOTYPE: Argentina. Tucuman: East slope of
umazinho, 26 April 1978, Singer B 10990 Taficillo, 23 February 1955, Singer T 2208
(INPA) (LIL)
floridanus Singer, 1946g: 134 argentinensis Singer, 1958g: 230
HOLOTYPE: USA. Florida: Highlands County, HOLOTYPE: Argentina. Tucuman: Rio de los
Highlands Hammock State Park, 15 August Sosas, 4 February 1955, Singer T 2139 (LIL)
1942, Singer F 217 (FH). [Published in 1948]. atriavellaneus var. coquimbensis Singer, 1969a:
geesterani Singer, 1961b: 45 155
HOLOTYPE: South Africa. Cape: Humansdorp HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16
District, 2 December 1949, Maas Geesteranus
April 1967, Singer M 6509 (SGO)
6736 (L)
atriavellaneus var. flavidopubescens Singer,
lindquistii Singer, 1960b: 216
1961a: 123, 131
HOLOTYPE: Argentina. Buenos Aires: Ensen-
ada, Punta Lara, 21 April 1960, Lindquist &
to Domingo, 1
August 1958, Dennis 1756a
Singer S 194 (LIL)
(LIL, K)
microleucus Singer, 1978h: 6
atriavellaneus var. parvus Singer, 1958g: 248
HOLOTYPE: Morona Santiago, 7 km
Ecuador.
from Mendez on the Mendez-Macas Rd., 2
San Javier, Taficillo, 4 March 1951, Singer T
August 1975, Dumont and others [Dumont
1389 (LIL)
no.] EC
2219 (NY)
ostreatoroseus Singer, 1961h: 10 beniensis Singer, 1958g: 285
HOLOTYPE: Brazil. Pernambuco: Camaragibe, HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri-
14 July 1960, Singer B 3407 (LIL) beralta, 30 March 1956, Singer B 2346 (LIL)
sutherlandii Singer, 1952c: 211 burserae Singer, 1958g: 268
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio
Grande, Estancia Nueva Argentina, 10 Feb- Suapi, 9 February 1956, Singer B 1061 (LIL)
ruary 1950, Singer M 151 (LIL) cervinus var. minor Singer, 1947c: 228
Type information: Spain. Catalonia: Ribera de
Sant Maurici, 1934.
PLUTEUS &
cervinus var. tucumanensis Singer in Singer
Digilio, 1951e: 254
agriensis Singer, 1978h: 46
HOLOTYPE: Argentina. Tucuman: near Taff del
Valle, Quebradita, 14 December 1949, Singer
T 772 (LIL) [published in 1952]
albostipitatus var. poliobasis Singer, 1973d: 64
HOLOTYPE: Mexico. Veracruz: near Tuxtlas, chrysophlebius ssp. sublaevigatus Singer, 1958g:
21 June 1961, Singer M 8117 (F) 278
amazonaicus Singer ex Singer, 1989a: 93 HOLOTYPE: Bolivia. La Paz: Nor-Yungas,
HOLOTYPE: Brazil. Amazonas: 30 km N of Charobamba, 13 February 1956, Singer B
1147 (LIL)
(INPA) cinerellus Singer, 1956c: 156
amphicystis Singer, 1958g: 213

HOLOTYPE: Bolivia. Pando: Manuripi, Con- Highlands Hammock State Park, 28 August
quista, 26 March 1956, Singer B 2249 (LIL) 1942, Singer F 427 (?)
angustisporus Singer, 1958g: 206 circumscissus Singer, 1958g: 226

HOLOTYPE: Bolivia. Pando: Madre de Dios, HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Car-
Las Piedras, 5 April 1956, Singer B 2497 men Pampa, 17 February 1956, Singer B
(LIL) 12185 (LIL)

PLUTEUS
defibulatm Singer, 1952c: 222 globiger Singer in Singer & Digilio, 1951e: 266
HOLOTYPE: Argentina. Tierra del Fuego: Rio [published in 1952]
Grande, Estancia Nueva Argentina, 22 Feb- HOLOTYPE: Argentina. Tucuman: Sierra de
ruary 1950, Singer M 435 (LIL) San Javier, Anta Muerta, 16 April 1950, Sing-
dennisii Singer ex Singer, 1989a: 97 er T 973 (LIL) [Published in 1952]
HOLOTYPE: Trinidad. Non Sangre
Pareil, glutinosus Singer, 1989a: 94
Grande, 11 October 1949, Dennis 152 (K) HOLOTYPE: Panama. Panama: near the El Lla-
no Carti road, 29 June 1975, Dumont P 1512
diptychocystis Singer, 1954b: 123, 124
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- (F)
pf National Park, Isla Victoria, 17 May 1952, haywardii Singer, 1956c: 147
Singer M 725 (LIL) Type information: Argentina. Tucuman, T 452
dominicanus Singer, 196 la: 129, 131 (LIL)
HOLOTYPE: Venezuela. Merida: Sierra de San- hendersoniensis Singer, 1989b: 795
to Domingo, 1
August 1958, Dennis 1772 HOLOTYPE: Henderson Islands. (South Pacif-
(LIL, K) ic), Northwest Beach, Schuster 00006 (F)
eliae Singer, 1958g: 290 hiemalis Singer, 1958g: 248

HOLOTYPE: Bolivia. Pando: Manuripi, Con- HOLOTYPE: Argentina. Tucuman: Rio Cochu-
25 March 1956, Singer & E.R, de la na, Marta Grassi, 29 September 1956, Singer
quista,
Sota B 2193 (LIL)
T 2838 (LIL)
hololeucus Singer, 1956c: 200
espeletiae Singer, 196 la: 120, 130
HOLOTYPE: Venezuela. Merida: Sierra de San- HOLOTYPE: Argentina. Tucuman: Sierra de
San Javier, Ciudad Universitaria, 31 January
to Domingo, 23 July 1958, Dennis 1720 (LIL,
K)
eucryphiae Singer, 1969a: 153
hongoi Singer, 1989a: 95
HOLOTYPE: Japan. Kyoto: Mt. Daimonji-
W slope, 7 May 1965, Singer M 5567 (BAFC) yama, 14 September 1984, Singer, Hongo and
others [Singer no.] A 4070 (F)
exilis Singer, 1989a: 94
horridus Singer, 1973d: 62
HOLOTYPE: USA. California: Marin County,
HOLOTYPE: Mexico. Veracruz: near Montepio,
14 January 1983, Singer N 5105 (F)
exilis var. austriacus Singer, 1989a: 94
23 June 1969, Singer M 8177 (F)
HOLOTYPE: Austria. Burgenland: Siegendorf,
hylaeicola Singer, 1989a: 95
19 September 1979, Singer C 9457 (WU)
Singer B 4201 (BAFC)
fallax Singer, 1958g: 294
iguazuensis Singer, 1956c: 201
HOLOTYPE: Argentina. Misiones: Iguazu, 26
le, Parque Aconquija, 24 March 1949, Singer November &
1949, Singer Digilio (LIL)
T272 (LIL)
keissleri Singer, 1929b: 106
fastigiatus Singer in Singer & Digilio, 1951e:
Type information: USSR. Caucasus.
259 [published in 1952]
kuthanii Singer, 1989a: 96
HOLOTYPE: Argentina. Tucuman: Santa Rosa, HOLOTYPE: Czechoslovakia. Slovakia: Cadca,
17 December 1950, Singer T 1054 (LIL) 27 July 1974, Kuthan & Singer [Singer no.]
[Published in 1952] C 5696 (F)
fernandezianm Singer, 1958g: 220 laetifrons var. bolivianus Singer, 1958g: 277
HOLOTYPE: Chile. Juan Fernandez: Masatier-
ra, Quebrada Agujereada, 12 March 1955, roico, 4 February 1956, Singer B 889 (LIL)
Sparre F 32 (LIL, S) laetus Singer, 1958g: 274
fibulatus Singer in Singer & Digilio, 1951e: 252 HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ivon,
[published in 1952] 3 April 1956, Singer B 2483 (LIL)
HOLOTYPE: Argentina. Tucuman: Ciudad, Jar- laetus var. mixtus Singer, 196 la: 126, 131
din del InstituteLillo, 22 February 1949, HOLOTYPE: Venezuela. Dto. Federal: Caracas,
Singer T 141 (LIL) [Published in 1952] 3 July 1958, Dennis 1130 (LIL)
fluminensis Singer, 1958g: 292 leucocyanescens Singer, 1973d: 64
HOLOTYPE: Brazil. Rio de Janeiro: Angra dos HOLOTYPE: Mexico. Veracruz: near Montepio,
Reis, 31 September 1952, Singer B 432 (F) 21 June 1969, Singer M 8130 (F)
glaucus Singer, 196 la: 114, 130 maculosipes Singer, 196 la: 122, 131
HOLOTYPE: Chile. Llanquihue: Puella, 20 HOLOTYPE: Venezuela. Miranda: Cortada del
March 1959, Singer M 1892 (LIL) Guayabo, 9 July 1958, Dennis 1154 (LIL, K)
82 Chapter Three
PLUTEUS
major Singer, 1989a: 96 riberaltensis var. missionensis Singer, 196 la:

HOLOTYPE: USA. Illinois: Cook County, For- 123, 131
est Reserve E of Wheeling, 23 June 1985, HOLOTYPE: Argentina. Misiones: Iguazii, 29
Singer N 5195 (F) February 1960, Guerrero 130 (LIL)
melanopotamicus Singer, 1989a: 96 ricardii Singer, 1969a: 158
HOLOTYPE: Brazil. Amazonas: Igarape do Tar- HOLOTYPE: Chile. Concepcion: Boca del Bio-
umazinho, 13 March 1983, Singer B 12426 Bio, 11 November 1967, M 7819
Singer
(F) (SCO)
mesosporus Singer, 196 la: 117, 130 rimosellus Singer in Singer & Digilio, 195 le:
262
to Domingo, 31 July 1958, Dennis 1756 (LIL, HOLOTYPE: Argentina. Tucuman: Rio de los
K) T 1089
Sosas, 1
January 1951, Singer (LIL)
minor var. quercuum Singer, 1973d: 64
[Published in 1952]
HOLOTYPE: Colombia. Cauca: road between
Cali and Popayan, 28 April 1968, Singer B rimosoafftnis Singer, 1956c: 211
6770
HOLOTYPE: Argentina. Tucuman, 19 Novem-
(F)
ber 1954, Singer T 2011 (LIL, K)
neophlebophorus Singer, 1958g: 262
HOLOTYPE: Bolivia. El Beni: Vaca Diez, riograndensis Singer, 1953d: 116
HOLOTYPE: Brazil. Rio Grande do Sul: Sao
(LIL) Leopoldo, 31 October 1951, Singer B 31
(LIL)
nigropallescens Singer, 1961a: 116, 130
HOLOTYPE: Venezuela. Merida: Sierra de San- riograndensis var. atromarginatus Singer, 196 la:
to Domingo, August 1958, Buza
1 & Dennis 128, 131
[Dennis no.] 1781 (LIL, K) HOLOTYPE: Argentina. Tucuman: Tafi del Val-
1958g: 266 le, 20 July 1958, Singer T 3141 (LIL)
oligocystis Singer,
HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio rubrotomentosus Singer, 1958g: 228
Yariza, 23 February 1956, Singer B 1420 HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Rio
(LIL) Yariza, 23 February 1956, Singer B 1412
paraensis Singer, 1973d: 65 (LIL)
HOLOTYPE: Brazil. Para: Estancia Pirelli, 7
rugososulcatus Singer, 1958g: 250
June 1966, Singer B 4238 (BAFC) HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Ca-
paraensis var. mexicanus Singer, 1973d: 65 taratas de San Juan, 28 January 1956, Singer
HOLOTYPE: Mexico. Morelos: near Tepoztlan, B 609 (LIL)
16 February 1969, Singer M 8487 (F) salicinus var. achloes Singer, 1956c: 183
pluvialis Singer, 1958g: 234 HOLOTYPE: USA. Michigan: Cheboygan Coun-
Bolivia.
ty, Douglas Lake, 7 July 1953, Singer N 468
quista, Rio Madre de Dios, 25 March 1956, (F)
Singer B 2194 (LIL) sanctixaverii Singer, 1958g: 264
polycystis Singer, 1958g: 271 HOLOTYPE: Argentina. Tucuman: San Javier,
HOLOTYPE: Argentina. Misiones: Iguazii, Re- 13 January 1957, Singer T 2878 (LIL)
fugio Yaguarete, Parque Nacional Iguazii, 17
1958g: 290
1957, Singer M 1066 (LIL)
sapiicola Singer,
April HOLOTYPE: Argentina. Salta: Giiemes, Que-
pouzarianus Singer, 1983a: 283 brada de Yaquiasme, 7 February 1957, Singer
HOLOTYPE: Czechoslovakia. Bohemia: Tach-
T 2960 (LIL)
ov, near Rozvadov, 22 September 1967, Pou-
sergii Singer, 1958g: 253
zar PR 628956 (PR)
HOLOTYPE: Bolivia. Pando: Madre de Dios,
pulverulentus var. pseudonanus Singer, 1958g: Las Piedras, 5 April 1956, Singer B 2513
289
(LIL)
HOLOTYPE: Argentina. Salta: Burruyacu, 17
February 1957, Singer T 2986 (LIL). Also see
spegazzinianus Singer, 1952c: 221
Trans. Brit. Mycol. Soc. 39, fig.57, 1956 ( =
Singer, 1956c)
riberaltensis Singer, 1958g: 255 ruary 1950, Singer M
303 (LIL)
HOLOTYPE: Vaca Diez, Ri-
Bolivia. El Beni: stephanobasis Singer, 1958g: 212
beralta, 30 March 1956, Singer B 2336 (LIL) HOLOTYPE: Argentina. Tucuman: S. M. Tu-
riberaltensis var. conquistensis Singer, 1958g: cuman, 21 April 1956, Singer T 2802 (LIL)
255 subfibrillosus Singer, 1956c: 187
Bolivia. HOLOTYPE: Brazil. Rio de Janeiro: Angra dos
quista, 24 March 1956, Singer B 2177 (LIL) Reis, 30 September 1952, Singer B 433 (F)

PLUTEUS
subminutus Singer, 1958g: 269 epipolia Singer, 1989a: 7

HOLOTYPE: Bolivia. La Paz: Nor Yungas, Rio HOLOTYPE: Brazil. Amazonas: 30 km N of
Yariza, 23 February 1956, Singer B 1419 Manaus, 23 March 1978, Singer B 10850
(LIL) (INPA)
substigmaticus Singer, 1958g: 273 poliomphax Singer, 1989a: 8
HOLOTYPE: Bolivia. Pando: Manuripi, Con- HOLOTYPE: Brazil. Amazonas: 30 km N of
24 March 1956, Singer B 2183 (LIL)
quista, Manaus, 18 May 1977, Singer B 9658 (INPA)
sulcatus Singer in Singer &
Digilio, 1951e: 265
HOLOTYPE: Argentina. Tucuman: Taff del Val- POLYPLOCIUM
le, Quebradita, 15 January 1950, Singer T 876
patagonicum Singer, 1969a: 370
(LIL) [Published in 1952] HOLOTYPE: Argentina. Neuquen: San Ignacio,
triplocystis Singer, 1973d: 63 4 November 1966, Singer M
6005 (BAFC)
HOLOTYPE: Mexico. Veracruz: near Montepio,
22 June 1969, Singer M 8141 (F) POROMYCENA
tucumanus Singer in Singer & Digilio, 195 le:
269 anastomosans Singer, 1945g: 220
HOLOTYPE: Argentina. Tucuman: Rio de los Type information: USA. Florida: Highlands
Sosas, 18 January 1950, Singer T 921 (LIL) County, Highlands Hammock State Park,
[Published in 1952] Singer (FH)

umbrinoalbidus Singer, 1953d: 134
HOLOTYPE: Rio Grande de Sul: Estacio
Brazil.
PORPHYRELLUS
Sao Salvador, 9 November 1951, Singer B 124
festivus Singer, 196 In: 18
(LIL) HOLOTYPE: Brazil. Pernambuco: Dois Irmaos,
variipes Singer, 1956c: 218 13 July 1960, Singer B 3384 (LIL)
HOLOTYPE: Argentina. Tucuman: Quebrada de rionegrensis Singer &
Araujo, 1978a: 436
la Higuera, January, Singer T 2100 (LIL) HOLOTYPE: Amazonas: Along Rio Ne-
Brazil.
variipes var. atrofibrillosus Singer, 1958g: 266 gro, 20 km downstream from Sao Gabriel des
HOLOTYPE: Argentina. Tucuman: Quebrada de Cachoeiras, 20 January 1978, Araujo 937
la Higuera, 25 January 1955, Singer T 2099 (INPA)
(LIL) zaragozae Singer & Garcia, 199 la: 69, 88
varzeicola Singer, 1989a: 97 HOLOTYPE: Mexico. Nuevo Leon: Agua del
HOLOTYPE: Brazil. Amazonas: Ilha Marchan- Toro, Municipio Zaragoza, 1 1
September
taria, 9 March 1983, Singer and others [Sing- 1982, Garcia 2294 (F)
er no.] B 12397 (F)
viscidulus Singer in Singer & Digilio, 1951e: PORPOLOMA Singer, 1952c: 197, 198
255
HOLOTYPE: Type species: Porpoloma sejunctum Singer
Argentina. Tucuman: near Tapia,
1 January 1949, Singer T 797 (LIL) [Pub- portentosum Singer, 1952c: 199
lished in 1952] HOLOTYPE: Argentina. Tierra del Fuego: Rio
xanthopus Singer, 1958g: 282 Grande, Estancia Nueva Argentina, 10 Feb-

HOLOTYPE: Argentina. Tucuman: San Javier, ruary 1950, Singer M
149 (LIL)
28 January 1957, Singer T 2916 (LIL) sejunctum Singer, 1952c: 198
xylophilus var. major Singer, 1958g: 210
HOLOTYPE: Argentina. Tucuman: Anta Muer- Grande, Estancia Nueva Argentina, 13-18
ta, 24 November 1950, Singer T 1042 (LIL) February 1950, Singer M 256 (LIL)
terreum Singer, 1952c: 201
yungensis Singer, 1958g: 232
HOLOTYPE: Bolivia. La Paz: Nor- Yungas, Rio HOLOTYPE: Argentina. Tierra del Fuego: Us-
Llolosa, 31 January 1956, Singer B 777 (LIL)

huaia, Lago Fagnano, 19 February 1950, Sing-
er M 372 (LIL)
PODABRELLA Singer, 1945c: 143 PSATHYRA

Type species: Agaricus microcarpus Berk. & codinae Singer, 1947c: 230
Br. Type information: Spain. Catalonia: Riu de Bon-
aigua, July 1934.
albida Singer, 1989a: 7 muricellata Singer, 1938e: 13
HOLOTYPE: Costa Rica. Guanacaste: near Ba- Type information: USSR. Leningrad, 1936,
gaces, October 1982, L D Gomez 18718 (F) Singer (LE?)
84 Chapter Three
PSATHYRELLA
PSATHYRELLA marthae Singer, 1969a: 183

acutissima Singer, 1969a: 186 pi National Park, above Puerto Manzano, 1 1
HOLOTYPE: Argentina. Rio Negro: Laguna April 1963, M. & R. Singer M 3380 (BAFC)
Frias, Puerto Alegre, 13 March 1959, Singer metuloidophora Singer, 1973d: 72
M 1756 (BAFC) HOLOTYPE: Argentina. Tucuman: Sierra de
aequatoriae Singer, 1978h: 50 San Javier, 16 January 1960, Singer T 3538

HOLOTYPE: Ecuador. Tungurahua: Rio Verde, (LIL)
26 April 1973, Singer B 7129 (F) moseri Singer, 1969a: 187
annulosa Singer, 1973d: 70 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
HOLOTYPE: Brazil. Para: Belem, 5 June 1965, pf National Park, above Puerto Manzano, 1 1
Singer B 4192 (BAFC) April 1963, Singer 3379 (BAFC) M
apora Singer, 1969a: 192 nothomyrciae Singer, 1969a: 183
HOLOTYPE: Chile. Valdivia: Hueycolla, 4 May HOLOTYPE: Chile. Valparaiso: Quintero, 5
1967, Singer M 6702 (SCO) June 1967, Villagrdn [Singer no.] M 6944
argillospora Rick ex Singer, 1973d: 71 (SGO)
HOLOTYPE: Colombia. Cauca: In region "Ga- patagonica Singer, 1969a: 194
briel Lopez", 4 April 1968, Singer B 6992 (F) HOLOTYPE: Argentina. Neuquen: Puerto Man-
chiloensis Singer, 1969a: 197 zano, 5 April 1963, Singer M
3275 (BAFC)
HOLOTYPE: Chile. Chiloe: Pudeto, May 1967, phaeocystidiata Singer, 1973d: 73
M. Singer & Hernandez [Singer no.] M 6792 HOLOTYPE: Argentina. Jujuy: Lagunas de
(SCO) Yala, 14 February 1966, Singer T 5138 (F)
crinipellis Singer, 1973d: 71 polycystidiosa var. exumbonata Singer, 1969a:

HOLOTYPE: Mexico. Chiapas: 15 km toward 190
west of San Cristobal de la Casas, 2 August HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
1969, Singer M 8825 (F) pi National Park, Arroyo Frutillas, 21 April
dactylocystis Singer, 1969a: 188 1965, Singer M 5104 (BAFC)

HOLOTYPE: Argentina. Rio Negro: Cerro Otto, polycystidiosa var. minor Singer, 1969a: 190
17 November 1964, Singer M 4012 (BAFC) HOLOTYPE: Argentina. Neuquen: Quetrihue,
dactylocystis f.rugulosiceps Singer, 1969a: 189 1 1 November 1966, Singer M
6070 (BAFC)
HOLOTYPE: Argentina. Neuquen: Puerto Man- polycystidiosa var. polycystidiosa Singer, 1969a:
3282 A (BAFC) 190
dactylocystis f. latispora Singer, 1969a: 189 HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
HOLOTYPE: Argentina. Neuquen: Cerro Colo- Chiverfa, 22 April 1964, Singer M 3409
rado, 9-10 March 1959, Singer M 1731 (BAFC)
(BAFC) rhizophorae Singer, 1973d: 74
fascicularis Singer, 1978h: 51 HOLOTYPE: USA. Hawaii: Oahu, 4 June 1968,
HOLOTYPE: Ecuador. Tungurahua, between the Kohlmeyer 2564 (Kohlmeyer)
Ulva bridge and San Antonio, 29 April 1973, riparia Singer, 1973d: 74
Singer B 7186 (F) HOLOTYPE: Argentina. Corrientes: Curuzu
hesleriaffinis Singer, 1978h: 52 Cuatia, 8 February 1964, Singer S 422 (BAFC)
HOLOTYPE: Ecuador. Napo: Lago Agrio, 16 segregis Singer, 1989a: 102
May 1973, Singer B 7507 (F) HOLOTYPE: USA. Michigan: Berrien County,
janauariensis Singer, 1989a: 101 29 September 1973, Singer N 4500 (F)
HOLOTYPE: Brazil. Amazonas: Lago Janauari, sepuchralis Singer, Smith &
Guzman, 19581: 26
18 April 1983, Singer B 12447 (F) HOLOTYPE: Mexico. Oaxaca: near San Augus-
laevissima var. nothofagi Singer, 1969a: 197 tin Loxicha, 14 September 1957, Guzman
HOLOTYPE: Chile. Valdivia: Cuesta Santa El- SA-1140 (MICH)
vira, 15 May 1967, Singer M 6918 (SGO) squarrosiceps Singer, 1978H: 53
lignatilis Singer, 1989a: 101 HOLOTYPE: Ecuador. Napo: Lago Agrio, 8
Brazil. May 1973, Singer 7276 (F)
Caracaraf, km 45, 11 July 1977, Singer B subandina Singer, 1969a: 195
10076 (INPA) HOLOTYPE: Argentina. Neuquen: Puerto Man-
macquariensis Singer, 1959b: 392 zano, 5 April 1963, Singer 3278 (BAFC)M
HOLOTYPE: Australia. Macquarie Island: Tent subconocyboides Singer, 1978h: 54
Hill, 15 November 1951, Bunt MB/51/B6 HOLOTYPE: Ecuador. Napo: Lago Agrio, 9
(LIL) May 1973, Singer B 7320 (F)

PSATHYRELLA
subprona Singer, 1969a: 193 PSEUDOOMPHALINA

HOLOTYPE: Chile. Valdivia: Hueycolla, near
arsitophylli Singer, 1973d: 17
Ocean, May 1967, Singer M 6578 (SGO)
1
HOLOTYPE: Colombia. Boyaca: Paramo quer-
tilcariensis Singer, 1969a: 192
rero above Picho, 26 July 1960, Singer B 3530
HOLOTYPE: Argentina. Jujuy: Tilcara, Gargan-
(LIL)
ta del Diablo, 21 March 1955, Singer T 2312
(LIL)
PSILOBOLETINUS Singer, 1945b: 250
tristis Singer, 1969a: 197
HOLOTYPE: Argentina. Rio Negro: Nahuel Type species: Phylloporus lariceti Singer
Huapi National Park, Brazo Tristeza, 7 May
1964, Singer M 3978 (BAFC) PSILOCYBE
valdiviana Singer, 1969a: 185
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, aequatoriae Singer, 1978h: 59
varzeae Singer, 1989a: 102
HOLOTYPE: Brazil. Amazonas: Lago Janauari, aggericola Singer &
Smith, 1958k: 142
18 March 1983, Singer B 12430 (F)
HOLOTYPE: Argentina. Tucuman: road to Tafi
del Valle, Rio de los Sosas, 13 December
1949, Singer T 760 (MICH)
PSEUDOBAEOSPORA
alpestris Singer, 1989a: 108
defibulata Singer, 1963g: 13 HOLOTYPE: Austria. N. O.: Hoch-Schneeberg,
HOLOTYPE: Argentina. Tucuman: San Javier, 16 August 1979, Singer C 9168 (WU)
28 February 1962, Singer & Horak T 3775 araucana Singer, 1969a: 239
(LIL) HOLOTYPE: Argentina. Neuquen: Puerto Man-
flavescens Singer, 1969a: 172 zano, 22 April 1964, Singer M
3402 (BAFC,
HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16 SGO)
6525 (SGO) baeocystis Singer & Smith, 1958k: 141
syringea Singer, 1963g: 15 HOLOTYPE: USA. Oregon: Eugene, 1 Novem-
HOLOTYPE: Peru. Cuzco: Tambomachai, 19 ber 1945, Sipe (MICH)
May 1958, Singer B 3001 (LIL) caerulipes var. gastonii Singer, 1958d: 236
HOLOTYPE: Mexico. Oaxaca: Huautla de Jim-
PSEUDOCLITOCYBE nez, 13 July 1958, Guzman Huerta GH-1128
A (LIL). Isotype: Institute Politnico, Mexico
beschidica Singer & Kuthan, 1980a: 70 D.F.
HOLOTYPE: Czechoslovakia. Moravia: Besky-
caesioannulata Singer, 1965i: 165
dy, Nature preserve Minosi near Horni Lom-
HOLOTYPE: Chile. Valdivia: near Hueycolla,
20 July 1974, Kuthan &
na, Singer [Singer 20 March 1963, Singer M 3248 (BAFC)
no.] C 5592 (F)
candidipes Singer &
Smith, 1958k: 141
martipanis Singer, 1969a: 64 HOLOTYPE: Mexico. Oaxaca: Huautla, 10 July
Chiveria, 7 M 6758 (SGO) 1957, Singer M
1514 (MICH)
May 1967, Singer
carbonaria Singer, 1965i: 166
PSEUDOHIATULA 31 March 1963, Singer M 3267 (BAFC)
callistosporoides Singer, 19615: 59 chilensis Singer, 1965i: 168
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
San Javier, 7 T 3196
January 1959, Singer 30 March 1963, Singer M
3241 (BAFC)
(LIL) chrysocystidiata Singer, 1973d: 82
guzmaniana Singer, 1973d: 33 HOLOTYPE: Bolivia. El Beni: Vaca Diez, 8
HOLOTYPE: Mexico. Morelos, Parque Nacion- March 1956, Singer B 1747 (BAFC)
al de Lagunas de Zampoala, 3 July 1962, Guz- collybioides Singer & Smith, 1958k: 141
man 2978 (F) HOLOTYPE: Argentina. Tucuman: Tafi del Val-
haedinopsis Singer, 1978h: 44 le, 25 February 1952, Singer T 1882 (MICH)
HOLOTYPE: Ecuador. Tungurahua: Rio Topo, cordobensis Singer, 1973d: 83
30 April 1973, Singer B 7227 (F) HOLOTYPE: Argentina. Jujuy: Lagunas de
panamensis Singer, 1955a: 772 Yala, 13 February 1965, Singer T 4059 (F)
HOLOTYPE: Panama. Zona Canalis: Corund, februaria Singer, 1989a: 108
13 August 1952, Martin & Welden 8335 (F, HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 7
IA) February 1956, Singer B 976 (LIL)
86 Chapter Three
PSILOCYBE
flammuliformis Singer, 1969a: 247 sierrae Singer, 1969a: 240

HOLOTYPE: Chile. Valdivia: Cordillera Pelada, HOLOTYPE: Chile. Maule: between Chovellen
6 May 1965, Singer M 5523 (BAFC) and Trehualemu, 3 August 1967, Singer M
isauri Singer, 1958d: 237 7214 (SGO)
HOLOTYPE: Mexico. Oaxaca: Huautla de Jim- squarrosipes Singer, 1959n: 580
nez, 13 July 1958, Guzman Huerta GH-1128 HOLOTYPE: Argentina. Tucuman: Infiernillo,
(LIL). Isotype: Instituto Politnico, Mexico 10-11 March 1955, Singer T 2254 (LIL)
D.F. strictipes Singer &
Smith, 1958k: 141
jujuyensis Singer, 1973d: 82 HOLOTYPE: USA. Oregon: near Welches,
HOLOTYPE: Argentina. Cordoba: Pampa de Gruber & Smith [Smith no.] 202448 (MICH)
Achala, 31 December 1966, Singer B 4313a subhyperella Singer, 1973d: 83
(F) HOLOTYPE: Colombia. Cundinamarca: Salto
laetissima Hausknecht & Singer, 1986e: 295 de Tequendama, 24 July 1960, Singer B 3502
HOLOTYPE: Austria. Vienna: Lobau, 13 Sep- (F)
tember 1980, Hausknecht & Egle 917.0 (WU). valdiviensis Singer, 1969a: 251
Isotype: F HOLOTYPE: Chile. Valdivia: Cordillera Pelada,
lazoi Singer, 1969a: 242 7 May 1965, Singer M 5473 (BAFC)
HOLOTYPE: Chile. Colchagua: Pumanque, yungensis Singer & Smith, 1958k: 142
PU-151 (SCO)
1967, Lazo HOLOTYPE: La Paz: Nor-Yungas, Ca-
Bolivia.
longinqua Singer, 1959n: 578 taratas San Juan, 28 January 1956, Singer B
HOLOTYPE: Australia. Macquarie Island, 1
648 (MICH)
May 1949, Bunt (LIL)
yungensis var. diconica Singer & Smith, 1958k:
marthae Singer, 1969a: 255 142
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, HOLOTYPE: Bolivia. La Paz: Nor-Yungas, Ca-
7 May 1967, R. & M. Singer (SGO). No col- taratas San Juan, 28 January 1956, Singer B
lection number given. 651 (MICH)
mesospora Singer, 1969a: 245
21 April 1965, Singer 5128 (BAFC)M PULVEROBOLETUS
muliercula Singer &
Smith, 1958k: 142
duckeanus Singer, 1983f: 95
HOLOTYPE: Mexico. Piedras Blancas, 28 July
M
1611 (MICH)
HOLOTYPE: Brazil. Amazonas: Reserva
1957, Singer
Ducke, 6 km from entrance, 25 km N of Ma-
omniumsanctorum Singer, 1969a: 249
naus, 28 June 1978, Freire & Singer [Singer
HOLOTYPE: Chile. Llanquihue: Puella, Lago B 11247 (INPA)
Todos los Santos, 22 March 1959, Singer M no.]
1992 (LIL) leucomycelinus Singer & Ivory, 1983f: 93

HOLOTYPE: Honduras. Siguatepeque, 12 Oc-
peladae Singer, 1969a: 254 tober 1976, Ivory S/246 (F)
7 May 1967, Singer M 6782 (SGO) mazatecorum Singer, 1973d: 102
HOLOTYPE: Mexico. Oaxaca: San Andres Hi-
pericystis Singer, 1989a: 109 M 8361
HOLOTYPE: Brazil. Amazonas: 25 km N of dalgo, 9 July 1969, Singer (F)
Manaus, 28 June 1978, Singer B 11261 paspali Singer & Grinling, 1967c: 373
HOLOTYPE: Zaire. Brazzaville: Djoue, no date,
(INPA)
Grinling 4 1,1 04
(BAFC)
peruviana Singer, 1959n: 583
HOLOTYPE: Cuzco: rosaemariae Singer, 1983f: 99
Peru. Lamay, 21 May
1958, Singer B 3032 HOLOTYPE: Brazil. Amazonas: Laguna Jan-
(LIL)
auari, 31 December 1978, Singer B 11458
praetervisa Singer, 1969a: 244
HOLOTYPE: Argentina. Neuquen: Lago Cor- (INPA)
rentoso, 12 March 1963, Singer M 3506 shoreae Singer & Singh, 197 la: 28
HOLOTYPE: Dehra Dun,
India. Uttar Pradesh:
(BAFC)
20 July 1968, Bakshi 9022 (FRI, F) (FRI =
pteridophytorum Singer, 1959n: 582 Forest Research Institute, India)
Sosas, 4 February 1955, Singer T 2132 (LIL)
semilanceata var. microspora Singer, 1969a: PYRRHOGLOSSUM Singer, 1944a: 367
247
HOLOTYPE: Chile. 28 March 1953, Singer M Type species: Agaricus (Crepidotus) pyrrhys
3236 (BAFC). Also see Singer, 1965i. Berk. & Curt.

PYRRHOGLOSSUM
lilaceipes Singer, 1961b: 77 RESUPINATUS

February 1956, Singer B 1225 (LIL) alboniger var. bisporus Singer, 1978h: 19
macrosporum Singer, 1973d: 92 HOLOTYPE: Venezuela. Sucre: NW of Irapa,
HOLOTYPE: La Paz: Nor-Yungas,
Bolivia. 17 Trail N of Manacal, 9 July 1972, Dumont et

al VE 4510 (NY)
recedens Singer, 1973d: 91 chilensis Singer, 1946g: 134
HOLOTYPE: Argentina. Jujuy: Lagunas de Type information: Chile. Magallanes: Arenas,:
Yala, 14 February 1966, Singer T 5143 near Punta, Thaxter (FH). [Published in 1948].
(F)
stipitatum Singer, 1948c: 37 omphalioides Singer, 1965i: 150

HOLOTYPE: Guiana. Bartica, 1
January 1924,
Under 620 (FH) March 1963, Singer

31 3262 (BAFC) M
subrhacodium Singer in Singer & Digilio,
1951e: 104
RADHGERA HOLOTYPE: Argentina. Tucuman: Rio de los
Sosas, 10 March 1950, Singer T 936 (LIL)
paulensis Singer, Wright & Horak, 1963d: 600,
[Published in 1952]
610
HOLOTYPE: Brazil. Sao Paulo: "Cerradao" de
Itirapina,along road from Sao Paulo to Sao
RHODOARRHENIA Singer, 1964b: 142
Carlos, 16 May 1962, Milanez & Fidalgo s.n.
Type species: Merulinus pezizoides Speg.
(BAFC)
albocremea Singer, 1964b: 143
RAJAPA Singer, 1945c: 142
Type species: Agaricus eurhizus Berk. (LIL, BAFC)

nobilis Singer, 1964b: 144
RAMARIA men Pampa, 19 February 1956, Singer B 1274
(LIL, BAFC)
griseobrunnea Singer, 1983f: 19
June 1966, Singer B 4216 (BAFC) RHODOCYBE
inedulis Singer, 1969a: 384
alutacea Singer, 1946e: 688
March 1963, Singer
Frfas, 21 M 3106 (BAFC) theson Hammock, 29 September 1942, Singer
valdiviana Singer, 1969a: 385
F 908 (FH)
angustispora Singer, 1989a: 122
vira, 15 May 1967, Singer M 6916 (SCO)
RESINOMYCENA Redhead & Singer, 1981b: 12470-12478 (F)
151 australis Singer, 1969a: 335

Peck pf National Park, 28 April
Type species: Agaricus rhododendri Arroyo Frutillas,

acadiensis Redhead &
Singer, 19815: 162 collybioides Singer in Singer & Digilio, 1951e:
HOLOTYPE: Canada. New Brunswick: Kouchi- 425
bouguac National Park, 24 September 1977, HOLOTYPE: Argentina. Tucuman: Rio Lules, 2
J.E.& S.A. Redhead 2558 (166073, DAOM) April 1949, Singer T 366 (LIL) [Published in
brunnescens Redhead &
Singer, 1981b: 163 1952]
HOLOTYPE: Canada. New Brunswick: Kouchi- conica Singer, 1989a: 122
bouguac National Park, 13 July 1977, J.E. & HOLOTYPE: Brazil. Amazonas: Road Manaus-
S.A. Redhead 2336 (165884, DAOM) Caracarai, km 125, 21 May 1978, Singer B
montana Redhead &
Singer, 1981b: 157 11049 (INPA)
HOLOTYPE: Canada. British Columbia: Rocky crepidotoides Singer, 1973d: 95
Mountain trench, Donald Station, 21 Septem- HOLOTYPE: Brazil. Paraiba: Joao Pessoa, 11
ber 1980, Redhead 3977 (178215, DAOM) July 1960, Singer B 3333 (BAFC)
88 Chapter Three
RHODOCYBE
galerinoides Singer, 1961b: 81 crinipelloides Singer, 1978h: 75

HOLOTYPE: Bolivia. El Beni: Vaca Diez, 14 HOLOTYPE: Ecuador. Napo: Lago Agrio, 8
March 1956, Singer B 1973 (LIL) May 1973, Singer B 7305 (F)
gilvoides var. convexa Singer, 1958d: 242 cryptochrous Singer, 1969a: 338
HOLOTYPE: Mexico. Oaxaca: Huautla de Jim- HOLOTYPE: Argentina. Neuquen: Villa La An-
nez, 10 July 1957, Singer B 1511 (LIL) gostura, 29 April 1964, Singer 3646 M
hawaiiensis Singer, 1989a: 123 (BAFC)
HOLOTYPE: USA. Hawaii: Kauai, 24 January dissimilis var. aporpus Singer, 1969a: 343
1981, Singer H 1 (F) HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16
ianthinocystis Singer in Singer & Digilio, April 1967, Singer M
6526 (SGO)
1951e: 427 dissimilis var. dissimilis Singer, 1969a: 342
HOLOTYPE: Argentina. Tucuman: Rio Cochu-
HOLOTYPE: Argentina. Neuquen: Lago Cor-
na, 20 February 1951, Singer T 1286 (LIL) rentoso, 25 April 1964, Singer 3487M
[Published in 1952] (BAFC)
lactariiformis Singer, 1961b: 81 dragonosporus Singer, 1965e: 46
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 17 HOLOTYPE: Bolivia. El Beni: Vaca Diez, 17
March 1956, Singer B 2070 (LIL) March 1956, Singer B 2106 (BAFC, LIL)
laeta Singer, 1961b: 80 farlowii Singer, 1945e: 49
HOLOTYPE: Bolivia. El Beni: Vaca Diez, 14 HOLOTYPE: USA. New Hampshire: Chocorua,
March 1956, Singer B 1951 (LIL) July 1941, Under & Singer (FH)
lignicola Singer, 1961b: 80 ferrugineogranulatus Singer, 1973d: 96

HOLOTYPE: Bolivia. Yungas: La Paz, Nor- HOLOTYPE: Colombia. Valle: Cali, Buenaven-
Yungas, Carmen Pampa, 20 February 1956,
Singer B 1495 (LIL) fraternus Singer, 1973d: 97
marasmioides Singer in & HOLOTYPE: Brazil. Pernambuco: Dois Irmaos,
Singer Digilio,
5 August 1960, Singer B 3074 (BAFC)
195 le: 424
HOLOTYPE: hololeucus Singer, 1969a: 348
Argentina. Tucuman: near Tapia,
12 February 1951, Singer T 1192 HOLOTYPE: Chile. Llanquihue: 15 km NE of
(LIL) [Pub-
lished in 1952]
Puerto Pargua, 9 May 1967, Singer M 6800
& (SGO)
mycenoides Singer in Singer Digilio, 1951e:
423 lacteoniger Singer, 1969a: 345
HOLOTYPE: Argentina. Neuquen: Villa La An-
del Taficillo, 4 March T 1396 gostura, 25 April 1964, Singer 3645 M
1951, Singer
(BAFC)
(LIL) [Published in 1952]
naucoria Singer in Singer & 1951e: lowyi var. lowyi Singer, 1973d: 97
Digilio,
425
Singer and others [Singer no.] B 4271 (F)
HOLOTYPE: Argentina. Tucuman: Tapia, 15
T 1485 lowyi var. microspora Singer, 1973d: 98
April 1951, Singer (LIL) [Published
in 1952]
June 1966, Singer & Emigdio de Melo [Singer
mssularia Singer, 1961b: 80 B 4227 (BAFC)
no.]
lyophylliformis Singer, 1973d: 98
RHODOPHYLLUS mesites Singer, 1969a: 342
HOLOTYPE: Argentina. Neuquen: Rinc6n, 13
austroanatinus var. austroanatinus Singer, 1969a: March 1963, Singer M 3534 (BAFC)
337 nitens var. defibulatus Singer, 1969a: 344
HOLOTYPE: Argentina. Rio Negro: Laguna HOLOTYPE: Chile. Valdivia: Cuesta Santa El-
Frias, 21 March 1963, Singer M 3093 (BAFC) vira, 14 May 1967, Singer M 6892 (SGO)
austroanatinus var. defibulatus Singer, 1969a: nubigenus Singer, 1969a: 348
337 HOLOTYPE: Argentina. Rio Negro: Nahuel
HOLOTYPE: Chile. Coquimbo: Fray Jorge, 16 Huapi National Park, near Paso de las Nubes,
April 1967, Singer M 6533 (SGO) 24 March 1963, Singer M
3180 (BAFC)
cantharelluloides Singer, 1965e: 45 patagonicus Singer, 1969a: 339
HOLOTYPE: Brazil. Paraiba: Joao Pessoa, 11 HOLOTYPE: Argentina. Rio Negro: Laguna
July 1960, Singer B 3360 (BAFC) Frias, 21 March 1963, Singer M 3099 (BAFC)

RHODOPHYLLUS
perbrevisporus Singer, 1973d: 99 arcyospora Singer, 1990e: 31

HOLOTYPE: Brazil. Pard: Estancia Pirelli, 7 HOLOTYPE: Costa Rica. San Jose": Jardin de
June 1966, Singer B 4217 (F) Dota, 6 km W of Empalme, 6 August 1986,
porpoarachnoides Singer, 1978h: 76 Singer B 14628 (F)

HOLOTYPE: Ecuador. Tungurahua: Rio Topo, atropurpurea van atropurpurella Singer, 1932:
30 April 1973, Singer B 7204 (F) 298
pruinosipes Singer, 1969a: 340 Type information: Germany. Bavaria: Aubinger
HOLOTYPE: Argentina. Neuquen: Puerto Man- Lohe near Munich.
3272 (BAFC) atropurpurea van atropurpuroides Singer, 1932:
pumanquensis Singer, 1969a: 338 299
HOLOTYPE: Chile. Colchagua: near Pumanque, Type information: Germany. Bavaria.
7 August 1967, Lazo PU-131 (SGO) atropurpurea var. krombholzii Singer, 1932:
substrictior Singer, 19651: 184 301
HOLOTYPE: Chile. Valdivia: Cordillera Pelada, No type information.
28-29 March 1963, Singer M 3190 (BAFC) atropurpurea var. rubripes Singer, 1932: 300
tephrocybe Singer, 1978h: 78 Type information: Austria. Lower Austria: be-
HOLOTYPE: Ecuador. Tungurahua: Puyo, 26 tween Vslau and Merkenstein.
April 1973, Singer B 7140 (F) aucarum Singer, 1975b: 243
RIMBACHIA May 1973, Singer B 7440 (F)

austrodelica Singer, 1969a: 352
palmigena Singer, 1966a: 11 HOLOTYPE: Chile. Valdivia: Cuesta Santa El-
HOLOTYPE: Bolivia. Pando: Madre de Dios, 27 April 1967, Singer

vira, 6544 (SGO) M
Las Piedras, 5 April 1956, Singer B 2503 austromontana Singer, 1989a: 126
(LIL, BAFC) HOLOTYPE: Costa Rica. San Jose: El Empal-
me, 23 July 1981, Singer B 12402 (F)
barlae var. marthae Singer, 1982b: 679
RIPARTITELLA Singer, 1947b: 85
Type information: Spain. Catalonia: Vail d'
Marasmius squamosidiscus Murr. Aran, Seuva above Salardii, September 1934,
Type species:
Singer (LE)
batistae Singer, 1955d: 228
RIPARTITES HOLOTYPE: Brazil. Pernambuco: Iputinga, 7
March 1955, 2042 (LIL)
amparae Singer, 1950c: 195
boyacensis Singer, 1963b: 247
HOLOTYPE: Argentina. Tucuman: Sierra de HOLOTYPE: Colombia. Boyaca Tunja, Arca-
San Javier, Anta Muerta, 19 June 1949, Singer
buco, Finca del Sefior Luis Antonio Porras, 29
T 576 (LIL) B 3593 (BAFC)
July 1960, Singer
brasiliensis Singer, 1946g: 108
RUSSULA Type information: Brazil. Rio Grande do Sul,
Rick (FH: Herbarium Patouillard). [Published
albonigroides Singer, 1946g: 109 in 1948].
HOLOTYPE: Liberia. Ganta, 19 April 1939, caucaensis Singer, 1989a: 127
Harley 77 (FH). [Published in 1948]. HOLOTYPE: Colombia. Cauca: Los Araoz, 12
alutacea pseudoalutacea Singer, 1934a: 450
f.
April 1968, Singer B 6046 (F)
Type information: Austria. Lower Austria: near cernohorskyi Singer, 1935b: 300
Hintersdorf.
Type information: China. Yunnan: Li Jiang,
amnicola Singer, 1983f: 269 around village Nguluko.
HOLOTYPE: Brazil. Amazonas, 14 December chamaeleon Singer, 1937: 227
1978, Singer B 11438 (INPA). B 1438 was
1
Type information: USSR. Leningrad: Strelna,
cited as type in Latin diagnosis, but B 1 1452
July 1936 (LE?)
was cited as type in English description. citrinaf.
separata Singer, 1938b: 136
aosma Singer, 1952b: 298 Type information: USA. Washington: S of Lake
HOLOTYPE: Trinidad. Siparia, 3 November Crescent, 6 October 1935, Smith (MICH?)
1949, Dennis 281 (LIL) columbiana Singer, 1963b: 248
appalachiensis Singer, 1948c: 39 HOLOTYPE: Colombia. Cundinamarca: Salto
HOLOTYPE: USA. Virginia: Mountain Lake, 1 de Tequendama, 23 July 1960, Singer B 3498
August 1946, Singer V 169 (FH) (BAFC)
90 Chapter Three
RUSSULA
costaricensis Singer, 1983f: 267 fusca ssp. echinospora Singer, 1939e: 270
HOLOTYPE: Costa Rica. Guanacaste: Parque Type information: USA. Michigan: Ann Arbor,
Nacional Santa Rosa, 25 July 1981, Gomez & July.
Singer [Singer no.] B 12438 (F) gelatinascens Singer, 1989a: 129
crassotunicata Singer, 1938b: 132 HOLOTYPE: Czechoslovakia. Bohemia: Brad-
Type information: USA. Washington: Olympic lo, 10 September 1970, Singer C 5115 (F)
National Forest, Hot Springs, 29 September
1935, Smith (MICH) HOLOTYPE: Costa Rica. Alajuela: San Ramon,
cremeo-avellanea Singer, 1936e: 288 24 July 1981, Singer B 12429 (F)
No type information. altaica f. emeticicolor
gracilis ssp. Singer,
cristulispora Singer, 1941b: 84 1938b: 144
Type information: USSR. Western Caucasus: Type information: USSR. Altai Mtns.: Lake Te-
Krasnodar, 2 August 1936, Vasilieva (LE)
lezkoje, 4 September 1937, Singer (LE?)
cyanoxantha f. peltereaui Singer, 1925c: 15 gracilis ssp. Altaica.ca Singer, 1938b: 143
No type information.
Type information: USSR. Altai Mtns. (LE?)
cyanoxantha f. pallida Singer, 1923a: 4 guayarensis Singer, 1983f: 205
No type information. HOLOTYPE: Bolivia. El Beni: Vaca Diez,
dadmunii Singer, 1948c: 39
Guayaramerfn, 9 March 1956, Singer B 1761
Type information: USA. Massachusetts: Cam- (F)
bridge, Harvard, Dadmun (FH) handelii Singer, 1935b: 319
delica var. centroamericana Singer, 1989a: 128
Type information: China. Yunnan: Li Jiang, near
village Nguluko, September-October.
11 July 1982, Gomez 18191 (F)
heterochroa Singer, 1983f: 273
depigmentata Singer, 1989a: 128 HOLOTYPE: Brazil. Amazonas: Road Manaus-
HOLOTYPE: Spain. Catalonia: Tossa, 24 Octo- km 125, 22 May 1978, Singer
Caracarai, B
ber 1979, Singer C 9610 (F)
11012 (INPA)
emetica f. knauthii Singer, 1926: 216 hoehnelii Singer, 1946g: 110
No type information.
Type information: Indonesia: Java. Tjibodas,
emetica ssp. alnijorullensis Singer, 1950b: 133 Hohnel (FH). [Published in 1948].
Type information: Distributed in Argentina (Tu-
hongai Singer, 1989a: 130
cuman, Catamarca) and Bolivia [Published in HOLOTYPE: Japan. Kyoto: Mt. Daimonji, 14
31 December 1950]
September 1984, Singer A 4074 (F)
emetica ssp. lacustris Singer, 1950b: 133
humboldtii Singer, 1963b: 245
HOLOTYPE: U.S.A. Florida: Alachua County,
HOLOTYPE: Colombia. Cundinamarca: above
near Newnan's Lake, 28 December 1943,
Pacho, "Los Robles", 22 July 1960, Singer B
Singer F 1662 (FH) [Published in 31 Decem-
3482 (BAFC)
ber 1950]
idroboi Singer, 1963b: 245
epitheliosa Singer, 1983f: 203
km N HOLOTYPE: Colombia. Cundinamarca: above
HOLOTYPE: Brazil. Amazonas: 30 of
Pacho, at "Los Robles", 26 July 1960, Singer
Manaus, 12 July 1978, Freire & Singer [Sing-
er no.] B 11272 (F)
B 3523 (BAFC)
exalbicans f. decolorata Singer, 1938b: 147 leguminosarum Singer, 1983f: 227
HOLOTYPE: USSR. Kurai and Czuiskie Alpy,
Caracarai, km 45, 12 January 1979, Singer B
July- August 1937 (LE?)
11485 (INPA)
ferreri Singer, 1936e: 284
No leucomodesta Singer, 1957b: 178
type information.
428 HOLOTYPE: USA. Florida: Alachua County,
ferrotincta Singer, 1945d:
HOLOTYPE: USA. Newnan's Lake, 23 July 1943, Singer F 2916
Alachua County,
Florida:
1889a (FH)
Gainesville, Singer F (FH)
liberiensis Singer, 1946g: 112
foetens var. minor Singer, 1938b: 135
No type information Type information: Liberia. Nengbe, Harley 50
(FH?). [Published in 1948].
fontqueri Singer, 1947c: 215
Type information: Spain. Catalonia, Pyrenees
lutea f. maxima Singer, 1932: 250
Mtns., 1934. Type information: USSR. Caucasus.
lutea f. montana Singer, 1934a: 450
fuegiana Singer, 1950b: 125
HOLOTYPE: Argentina. Tierra del Fuego: Rio Type information: Austria. Nurnbergerhiitte.
Grande, Estancia Nueva Argentina, 13 Feb- lutea var. subcristulata Singer, 1938b: 156
254 (LIL). [Published Type information: USSR. Caucasus, Vasilieva
in 31 December 1950]. (LE?)

RUSSULA
macropoda Singer, 1939e: 277 nitida f. pseudoamethystina

Singer, 1932: 261
Type information: Type may be in LE. Type information: Germany-Italy.
maculata var. decipiens Singer, 1930: 211, 212 nitida var. transiens Singer, 1938b: 160
Type information: Czechoslovakia, near Prague, Type information: USSR. Tian Shan, 1
Septem-
1927, Holuinsky ber 1936, Nevodovski (LE?)
mairei Singer, 1929f: 308 nondorbingi Singer, 1958q: 275
Type information: Austria. (WU?) Type information: New Guinea. Binj, Christian
major Singer, 1954b: 153 &

Dennis (MICH)
HOLOTYPE: Argentina. Neuque"n: Nahuel Hua- nothofaginea Singer, 1950b: 127
pf National Park, Quetrihue, 13 May 1952, HOLOTYPE: Argentina. Tierra del Fuego: Rio
Singer M 521 (LIL) Grande, Estancia Nueva Argentina, 16 Feb-
mallophora Singer, 19505: 130 ruary 1950, Singer M

297 (LIL). [Published
HOLOTYPE: in 31 December 1950].
Argentina. Tierra del Fuego: Lago
Fagnano (Kamel), 19 February 1950, Singer obtecta Singer, 1939e: 233, 234
M 369 (LIL). [Published in 31 December Type information: USA. Tennessee: Great
1950]. Smoky Mountain National Park, Indian Cap,
mariae var. subflavida Singer, 1939e: 244 August, Smith
Type information: USA. Collected in 18 August oreina Singer, 1938b: 142

1931. Type information: USSR. Altai Mtns. (LE?)
mesospora Singer, 1938b: 161 oxydabilis f. aberrans Singer, 1938b: 161
Type information: USSR. Altai Mtns.: Lake Te- Type information: USSR. Altai Mtns.: Lake Te-
August (LE?)
letzkoje, lezkoje, September 1937, Singer (LE?)
metachromatica Singer, 1952b: 297 pachycystis Singer, 1983f: 236

HOLOTYPE: Venezuela. Aragua: Mt. Periquito, HOLOTYPE: Brazil. Amazonas: Road Manaus-
Rancho Grande, Maracay, 19 November 1949, Caracaraf, km 45, 25 June 1980, Singer B
Dennis 407 (LIL) 12123 (INPA)
metachromatica ssp. tarumaensis Singer, 1983f: pantoleuca Singer, 1957b: 239

HOLOTYPE: USA. Massachusetts.: Woburn, 18
245
Brazil.
September 1943, Singer (FH)
umazinho, 16 April 1980, Singer B 12105 patouillardii Singer, 1935c: 89
(INPA)
Type information: Madagascar. Maromandia
(PC)
mlcrospora Singer, 1939e: 262
peckii Singer, 1943c: 147
Type information: USA. Michigan: Whitemore
Type information: USA. New York: North Elba
Lake, 30 July 1923, Smith
or Piseco.
minutula var. robusta Saini, Atri & Singer,
persobria Singer, 1939e: 228, 229
1982c: 239
Type information: USA. Oregon, October, Kauff-
HOLOTYPE: India. Himachal Pradesh: Simla, man (MICH?)
Summer Hill, 30 July 1979, Atri PUN 274
piceetorum Singer, 1963f: 294, 301
(PUN). Isotype: F HOLOTYPE: Austria. Tyrol: St. Leonhard im
mitissima Singer, 1989a: 130 22 September 1960, Singer C 3006
Pitztal,
HOLOTYPE:Costa Rica. Alajuela: San Ramon,
(BAFC)
24 July 1981, Singer B 12428 (F)
pluteoides Singer, 1989a: 130
montivaga Singer, 1950b: 131 HOLOTYPE: Costa Rica. Alajuela: San Ramon,
Type information: Distributed in Argentina (Ca- 26 July 1981, Singer B 12500 (F)
tamarca, Jujuy) and Bolivia. [Published in 31
pluvialis Singer, 1983f: 221
December 1950]. HOLOTYPE: Brazil. Amazonas: Manaus, 5 June
nanella Singer, 1983f: 271 1977, Singer B 9933 (INPA)
polycystis Singer, 1939e: 238, 239
Caracarai, km 106, 4 April 1978, Singer B
Type information: USA. Tennessee: Great
10923 (INPA) Mt. Le
Smoky Mountain National Park,
neodiscopoda Singer, 1983f: 211 Conte, 23 August.
HOLOTYPE: Bolivia. El Beni: Vaca Diez, psammophila Singer, 1989a: 131
Guayaramerin, 14 March 1956, Singer B 1943 HOLOTYPE: USA. Illinois: Lake County, 1 Au-
(F) gust 1982, Singer N 5046 (F)
nitida var. oirotica Singer, 1938b: 160 pseudolepida Singer, 1939e: 251
Type information: USSR. Altai Mtns.: Teletzkoe, Type information: USA. Maryland: Takoma, 8
9 September 1937, Singer (LE?) August 1919, Kauffman
92 Chapter Three
RUSSULA
puellaria var. caucasica Singer, 1932: 274 sphagnophila var. subingrata Singer, 1930: 209,
Type information: USSR. Caucasus. 212
pyrenaica Blum ex Singer, 1982b: 686, 687 Type information: Austria or Czechoslovakia,
HOLOTYPE: Spain. Catalonia: La Salut, 25 Oc- 1925.
tober 1979, Singer C 9621 (F) steinbachii Cernohorsky &
Singer, 1934a: 457
"This taxon is established by a watercolor in Vi-
quercusoleoidis Singer, 1983f: 255
HOLOTYPE: Costa Rica. Guanacaste: Parque enna Natural History Museum."
Nacional Santa Rosa, 25 July 1981, Gomez & subminutula Singer, 1947c: 237
Singer [Singer no.] B 12439-19439a (F) Type information: Spain. Catalonia: Riu de Pe-
tianschanica 1938b: guera, July 1934.
rhodopoda ssp. Singer,
147 subtorulosa Singer, 1982b: 684, 685
HOLOTYPE: Switzerland. Bern: Gstaad, 17 Au-
Type information: USSR. Kazakhstan: near
gust 1970, Singer C 5085 (F)
Ata, 24 July 1937, Kuznetzova & Nev-
Alma
odovski (LE?) subvelata Singer, 1929b: 90
rosacea var. americana Singer, 1938b: 146 Type information: USSR. Caucasus.
tennesseensis Singer, 1939e: 229, 230
Type information: USA. Washington: Olympic
Type information: USA. Tennessee: Great
National Forest, Hot Springs, 23 September
Smoky Mountain Natl Park, Cades Cove, Au-
1935, Smith (MICH?)
gust, Smith (MICH?)
roseipes ssp. dictyospora Singer, 1938b: 155
variegata Singer, 1957b: 186
Type information: USSR. Western Siberia: Altai HOLOTYPE: USA. Florida: Alachua County,
Mtns., Kebesen, July- September, Singer (LE?)
Sugarfoot Hammock, 4 July 1943, Singer F
roseipes ssp. thermophila Singer, 1957b: 257 2544 (FH)
HOLOTYPE: USA. Florida: Alachua County,
venezueliana Singer, 1952b: 295
Gainesville, 10 July 1943, Singer F 2682 (FH) HOLOTYPE: Venezuela. Dto. Federal: Caracas,
schiffheri Singer, 1929b: 88 18 November 1949, Dennis 408 (LIL)
Type information: USSR. Caucasus. verna Singer, 1983f: 216
semililacea Singer, 1989a: 132 HOLOTYPE: Brazil. Amazonas: 30 km N of
HOLOTYPE: Colombia. Cundinamarca: near Manaus, 3 June 1977, St. John & Singer
Boyaca, 8 April 1968, Singer B 6013 (F) [Singer no.] B 9918 (INPA)
septentrionalis Singer, 1936f: 151 vinosa ssp. occidentalis Singer, 1946g: 114
Type information: USSR. Umgebung des Dor- No type information. [Published in 1948].
fes: Schtscheljagur, Flussgebiet der Petschora, \vahgiensis Singer, 1960J: 126
23 August 1926, Zinserling HOLOTYPE: New Guinea. Admin. Australia:
serotina f. minor Singer, 1941b: 84 Rio Wahgi Valle, Dwyer (MICH)
Type information: USSR. Western Caucasus: xerampelina var. semirubra Singer, 1957b: 218,
Krasnodar, 27 July 1936, Vasilieva (LE?) 219
smaragdina var. innocua Singer, 1935b: 304 HOLOTYPE: USA. Massachusetts, 31 August
Type information: Austria. Burgenland: Wiener- 1945, Roxbury, Singer & Under s. n. (FH)
wald, August.
smithii Singer, 1938b: 140 SCUTIGER
Type information: USA. Washington: Olympic
National Forest, 14 September 1935, Smith brasiliensis Singer, 1983f: 22
(MICH?) HOLOTYPE: Brazil. Amazonas: Road Manaus-
sororiicolor Singer, 1983f: 250 Caracaraf, km 44, Estacao de Silvicultura

Brazil. Tropical, 6 January 1980, Webber 139 (INPA)
(INPA) SCYTINOPOGON Singer, 1945c: 139
sphagnophila var. heterosperma Singer, 1938b:
150 Type species: P ferula pallescens Bres.
Type information: USSR. Siberia: Oirotia, near
lake Teletzkoje, August, Singer (LE?)
SETCHELLIOGASTER
sphagnophila var. subheterosperma Singer,
1938b: 150 tetrasporum Singer, 197 Id: 467
Type information: USSR. Altai Mtns.: Lake Te- HOLOTYPE: Chile. Malleco: Cordillera Na-
letzkoje, September, Singer (LE?) huelbuta, 16 May 1967, Singer M 6923 (SGO)

SIMOCYBE
SIMOCYBE tucumana Singer, 1961b: 72

aestivalis Singer, 1973a: 49, 516 San Javier, 26 February 1961, Singer T 3661
HOLOTYPE: Mexico. Veracruz: 7 km S of (LIL)
Montepfo, Estacion Biologica de las Tuxtlas, urosphaera Singer, 1969a: 289

8080 (F) HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
albovelutina Singer, 1989a: 119 pf National Park, Arroyo Frutillas, 28 April
HOLOTYPE: Venezuela. Monagas: Caripe, 18 1964, Singer M 3624 (BAFC)
July 1972, Dumont and others [Dumont no.]
VE 5310 (NY) SKEPPERIELLA
alneti Singer, 1989a: 120
populi Singer, 1963e: 210
HOLOTYPE: Argentina. Jujuy: Lagunas de HOLOTYPE: Argentina. Buenos Aires: Parque
Yala, 10 February 1966, Singer T 5024 (F) Pereyra, 20 June 1962, Singer S 200 (BAFC)
amazonica Singer, 1973a: 513, 516
HOLOTYPE: Bolivia. El Beni: Vaca Diez, Ri- SMITHIOMYCES Singer, 1944a: 366
beralta, 29 March 1956, Singer B 2319 (LIL?)
Latin diagnosis says the type at F, but the type Type species: Leucomyces mexicanus Murr.
cannot be located at F.
coniferarum Singer, 1989a: 120

STIGMATOLEMMA
HOLOTYPE: Switzerland. Vaud: Bois d' Apples,
hyalinum Singer, 1989a: 43
5 August 1970, Singer C 5040 (F) HOLOTYPE: Brazil. Amazonas: 30 km N of
coniferarum quercuum Singer, 1989a: 120
var. Manaus, 30 January 1978, Singer B 10565
HOLOTYPE: USA. Illinois: White Pine State (INPA)
Park, 1 September 1975, Singer N 7541 (F)
coroicensis Singer, 1973a: 507, 516 STROBILOMYCES
B 999 confusus Singer, 1945a: 108
roico, 8 February 1956, Singer (F)
corticalis Singer, 1978h: 69
Flatwood, east of Gainesville, July 1943,
HOLOTYPE: Ecuador. Napo: Lago Agrio, 13 F
2782 (FH)
Singer
pauper Singer, 1983f: 126
junquillea Singer, 1961b: 72 HOLOTYPE: Brazil. Amazonas: Road Manaus-
HOLOTYPE: Argentina. Tucuman: Sierra de Caracaraf, km 112, 4 April 1978, Singer B
San Javier, 17 February 1955, Singer T 2176 10915 (INPA)
(LIL)
pterosporus Singer, 1945a: 114
magna Singer, 1978h: 70 HOLOTYPE: Liberia. Du River, Firestone # 3,
HOLOTYPE: Ecuador. Napo: Lago Agrio, 8 Under 312 (FH)
microlepis Singer, 1978h: 71 STROBILURUS Singer, 1962b: 409
HOLOTYPE: Ecuador. Pichincha: Santo Domin-
Type species: Agaricus conigenoides Ellis
go de los Colorados, 21 April 1973, Singer B
7089 (F) esculentus var. montezumae Singer, 1973d: 32
ovalis Singer, 1961b: 73 HOLOTYPE: Mexico. Rio Frio, 13 July 1969,
HOLOTYPE: Venezuela. Horto Bogotensi, no Singer s.n. (F)
Dennis 1072 (LIL, K)

date,
panamensis Singer, 1989a: 121 STROPHARIA

HOLOTYPE: Panama. Panama: Veraguas, 18
&
aeruginosa var. neuquenensis Moser Singer,
June 1975, Dumont PA 560 (NY)
1969a: 229
peullensis Singer, 1969a: 290 HOLOTYPE: Argentina. Neuquen: Rinc6n, 13
HOLOTYPE: Chile. Llanquihue": Puella, 25 March 1963, Singer & Moser [Singer no.] M
March 1963, Singer M 3189 (BAFC) 3520 (BAFC)
polytropoides Singer, 1961b: 74 andina Singer, 1978h: 60
HOLOTYPE: Argentina. Tucumdn: Playa Gran- HOLOTYPE: Ecuador. Cotopaxi: Paramo de
de, 24 January 1951, Singer T 1120 (LIL) Cotopaxi, 20 May 1973, Singer B 7085 (F)
reductoaffinis Singer, 1961b: 71 flocculosipes Singer, 1969a: 230
HOLOTYPE: USA. Florida: Dade County, 11 HOLOTYPE: Chile. Santiago: Cerro Robles, 19
December 1942, Singer F 1618 (F) June 1967, Singer M 6975 (SGO)
94 Chapter Three
SUILLUS
SUILLUS dombeyi Singer, 1960d: 385

brevipes var. aestivalis Singer, 19455: 267 Huapi National Park, Arroyo Blest, 17 March
Type information: USA. Florida: Alachua Coun- 1959, Singer M
1861 (LIL)
Gainesville.
ty,
fragile var. cortinariopsis Singer, 1969a: 364
chiapasensis Singer, 1973d: 99 HOLOTYPE: Chile. Llanquihue: Salto de Petro-
HOLOTYPE: Mexico. Chiapas: 5 km W of San hue, 1 1
May 1967, Singer M 6838 (SGO)
Cristobal de las Casas, 2 August 1969, Singer
magellanicum Singer, 1951c: 219
M 8908 (F) HOLOTYPE: Argentina. Tierra del Fuego: Rio
cothurnatus ssp. aestivalis Singer, 1945b: 261 Grande, Estancia Nueva Argentina, Lago Fag-
HOLOTYPE: USA. Florida: Highlands County, nano, 10 February 1950, Singer M
515 (LIL)
Highlands Hammock State Park, 2 September subalbidum Singer, 19631: 24
1942, Singer F 478 (FH) HOLOTYPE: Chile. Magallanes: Punta Arenas,
cothurnatus ssp. hiemalis Singer, 1945b: 261, 5 March 1906, Thaxter, Fungi Hypogeus 4
263 (FH)
tylostomatoides Singer, 1969a: 365
Gainesville, December 1942-January 1943, HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
Singer F 1666 (FH)
pi National Park, above Puerto Manzano, 26
granulatus ssp. snellii Singer, 1945e: 40 April 1964, Singer M
3576 (BAFC)
Type information: USA. New Hampshire: Cho- violaceum Singer, 1951c: 216
corua, Linder & Singer (FH) HOLOTYPE: Argentina. Tierra del Fuego: Lago
hirtellus ssp. cheimonophilus Singer, 1945b: Fagnano (Kamel), 19 February 1950, Singer
274 M 362 (LIL)
ty, 30 January 1943, Singer F 1772? (FH) TRICHOLOMA
hirtellus ssp. thermophilus Singer, 1945b: 275
HOLOTYPE: USA. Florida: Highlands County, altaicum Singer, 1943a: 72
Highlands Hammock State Park, August-Sep- Type information: USSR. Altai Mtns. Central:
tember 1942, Singer F 543 (FH) Alpes Czuicae, August.

lithocarpisequoiae Singer, 1959n: 589 cortinatellum Singer, 1954b: 107, 108
HOLOTYPE: USA. California: Marin County, HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
Muir Woods, 30 January 1958, Singer N 1531 pi National Park, Quetrihue, 15 May 1952,
(LIL) Singer M 636 (LIL)
nueschii var. caerulescens Singer, 1961b: 82 cortinatum Singer, 1952c: 192
HOLOTYPE: USSR. Kurai, 1937, Singer & HOLOTYPE: Argentina. Tierra del Fuego: Rio
Vasilieva s.n. (LE) Grande, Estancia Nueva Argentina, 22 Feb-
ruber Singer & Sipe, 1946g: 103 ruary 1950, Singer M
433 (LIL)
HOLOTYPE: USA. Oregon: Odell Creek, Sipe diemii Singer, 1954b: 108
333 (FH). [Published in 1948]. HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
pi National Park, Quetrihue, 14 May 1952,
TECTELLA Singer M
600 (LIL)
elvirae Singer, 1969a: 38
phellodendri Singer, 1943a: 156

Type information: USSR. Khaborovski Krai. 27 April 1967, Singer

vira, M 6545 (SGO)
fagnani Singer, 1952c: 194
HOLOTYPE: Argentina. Tierra del Fuego: Lago
THAXTEROGASTER Singer, 1951c: 216
366 M
Fagnano, 19 February 1950, Singer
(LIL)
Type species: Thaxterogaster magellanicum
Singer.
fusisporum Singer, 1943a: 71
Type information: USSR. Oirotia: Altai Mtns.
bibulum Singer, 1969a: 361 Central, July-August.

HOLOTYPE: Argentina. Rio Negro: Laguna glaucoalbum Singer, 1948c: 28
Frias and Pas de las Nubes, 3-4 May 1964, HOLOTYPE: USA. Virginia: Mountain Lake,
Singer M 3951 (BAFC) Wilson V-128 (FH)
brevisporus Singer, 1960d: 386 moseri Singer, 1973d: 17

HOLOTYPE: Argentina. Rio Negro: Nahuel HOLOTYPE: Mexico. Dto. Federal, Paso de
Huapi National Park, Los Cantaros, 1 5 March Cortes, 23 July 1969, Singer M
8521 (F). Also
1959, Singer M
1817 (LIL) published in 1989a: 11.

TRICHOLOMA
mostnyae Singer, 1969a: 38 strophosa Singer, 1969a: 319

HOLOTYPE: Chile. Valdivia: Cuesta Santa El- HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
vira, 14 May 1967, Singer M 6883 (SGO) pi National Park, Arroyo Frutillas, 28 April
ochraceorobustum var. cautinense Singer, 1969a: 1964, Singer M
3639 (BAFC)
37 thermophila Singer, 1946g: 145
HOLOTYPE: Chile. Cautin: Pidenco, 26 April HOLOTYPE: USA. Florida: Highlands County,
1967, Singer M 6540b (SGO) Highlands Hammock State Park, 2 August
patagonicum Singer, 1954b: 109 1942, Singer F 20, F 20a (FH). [Published in
HOLOTYPE: Argentina. Neuquen: Nahuel Hua- 1946].

Singer M 684 (LIL) TYLOPILUS
pseudosordidum Singer, 1945d: 434
HOLOTYPE: USA. Florida: Dade County, Mi- acutesquamosus Singer, 1983f: 117
ami, 29 September 1942, Singer F 900 (FH) HOLOTYPE: Brazil. Amazonas: Road Manaus-
Caracaraf, km 45, 12 January 1979, Singer B
TRICHOLOMOPSIS 11483 (INPA)
arenarius Singer, 1978a: 432
araucariae Singer, 1953d: 119
HOLOTYPE: Brazil. Parecf, 2 November 1951,
Caracarai, km 45, 3 February 1978, Singer B
Singer B 50 (LIL) 10590 (INPA)
elatipes Singer, 1989a: 10 &
brachypus Singer, Garcia Gomez, 199 la: 47,
89
Manaus, 23 June 1977, Singer & Araujo
B 10006 (INPA) HOLOTYPE: Mexico. Durango: between Potre-
[Singer no.]
ro Mesa Larga and Arroyo Presa Los Altares,
elegans var. minor Singer, 1973d: 13
in Reserva de la Biosfera de la Michilia, 4
HOLOTYPE: Brazil. Para: near Belem, 5 June
B 4196 (BAFC) September 1983, Valenzuela 2470 (F). Isotype
1966, Singer
in ENCB
humboldtii Singer, Ovrebo & Hailing, 1990c:
cellulosus Singer, 1947a: 105
456
Type information: Liberia. Nengbe, Harley 60
HOLOTYPE: Colombia. Antioquia: San Jose de
(FH)
laMontana, Along road from San Jose de la
corneri Singer, 199 la: 50, 89
Montana to Labores, 26 November 1986,
Ovrebo 2554 (HUA). Isotype: CSU and NY.
HOLOTYPE: Costa Rica. San Jose: 6 km of W
Empalme, 2 July 1983, Gomez & Alfaro
tricholomatoides Singer, 1989a: 1 1
20582 (F)
HOLOTYPE: Brazil. Amazonas: Ilha Janauari,
18 March 1983, Singer B 12429 (F)
costaricensis Singer &
Gomez, 199 la: 36, 89
HOLOTYPE: Costa Rica. San Jose: 1.1 km E of
"La Perla" restraurant, Empalme, July 1983,
TUBARIA Gomez & Alfaro 20621 (F)
agrocybeoides Singer, 1941e: 96 gomezii Singer, 199 la: 30, 89
Type information: USSR. Altai Mtns.: near Lake HOLOTYPE: Costa Rica. Cartago: near San
Kurai, 16 August 1937, Singer (LE?) Cristobal, 2 km SE of junction of Highway
furfuracea novembris Singer, 1969a: 322
var. No. 2 to San Cristobal, November 1983, Go-
HOLOTYPE: Argentina. Neuquen: Lago Lacar, mez & Alfaro 22018b (F)
6009 (BAFC) guanacastensis Singer, 1983f: 110
omphaliopsis Singer in Singer & Digilio,
1951e: 397 7 km NW of town, October 1982, Gomez
HOLOTYPE: Argentina. Tucuman: Ciudad, 7
18724 (F)
January 1949, Singer T 89 (LIL) [Published hondurensis Singer &

Ivory, 1983f: 109
in 1952] HOLOTYPE: Honduras. Siguatepeque, 12 Oc-
pentstemonis Singer, 1957a: 368 tober 1976, Ivory S/249 (F)
HOLOTYPE: Mexico. Dto. Federal, Paso de jalapensis Singer &
Garcia, 199 la: 16, 89
Corts, 21 July 1957, Singer M
1560 (MICH) HOLOTYPE: Mexico. Jalapa: near border of Ve-
pseudoripartites Singer in Singer & Digilio, racruz, NE of Lencero, 14-19 July 1985, Gar-
195 le: 396 cia 4767 (F)
HOLOTYPE: Argentina. Salta: Corralitos, 24 lividobrunneus Singer, 1973d: 103
March 1951, Singer T 1419 (LIL) [Published HOLOTYPE: Mexico. Oaxaca: San Augustin,
in 1952] 10 July 1969, Singer 8393 (F) M
96 Chapter Three
TYLOPILUS
minor Singer, 19451: 799 williamsii Singer & Garcia, 199 la: 33
Type information: USA. Florida: Alachua Coun- HOLOTYPE: Mexico. Veracruz: 5 km NE of
ty, Gainesville. Lencero, road Jalapa- Veracruz, 14 July 1985,
mitissimus Singer &
Gomez, 199 la: 17, 89 Garcia 4759 (F)
7 June 1983, Singer & Gomez [Singer no.] B VELOPORPHYRELLUS G6mez & Singer,
14090 (F) 1984f: 293
montanus Singer, 1989a: 125
HOLOTYPE: Costa Rica. San Jose: La Chonta, Type species: Veloporphyrellus pantoleucus
1 1
July 1982, Gomez 19199 (F) Gomez & Singer
montoyae Singer &
Garcia, 199 la: 19, 89
HOLOTYPE: Mexico. Veracruz: Road San An- pantoleucus Gomez &
Singer, 1984f: 293
dres-Tlalnehuayocan, Plan de Sedeno, 28 June
HOLOTYPE: Costa Rica. Cartago: near San
1986, Montoya-Bello 699 (F) Cristobal, 1 .5 km SE of junction of dirt road

to San Cristobal and Rt. 2, November 1983,
nicaraguensis Singer & Ivory, 1983f: 107
HOLOTYPE: Gomez 22007 (F)
Nicaragua. Slilma Sia, May 1976,
Ivory S/20 (F)
peralbidus var. rhodoconius Singer, 1945i: 799 VOLVARIELLA
ty, Gainesville. lepiotospora Singer, 1955a: 774
potamogeton Singer, 1978a: 433 F
HOLOTYPE: Amazonas: Ponta Negra,
Brazil.
theson Hammock, 20 October 1942, Singer
1143 (F)
18 December 1977, Singer & Prance [Singer
no.] B 10351 (INPA) macrospora Singer, 1989a: 92
potamogeton var. aquarius Singer, 1983f: 122 B 9863 (F)
HOLOTYPE: Brazil. Amazonas: Igarape do Tar- Manaus, 11 May 1977, Singer
B psammophila Singer, 1989a: 92

umazinho, 14 December 1978, Singer
11433 (INPA)
HOLOTYPE: Argentina. Buenos Aires: Parque
Ezeiza, 1966, Havrizco [Singer no.] 703
potamogeton var. mitis Singer, 1989a: 125
HOLOTYPE: Brazil. Amazonas: Igarape do Tar- (BAFC)
rondoniensis Singer, 1989a: 93
HOLOTYPE: Brazil. Rondonia: Ouro Preto, 17
(INPA)
B 10969 (INPA)
April 1978, Singer
sanctaerosae Singer, 1983f: 112
HOLOTYPE: Costa Rica. Guanacaste: Parque subxerophytica Singer, 1989a: 93
HOLOTYPE: Argentina. Tucuman: Pie del Per-
iquillo, 30 December 1951, Singer T 1687
Singer [Singer no.] B 12436 (F)
(LIL). Deposited as "Volvariella taylori"
subcellulosus Singer &
Garcia, 199 la: 32, 90
HOLOTYPE: Mexico. Tamaulipas: Rancho El
Cielo, 9 July 1984, Garcia 3781 (F) VOLVOLEPIOTA
subniger Singer, Garcia & Gomez, 199 la: 38,
albida Singer, 1959i: 12
90
HOLOTYPE: Argentina. Tucumdn: Sierra de
HOLOTYPE: Costa Rica. San Jose: Jardin de &
San Javier, 9 January 1959, Singer, Lowy
Dota, 6 km W of Empalme, 8 June 1984,
Guerrero [Singer no.] T 3202 (LIL)
Singer B 14555a (F)
tabacinus var.amarus Singer, 1945i: 799
Type information: USA. Florida: Alachua Coun- WERAROA Singer, 1958b: 46
ty, Gainesville.
tabacinus var. dubius Singer, 1945i: 799
Type species: Secotium novaezelandiae Cun-
ningham
ty, Gainesville.
patagonica Singer & Wright, 1959d: 607
venezuelae Singer & Digilio, 1960i: 163 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
HOLOTYPE: Venezuela. Miranda: Los Guaya- pi National Park, Lago Traful, no date, An-
bitos, 8 June 1958, Dennis 1001 (K, LIL) dreis & Wright BAFC 20.080 (BAFC)
vinaceogriseus Singer, Garcia & Gomez, spadicea Singer, 1960d: 389

199 la: 40, 90 HOLOTYPE: Argentina. Neuquen: Nahuel Hua-
HOLOTYPE: Costa Rica. San Jose: La Chonta, pf National Park, Quetrihue, 3 May 1959,
17 June 1983, Singer B 14360 (F) Diem 2875 (LIL)

XANTHOCHROUS
XANTHOCHROUS linden Singer, 1945b: 296

Type information: Liberia, near Banga, October,
cuticularis f.
platani Singer, 19361: 423 Under 1211 (FH)
No type information. No Latin diagnosis. &
microsporus Singer Grinling, 1967c: 372
HOLOTYPE: Zaire. Brazzaville: near Djoue riv-
December 1965, Grinling 51,204 (BAFC)

XANTHOCONIUM Singer, 1944a: 361
er,
radicicola Singer, 1978a: 430

Type species: Gyroporus stramineus Murr. HOLOTYPE: Amazonas: Along Rio Ne-
Brazil.
gro, 17 January 1978, Araujo 916 (INPA)

scrobiculatus Singer, 1983f: 81
XEROCOMUS HOLOTYPE: Brazil. Amazonas: Road Manaus-
Caracaraf, km 45, 24 March 1978, St. John &
amazonicus Singer, 1978a: 429
Manaus, 20 December 1977, Singer & Araujo

truncatus Singer, Snell & Dick, 1959m: 573
HOLOTYPE: USA. North Carolina: Highlands,
26 July 1941, Coker 12426 (UNC, FH)
amazonicus var. obscuratus Singer Araujo, &
1983f: 78
HOLOTYPE: Brazil. Amazonas: Ponta Negra,
XEROMPHALINA
30 December 1978, Singer B 11451 (INPA)
bakshii Singer &
Singh, 197 la: 31 amtroandina Singer, 1965d: 303, 309
HOLOTYPE: India. Uttar Pradesh: Dehra Dun, HOLOTYPE: Rio Negro:
Argentina. Laguna
= For-
July 1968, Bakshi 9021 (FRI, F). (FRI Frias, 21 March 1963, Singer M 3120 (BAFC)
est Research Institute, India)
helbergeri Singer in Singer & Digilio, 195 le:
campinaranae Singer, 1983f: 66 240
Caracaraf, km 125, 4 April 1978, Singer B
del Taficillo, 29 July 1951, Helberger [Singer
10917 (INPA)
no.] T 1587 (LIL) [Published in 1952]
chapinii Singer, 1983f: 74
Brazil.
mesopora Singer, 1938a: 196
Type information: USSR. Kazakhstan: Aktura,
umazinho, 12 March 1980, Chapin, Araujo &
Czuiskie Alpy, 2 August 1937, Vasilieva
(LE?)
flavus Singer & Kuthan, 1976h: 153
tropicalis Singer, 1989a: 89
HOLOTYPE: Czechoslovakia. Southern Mora-
via: Zdrava Voda, zadanicky, 4 August 1974,
Kfiz & C 5759 Itacoatiara, km 21, 18 November 1978,

Singer [Singer no.] (F)
Aguiar &
Souza [Singer no.] B 11427 (INPA)
globuliger Singer, 1983f: 68
HOLOTYPE: Brazil. Amazonas: Road Manaus- yungensis Singer, 1989a: 89
km 125, 22 B HOLOTYPE: Bolivia. La Paz: Nor-Yungas, 1
Caracaraf, May 1978, Singer
11015 (INPA) February 1956, Singer B 795 (F)
hemixanthus Singer, 1945b: 292

HOLOTYPE: USA. Florida: Alachua County, 1
XERULA
July 1943, Singer F 2473? (FH)
hypoxanthus Singer, 1945b: 289 verruculosa Singer, 1953b: 6
HOLOTYPE: Argentina. Catamarca: Rio Chac-
ras, 10 February 1952, Singer T 1845 (LIL)
1942, Singer F 626 (FH)
illudens ssp. xanthomycelinus Singer, 1945b:
294 XERULINA Singer, 1961b: 59
Gainesville, May 1943, Singer F 2167 (FH) Type species: Lentinus chrysopeplus Berk. &
indicus Singer, 1946g: 104 Curt.
Type information: USA. Roxb Ladhar St., Sultan
Ahmed 44 (FH). [Published in 1948].
inundabilis Singer, 1983f: 70 ZELLEROMYCES Singer & Smith, 1960h: 18
HOLOTYPE: Brazil. Amazonas: Igarape do Tar-
umazinho, 14 December 1978, Singer B Type species: Zelleromyces cinnabarinus Sing-
11432 (INPA) er & Smith
98 Chapter Three
ZELLEROMYCES
cinnabarinus Singer & Smith, 1960H: 19 oregonensis Singer & Smith, 1960h: 19
HOLOTYPE: USA. Louisiana: Jackson County, HOLOTYPE: USA. Oregon: Comstock, 23 Oc-
26 December 1931, Smith & Brown s.n. tober 1937, Zeller 8333 (NY)
(MICH)
gilkeyae Singer & Smith, 1960h: 21
HOLOTYPE: USA. Oregon: near Reedsport, 22
October 1938, Gilkey [Zeller no.] 8498 (NY)

The Publications of Rolf Singer
Gregory M. Mueller
Qiuxin Wu
James H. Ginns
Scott A. Redhead
intended to be a complete list of the

is
1922b Pflanzengeographische Beobachtungen an ober-
Thispublications of Rolf Singer, not only those bayerischen und oberpfalzischen Hymenomyceten.
Zeitschrift fur Pilzkunde 1: 63-66.
describing new taxa. Publications are listed by the 1922c Uber die Gattung Psalliota. Zeitschrift fur Pilz-
year stated on the publication, and a suffix (a, b, kunde 1: 21-24.
c, etc.) designates multiple publications within a 1922d Uber den Wert der Gattung Rozites (Karsten).
year. Whenthe year stated on the publication is Annales Mycologici 20: 296-299.
different from the year of actual publication (e.g., 1922e Zur Pilzflora der Oberpfalz. Zeitschrift fur Pilz-
kunde 1: 48.
1950J), the actual date of publication is noted in
1923a Die Taublinge Mitteleuropas. Bin monogra-
[ ] at the end of the citation. Titles of papers are
phischer Beitrag zur Pilzflora Deutschlands und der
primarily given in their original languages.
Only angrenzenden Lander. Zeitschrift fur Pilzkunde 2: 1-
a few papers in Russian are translated into En- 20.
glish,and these are presented in [ ]. Coauthorships 1923b Eine fragliche Psilocybe. Zeitschrift fur Pilzkun-
are noted at the end of the citation, and names of de 2: 39-40.
the coauthors that appeared in front of Singer's 1923c Russula xerampelina (Schff.) Fr. Zeitschrift fur
Pilzkunde 2: 172-174.
name on the publication are underlined.
1923d Interessantes Uber Winterpilze. Zeitschrift fur
Two aspects of the format used in the following Pilzkunde 2: 47.
publication list need explanation. First, although 1923e Uber deutsche Namen fur die Pilze. Zeitschrift
the papers are presented in chronological order to fur Pilzkunde 2: 47.
year, within a year they are not necessarily listed 1923f Berichtigungen zu "Die Taublinge Mitteleuro-
chronologically. Publications are listed in the or- pas." Zeitschrift fur Pilzkunde 2: 113.
der that they appear in Dr. Singer's bound reprints 1923g Pflanzengeographische Beobachtungen an ober-
since dates (year plus letter) were assigned as data bayerischen und oberpfalzischen Hymenomyceten. 2.
Reihe. Zeitschrift fur Pilzkunde 2: 115.
were added to the database of new taxa presented
1924a Kritische Formenkreise aus der Gattung Russula.
in the preceding chapter. Second, we cite publi- I. Zeitschrift fur Pilzkunde 3: 73-78.
cations as they appear on the actual article, so

1924b Kritische Formenkreise aus der Gattung Russula.
some titles do not conform to current, standard II. Zeitschrift fur Pilzkunde 3: 107-1 12.
journal citations such as given in the Botanico- 1924c Berichtigung. Zeitschrift fur Pilzkunde 3: 119.
Periodicum-Huntianum (B-P-H). 1 925a Zur flws.YM/a-Forschung. Zeitschrift fur Pilzkunde
5: 73-80.
1925b Pflanzengeographische Beobachtungen an ober-

bayerischen und oberpfalzischen Hymenomyceten. 3.
1922a Collybia velutipes (Curt.) n.f. aestivalis und das Reihe. Zeitschrift fur Pilzkunde 4: 37-44.
periodische Pilzwachstum in den vier Jahreszeiten. 1925c Kritische Formenkreise aus der Gattung Russula.
Zeitschrift fur Pilzkunde 1: 40-41. III. Zeitschrift fur Pilzkunde 5: 13-18.
The Publications of Rolf Singer 101

1926 Monographic der Gattung Russula. Ire edition. 1934a Vorarbeiten zur Russulaflora osterreichs. Annales
Hedwigia66: 163-260. Mycologici 32: 448-463 (with Th. Cernohorsky).
1927a Die haufigeren Basidiomyceten der Umgebung 1934b Russula variata Bann. und Russula lactea Fr.
von Otz (Tirol). Zeitschrift fur Pilzkunde 6: 145-147. Feddes Repertorium 33: 350-352.
1927b Wie bestimmt man frische Taublinge? Zeitschrift 1935a Etude systematique sur les Melanoleuca d'Eu-
fur Pilzkunde 6: 169-176. rope et cle des especes observees en Catalogne. Ca-
1927c Russula vesca und Russula constans. Zeitschrift vanillesia7: 122-132.
fur Pilzkunde 6: 122-123. 1935b Supplemente zu meiner Monographic der Gat-
1928a Drei seltene, ausserhalb Deutschlands heimische tung Russula. Annales Mycologici 33: 297-352.
Russulae. Zeitschrift fur Pilzkunde 7: 42-44. 1935c Sur quelques Russules exotiques. Annales de
1928b Die Taublinge der otztaler Alpen. Zeitschrift fur Cryptogamie exotique 8: 88-93.
Pilzkunde 7: 75-77. 1935d Sur la classification des Russules. Bulletin de la
1928c Ziele der systematischen Hymenomycetenfor- Societe Mycologique de France 51: 281-304.
schung. Zeitschrift fur Pilzkunde 7: 161-167. 1935e Russula schiffneri Sing. Bulletin de la Societe
1928d Die Taublingsexkursion am 6 September 1927- Mycologique de France 51: 102-103.
anla'Blich des Miinchner Kongresses. Zeitschrift fiir 1936a Melanoleuca kavinae (Pil. et Ves.) Sing. nov.
Pilzkunde 7: 114-115. comb., de la serie des Alboflavidae Sing. Revue de
1928e Neue Mitteilungen iiber die Gattung Russula. I. Mycologie 1: 38-39.
R. sardonia Fr. oder der Formenkreis R. queletii Fr.
1936b Contribution a 1'etude des Russules. II. Obser-
und II. Die gelben hellsporigen Piperatae. Hedwigia vations floristiques. Bulletin de la Societe Mycolo-
48: 191-202.
gique de France 52: 111-114.
1928f Besprechung einiger Arbeiten aus dem Bulletin
1936c Les Russules de 1'herbier de Boudier. Revue de
trimestriel de la Societe Mycologique de France.
Mycologie 1: 19-24.
Tome XLII, 1926. Zeitschrift fur Pilzkunde 7: 14-15.
1936d Notes sur quelques Basidiomycetes. I. Revue de
1928g Berichtigung. Zeitschrift fur Pilzkunde 7: 30.
Mycologie 1: 75-84.
1928h Verschiedene Beitrage zur Geniessbarkeit des
1936e Notes sur quelques Basidiomycetes. II. Revue de
netzstieligen Hexen-Rohrlings. Ein gefa'hrlicher Gift-
Mycologie 1: 279-293.
pilz? Zeitschrift fur Pilzkunde 7: 85-87.
1936f Studien zur Systematik der Basidiomyceten. I.
1929a Beitrag zur systematischen Stellung des Pleuro-
Beihefte zum Botanischen Centralblatt 56: 137-156.
tus nidulans. Archiv fiir Protistenkunde 65(3): 314-
320. 1936g Studien zur Systematik der Basidiomyceten. II.
Beihefte zum Botanischen Centralblatt 56: 157-174.
1929b Pilze aus dem Kaukasus. Ein Beitrag zur Flora
I.
des siidwestlichen Zentralkaukasus. Beihefte zum Bo- 1936h Das System der Agaricales. I. Annales Mycolo-
tanischen Centralblatt 46(2): 71-113. gici 34: 286-378.
1929c Sitzung der mathematisch-naturwissenschaftlich- 1936i Bemerkungen iiber einige Basidiomyceten. An-
en Klasse vom 10 Janner 1929. Akademie der Wis- nales Mycologici 34: 423-434.
senschaften in Wien. No. 1: 1-4.
1936J Sur quelques Clavaires collectiones dans les or-
1929d Sitzung der mathematisch-naturwissenschaftlich- angeries de Tlnstitut Botanique de 1' Academic des
en Klasse vom 24 Oktober 1929. Akademie der Wis- Sciences de 1'USSR a Leningrad. Journal Botanique
senschaften in Wien. No. 20: 1-2. de L'URSS 21: 723-725 (with A. S. Bondartsev).
1929e R. coerulea, R. amoenata a R. amara Ku. (ne 1937 Notes sur quelques Basidiomycetes. III. Revue de
Mre). Casopis deskoslovenskych houbafu 9: 77-79 Mycologie 2: 226-242.
(with J. Zvara). 1938a Notes sur quelques Basidiomycetes. IV. Revue
1929f Eine neue Russula-ALrt: Russula mairei nov. spec. de Mycologie 3: 187-199.
Archiv fUr Protistenkunde 65(3): 306-314.
1938b Contribution a 1'etude des Russules. HI. Quel-
1929g Neue Mitteilungen iiber die Gattung Russula. III- ques Russules americaines et asiatiques. Bulletin de
IV. Hedwigia 67: 253-261. la Societe Mycologique de France 54: 132-177.
1929h Russula mairei Singer. Bulletin de la Societe My- 1938c Sur les genres Ixocomus, Boletinus, Phylloporus,
cologique de France 45: 103-105. Gyrodon et Gomphidius. Revue de Mycologie 3: 35-
1930 Contributions a 1'etude des Russules. Note sur 53.
deux variety's nouvelles. Bulletin de la Societe My- 1938d Sur les genres Ixocomus, Boletinus, Phylloporus,
cologique de France 46: 209-212. Gyrodon et Gomphidius. 2. Les Boletinus. Revue de
dem Kaukasus. II. Ein Beitrag zur Flora
1931 Pilze aus Mycologie 3: 157-177.
Swanetiens und einiger angrenzender Ta'ler. Beihefte 1938e De nonnullis Basidiomycetibus. I. BotaniCeskie
zum Botanischen Centralblatt 48: 5 3-542.1
materialy Otdela sporovyh rastenij BotaniCeskogo in-
1932 Monographic der Gattung Russula. Beihefte zum stitutaimeni V. L. Komarova Akademii Nauk SSSR
Botanischen Centralblatt 49: 205-380. 10-12: 4-18.
1933 Mykologische Studien im Hochgebirge des Kau- 1938f Les Russules de Catalunya i clau per a llur de-
kasus. Schweizerische Zeitschrift fiir Pilzkunde 11: terminacio. Cavanillesia Rerum Botanicarum Acta
157-160. (Barcinone) 8: 144-159.
102 Chapter Four

1938g Uber Larchen, Zirben-und Birkenrdhrlinge. 1943b A monograph on the genus Leucopaxillus Bour-
Schweizerische Zeitschrift fur Pilzkunde 16: 123-126, sier. Papers of the Michigan Academy of Science, Arts
134-137, 148-150. & Letters 28: 85-132, 8 plates (with A. H. Smith).
1938h [The production of ink from the spores of fungi]. 1943c Type studies on Basidiomycetes. II. Mycologia
Priroda 1: 121-123. 35: 142-163.
1938i Reviewof: [Mushrooms, procurement and pro- 1943d K yestestvennoy sisteme trutovykh gribov [On
the natural system of Polyporaceous fungi]. Soviet-
cessing],by Prof. L. A. Lebedeva. Priroda 7-8: 210-
212 (with V. Transchel, A. S. Bondartsev, and L. Vas- skaia Botanika 1943(1): 29-43 (with A. S. Bondart-
ilieva). sev).
1939a Notes sur quelques Basidiomycetes. V. Revue de 1944a New genera of fungi. Mycologia 36: 358-368.
Mycologie 4: 64-72. 1 944b Notes on taxonomy and nomenclature of the pol-
1939b O novykh dannykh dlia sistematiki i
filogenii ypores. Mycologia 36: 65-69.
Agaricales kak potomkov Gasteromycetes [On new 1944c On some Basidiomycetes new for the United
data for the taxonomy and phylogeny of the Agari- States. Mycologia 36: 552-554.
cales as descendants of the Gasteromycetes]. Soviet- 1945a The Boletineae of Florida with notes on extra-
skaia Botanika 1939(1): 95-98. limital species. I. The Strobilomycetaceae. Farlowia2:
1939c Phylogenie und Taxonomie der Agaricales. 97-141.
Schweizerische Zeitschrift fur Pilzkunde 17: 23-28, 1945b The Boletineae of Florida with notes on extra-
35-39, 52-57, 71-73, 84-87, 97-101. limital species. II. The Boletaceae (Gyroporoideae).
1939d [On anti-Darwinistic tendencies in contemporary Farlowia 2: 223-303.
taxonomies of higher basidiomycetous fungi]. Soviet- 1945c New genera of fungi. II. Lloydia 8: 139-144.
skaia Botanika 1939(6-7): 139-144. 1945d New and interesting species of Basidiomycetes.
1939e Contribution a F etude des Russules. IV: Qu- I.
Mycologia 37: 425-439.
elques Russules americaines et asiatiques (suite). Bul- 1945e Notes on Farlow's Agaricales from Chocorua.
letin de la Societe Mycologique de France 55: 226- Farlowia 2: 39-51.
283.
1945f A monograph of the genus Cystoderma. Papers
1939f Review of: Les Lactario-Russules du Domaine of the Michigan Academy of Science, Arts & Letters
Oriental de Madagascar, by R. Heim. 196 pp. Paris. 30: 71-124 (with A. H. Smith).
Sovietskaia Botanika 1939(1): 128.
1945g The Laschia-complex (Basidiomycetes). Lloydia
1940 Notes sur quelques Basidiomycetes. VI. Revue de 8: 170-230.
Mycologie 5: 3-13. 1945h The taxonomic position of Polyporoletus subli-

194 la Zur Systematik der Polyporaceen. Annales My- vidus. Mycologia 37: 124-128 (with W. H. Snell and
cologici 39: 43-65 (with A. S. Bondartsev). W. L. White).
1941b De nonnullis Basidiomycetibus. II. Botanideskie 1945i New Boletaceae from Florida. Mycologia 37:
materialy Otdela sporovyh rastenij Botanideskogo in-
797-799.
stituta imeni V. L. Komarova Akademii Nauk SSSR 1946a The Boletineae of Florida with notes on extra-
5(4-6): 84-86. limital species. IV. The lamellate families (Gomphi-
1941c Is shiitake a Cortinellusl Mycologia 33: 449- diaceae, Paxillaceae and Jugasporaceae). Farlowia 2:
451. 527-567.
194 Id O 1946b Proposals concerning the nomenclature of the gill

ponyatii vida u vysshich basidialnych gribov
[On the concept of species in higher Basidiomycetes]. fungi including a list of proposed lectotypes and gen-
Sovietskaia Botanika 1940(5-6): 262-269. era conservanda. Mycologia 38: 240-299 (with A. H.
Smith).
1941e Cortinariaceae novae et minus cognitae. Botani-
1946c Type studies on agarics. II. Lloydia 9: 114-131.
Ceskie materialy Otdela sporovyh rastenij Botani5es-
1946d The taxonomic position of Pholiota mutabilis
kogo instituta imeni V. L. Komarova Akademii Nauk
SSSR 5(7-9): 92-100. and related species. Mycologia 38: 500-523 (with A.
H. Smith).
1942a A monographic study of the genera Crinipellis
and Chaetocalathus. Lilloa 8: 441-534 [published in 1946e Two new species in the Agaricales. Mycologia
38: 687-690.
1943]
1942b Das System der Agaricales. II. Annales Mycol- 1946f A new
species of Hydropus (Ktihn.) Sing. (Agar-
ogici 40: 1-132.

icales).
1942c Type studies on agarics. Lloydia 5: 97-135. 1946g New and interesting species of Basidiomycetes.
II. Papers of the Michigan Academy of Science, Arts
1942d Type studies on Basidiomycetes. I. Mycologia & Letters 32: 103-150. [published in 1948]
34: 64-93.
1947a The Boletoideae of Florida. The Boletineae of
1942e Roaming Russia's Caucasus. National Geograph- Florida with notes on extralimital species. III. The
ic 82: 91-121. American Midland Naturalist 37: 1-135.
1942f The Caucasus: Soviet training ground for moun- 1947b New genera of fungi. III. Mycologia 39: 77-89.
taineers. Appalachia 94: 149-154. 1947c Champignons de la Catalogne. Especes obser-
1943a Das System der Agaricales. III. Annales Mycol- vees en 1934. Collectanea Botanica (Barcelona) 1:
ogici 41: 1-189. 199-246.

1947d Coscinoids and coscinocystidia in Linderomyces 19501 Should Dr. D. Rogers' proposals for the regu-
P.
lateritius. Farlowia 3: 155-157. lation for fixing generic types be accepted as pub-
1947e Type studies on Basidiomycetes. III. Mycologia
lished in Mycologia 41: 219, 1949? Mycologia 42:
195-197.
39: 171-189.
1947f Additional notes on the genus Leucopaxlllus. My- 195 la Antibiotic activity and formation of ectotrophic
cologia 39: 725-736 (with A. H. Smith), [published mycorrhiza in relation to natural affinity in the clas-
in sification of the Basidiomycetes. Seventh International
1948]
Botanical Congress Proceedings. 401-402.
1947g Contributions towards a monograph of the genus
Crepidotus. Lilloa 13: 59-95. [published in 1948]
195 Ib New genera of fungi. V. Mycologia 43: 598-604.
1948a New genera of fungi. IV. Mycologia 40: 262- 1951c Thaxterogaster-a new link between Gasteromy-
264. cetes and Agaricales. Mycologia 43: 215-228.
1948b Review of: North American Species of Mycena, 195 Id Type studies on Basidiomycetes. V. Sydowia 5:
by A. H. Smith. Bulletin of the Torrey Botanical Club 445-475.
75: 117-119. 1951e Prodromo de la Flora Agaricina Argentina. Lilloa
1948c Diagnoses fungorum novorum Agaricalium. Sy- 25: 5-461 (with A. P.Digilio). [published in 1952]
dowia 2: 26-42.
195 If Die Nomenklatur der hoheren Pilze. Schweizer-
1948d Notes on the genus Cystoderma. Mycologia 40: ische Zeitschrift fur Pilzkunde 29: 204-228.
454_460 (with A. H. Smith).
1952a Type studies on Basidiomycetes. VII. Sydowia 6:
1948e Emendations to our proposals concerning the no- 344-351.
menclature of the gill fungi. Mycologia 48: 627-629
1952b Russulaceae of Trinidad and Venezuela. Kew
(with A. H. Smith).
Bulletin 1952: 295-301.
1949a The genus Gomphidius Fries in North America.
1952c The agarics of the Argentine sector of Tierra del
1949b [Materials for the flora of pileate fungi of the Fuego and limitrophous regions of the Magallanes
area. I. White and pink spored groups. Sydowia 6:
"Kivach" Game Preserve of the Karelo-Finnish SSR].
165-226.
In: M. V. Freundling. Izv. Karelo-Finsk. fil. AN SSSR
4: 84-97. 1952d Le genre Gloiocephala Massee. Revue de My-
1949c The Agaricales in modern taxonomy. Lilloa 22: cologie 17: 161-164.
1-832. [published in 1951] 1952e Type studies on agarics. III. Lilloa 25: 463-514.
1950a Newand interesting species of Basidiomycetes. 1952f Review of: A

dictionary of color. 2nd ed. My-
III. Sydowia4: 130-157. cologia 44: 267-269 (with A. P. Digilio).
1950b Les Russules de PArgentine. Revue de Mycol- 1953a The agarics of the Argentine sector of Tierra del
ogie 15: 125-137. Fuego and limitrophous regions of the Magallanes
1950c Die Hoheren Pilze Argentiniens. Schweizerische area. II. The brown-spored genera (except Cortinar-
Zeitschrift fur Pilzkunde 28: 181-196. ius). Sydowia 7: 206-265.
1950d Zwei neue Pilzarten den Alpen. Schweizeri-
in 1953b Quelques Agarics nouveaux de 1'
Argentine. Re-
sche Zeitschrift fur Pilzkunde 28: 196-200. vue de Mycologie 18: 3-23.
1950e Supplementary notes on the genera Campanella 1953c Four years of mycological work in southern
and Favolaschia. Lloydia 13: 249-258. South America. Mycologia 45: 865-891.
1950f Novye osnovy dlya classification Panus y rod- 1953d Type studies on Basidiomycetes. VI. Lilloa 26:
stvennykh rodov [New bases for the classification of 57-159.
Panus and related genera]. Trudy BotaniCeskogo In-
1953e Review of: Les Agaricales. II. Russulacees, Hy-
stituta imeni V. L. Komarova Akademii Nauk SSSR,
ser. II, 6: 375-387.
grophoracees, Gomphidiacees, Paxillacees, Boleta-
cees, by P. Konrad and A. Maublanc. Mycologia 45:
1950g K izutcheniyu roda Amanita v SSSR [Amanita 477-478.
in the USSR]. Trudy Botanieskogo Instituta imeni V.
Komarova Akademii Nauk SSSR, 1953f Review of: Russula-Monographie (Die Pilze Mit-
L. ser. II, 6: 388-
401. teleuropas, Band III), by J. Schaffer. Mycologia 45:
478-480.
1950h Naucoria Fries y blizkiye rody v SSSR [Nau-
coria Fries and related genera in the USSR]. Trudy 1954a Part VI. Fungi, pp. 451-462. In: Polunin, N., ed.,
The Cryptogamic Flora of the Arctic. Botanical Re-
BotaniCeskogo Instituta imeni V. L. Komarova Aka-
demii Nauk SSSR, ser. II, 6: 402-498. view 20: 361-476.
1950i New genera of fungi. VI. Descolea antarctica, 1954b Agaricales von Nahuel Huapi. Sydowia 8: 100-
157.
genero y especie nuevos de Tierra del Fuego. Lilloa
23: 255-258. 1954c Uber Pilzbestimmung. Schweizerische Zeitschrift
1950J Type studies on Basidiomycetes. IV. Lilloa 23: fur Pilzkunde 32: 101-108.
147-246. [published in 1952] 1954d Review of: Flore analytique des Champignons
1950k [A manual of the collection of basidiomycetous Superieurs (Agarics, Bolets, Chanterelles) comprenant
fungi for their scientific study.] Trudy BotaniCeskogo les especes de 1' Europe occidentale et centrale ainsi
Instituta imeni V. L. Komarova Akademii Nauk SSSR, la plupart de celles de 1'Algerie et du Maroc, by R.
ser. II, 6: 499-543 (with A. S. Bondartsev). Kiihner and H. Romagnesi. Mycologia 46: 129-132.
104 Chapter Four

1954e New data of affinities of certain Agaricales and 1956i Die Pilznomenklatur seit 1954. Schweizerische
certain Gasteromycetes of South America. Eighth In- Zeitschrift fur Pilzkunde 34: 33-43.
ternational Botanical Congress Proceeding. 29-30. 1957a Fungi Mexicani, series prima-Agaricales. Sy-
1954f Pages 554-555, 565-566, 571. In: Hueck K. Der dowia 11: 354-374.
Anden-Erlenwald (die Alnus jorullensis-Assoziation) 1957b New and interesting species of Basidiomycetes.
in der Provinz Tucuman (Argentinien). Festschrift fur V. 11: 141-272.
Sydowia
Erwin Aichinger zum 60. Geburtstag. Sonderfolge der
1957c Las Boletaceas Austrosudamericanas. Lilloa 28:
Schriftenreihe Angewandte Pflanzensozilologie 1:
247-268 (with A. P. Digilio).
512-572.
New and interesting species of Basidiomycetes. 1957d The genus Galerina: An outline of its classifi-
1955a
cation.Sydowia 11: 446-453 (with A. H. Smith).
IV. Mycologia 47: 763-777.
1957e Un nuevo hongo fosil de los bosques petrificados
1955b Agaricales from Tristan da Cunha. Results of the
de Santa Cruz, Prov. Patagonia. Ameghiniana (Buenos
Norwegian Scientific Expedition to Tristan da Cunha
1937-1938. 38: 15-18. Aires) 1: 40-41 (with S. Archangelsky).
Un 1957f Sacred mushrooms inspire medical research. Chi-

1955c Clitocybe cortique et un Clitocybe faux. Bul-
letin de la Societe Mycologique de France 71: 147- cago Natural History Museum Bulletin 28(12): 7.
152. 1957g Neue Literatur: The Russulas, by W. R. Arde.
1955d New species of Agaricales from Pernambuco. Sydowia 11: 473-474.
Anais da Sociedade de Biologia de Pernambuco 13: 1957h New genera of fungi X-Pachylepyrium. Sydowia
225-229. 11: 320-322. [published in 1958]
1955e New species of Galerina. Mycologia 47: 557- 1957i Neuere Tendenzen in der Systematik und Klas-
596 (with A. H. Smith). sifikation der Basidiomyceten. Schweizerischen Zeit-
1955f Type studies on Basidiomycetes. VIII. Sydowia schrift fur Pilzkunde 35: 33-43.
9: 367-431. 1958a New genera of fungi. VIII. Notes concerning the

sections of the genus Marasmius FT. Mycologia 50:
1955g Basidioneuromycosis on man. Anais da Socie-
dade de Biologia de Pernambuco 13: 52-60 (with A. 103-110.
C. Batista and J. A. Maia). 1958b New genera of fungi. IX. The probable ancestor
1955h "Asiatische" und "Amerikanische" Pilze in Eu- of the Strophariaceae: Weraroa gen. nov. Lloydia 21:
ropa. Zeitschrift fur Pilzkunde 21: 1-4. 45-47.
1955i The nomenclature of Armillaria, Hypholoma and 1958c Studies on secotiaceous fungi. VIII. A new genus
Entoloma. Mycologia 47: 147-149. in the Secotiaceae related to Gomphidius. Mycologia
50: 927-938 (with A. H. Smith).
1955J Staude redivivus. Mycologia 47: 270-272.
1955k "Basidiomycetes of South America" Report of 1958d Fungi Mexicani, series secunda- Agaricales. Sy-
committee on research. Yearbook of the American dowia 12: 221-243.
Philosophical Society for 1954, Geological & Biolog- 1958e Studies towards a monograph of the South Amer-
ical Sciences. 166-168. ican species of Marasmius. Sydowia 12: 54-148.
19551 Die Pilznomenklatur seit 1954. Schweizerische 1958f Pilze, die Zerebralmyzetismen verursachen.
Zeitschrift fur Pilzkunde 33: 137-143. Schweizerische Zeitschrift fur Pilzkunde 36: 81-89.
1955m Die Pilznomenklatur seit 1954. Schweizerische
1958g Monographs of South American Basidiomycetes,
especially those of the east slope of the Andes and
1955n Le genre Melanomphalia Christiansen. Revue de Brazil. I. The genus Pluteus in South America. Lloy-
dia 21: 195-299.
Mycologie20: 12-17.
1956a New genera of fungi. VII. Mycologia 48: 719- 1958h Mycological investigation on Teonandcatl, the
727. Mexican hallucinogenic mushroom. Part II. A taxo-
1956b A nomic monograph of Psilocybe, Section Caerulescen-
fungus collected in the Antarctic. Sydowia
tes. Mycologia 50: 262-303 (with A. H. Smith).
Beihefte I: 16-23.
1956c Contributions towards a monograph of the genus 1958i New species of Galerina. Mycologia 50: 469-
Pluteus. Transactions of the British Mycological So-
489 (with A. H. Smith).
ciety 39: 145-232. 1958J A petrified basidiomycete from Patagonia. Amer-
1956d Versuch einer Zusammenstellung der Arten der ican Journal of Botany 45: 194-198 (with S. Archan-
Gattung Phaeomarasmius. Schweizerische Zeitschrift gelsky).

fur Pilzkunde 34: 44-47, 53-65. 1958k New species of Psilocybe. Mycologia 50: 141-
1956e Boletus luridus in North America. Mycologia 48: 142 (with A. H. Smith).
768-769. 19581 A new species of Psathyrella. Lloydia 21: 26-28
1956f The Armillariella mellea group. Lloydia 19: 176- (with A. H. Smith and G. Guzman Huerta).
187. 1958m Studies on secotiaceous fungi. I. A monograph
1956g The Pleurotus-hirtus-comp\e\. Mycologia 48: of the genus Thaxterogaster. Brittonia 10: 201-216
852-859. (with A. H. Smith).
1956h Campbellia africana and Versipellis variegata. 1958n Studies on secotiaceous fungi. II. Endoptychum
Taxon 5: 30-33. depressum. Brittonia 10: 216-221 (with A. H. Smith).

1958o Studies on secotiaceous fungi. III. The genus 1960f Monographs of South American Basidiomycetes
Weraroa. Bulletin of the Torrey Botanical Club 85: especially those of the east slope of the Andes and
324-334 (with A. H. Smith). Brazil. III. Reduced marasmioid genera in South
America. Sydowia 14: 258-280.
1958p The meaning of the affinity of the Secotiaceae
with the Agaricales. Sydowia 12: 1-43. 1960g Persoon's Synopsis 1801 as starting point for all
1958q Observations on agarics causing cerebral myce- fungi. Taxon 9: 35-37.

tisms. Mycopathologia et Mycologia Applicata 9: 1960h Studies on secotiaceous fungi. IX. The Astrogas-
261-284 (with S. I. Stein, R. W. Ames, and A. H. traceous series. Memoirs of the Torrey Botanical Club
Smith). 21: 1-112 (with A. H. Smith).
1958r Mycological investigation on Teonanactl, the 1960i Las boletaceas de Sudamerica tropical. Lilloa 30:
Mexican hallucinogenic mushroom. Part I. The his- 141-164 (with A. P. Digilio).
tory of Teonanacatl, field work and culture work. My- Sobre algunas especies de hongos presumible-
1960J
cologia 50: 239-261. mente psicotropicos. Lilloa 30: 117-127.
1958s Libri novi: Immunodiagnostika Aktinomicoza
1960k Ectotrophic forest tree mycorrhizae and forest
[Immunodiagnostic of Actinomycosis], by D. I. Asnin. communities. Ecology 41: 549-551 (with J. H. Mo-
Mycopathologia et Mycologia Applicata 9: 259-260. rello).
1959a Contributions towards a monograph of the genus
196 la Monographs of South American Basidiomycetes
Pluteus. II. Transactions of the British Mycological
Society 42: 223-226. Brazil. IV. Inocybe in the Amazone region with a sup-
1959b New and interesting species of Basidiomycetes. plement to part I (Pluteus in South America). Sydowia
VI. Mycologia 51: 375-400. 15: 112-132.
1959c Basidiomycetes from Masatierra (Juan Fernandez 1961b Diagnoses fungorum novorum Agaricalium. II.
Islands, Chile). Arkiv for Botanik, sen 2, 4: 371-400.
Sydowia 15: 45-83.
1959d A new species of the genus Weraroa from South 1961c Two genera of fungi new for South America.
America. Darwiniana 11: 607-610 (with J. Wright). Vellozia 1: 14-19. [published in 1962]
1959e Studies in secotiaceous fungi. VI. Setchelliogas-
196 Id Macroleplota ollvascens. Schweizerische Zeit-
ter Pouzar. Madrono 15: 73-79 (with A. H. Smith).
schrift fur Pilzkunde 39: 154-155 (with M. Moser).
1959f Studies on secotiaceous fungi. IV. Gastroboletus,
1961e Type studies on Basidiomycetes. X. Persoonia 2:
Truncocolumella, and Chamonixia. Brittonia 11: 205-
1-62.
223 (with A. H. Smith).
196 If Type studies on agarics. IV Sydowia 15: 133-
1959g Studies on secotiaceous fungi. V. Nivatogastrium 151.
gen. nov. Brittonia 11: 224-228 (with A. H. Smith).
1959h Hongos alucinogenos. Boletin de Academia Na- 1961g Mushrooms and truffles: botany, cultivation, uti-
lization. Leonard Hill, London. 272 pp.
cional de Ciencias, Cordoba, 41: 31-46.
1959i Dos generos de hongos nuevos para Argentina. 1961h Fungi of northern Brazil. Universidade do Recife
Institute de Micologia 304: 1-26.
Boletin de la Sociedad Argentina de Botanica 8: 9-
13. 1961i Review of: Die Gattung Phlegmacium (Schleim-
1959J Micorrhizas y comunidades forestales. Ninth In- kb'pfe). (Die Pilze Mitteleuropas, Band IV), by M.
ternational Botanical Congress Proceedings 2: IIA Moser. Mycologia 52: 823-825.
(with J. Morello). 1961J Nomina conservanda proposita (Rhodophyllus).
1959k Type studies on Basidiomycetes. IX. Sydowia Taxon 10: 249-251.
13: 235-238. 1962a Monographs of South American Basidiomycetes
19591 Libri novi: Candidosy (vozbuditeli, klinika, i ep- especially those of the east slope of the Andes and
idemologia) [Candidoses (causative agents, clinical Brazil. V. Gasteromycetes with agaricoid affinities (se-
and epidemiology)], by P. N. Kashkin. Myco-
data, cotiaceous Hymenogastrineae and related forms). Bol-
etin de la Sociedad Argentina Botanica 10: 52-67.
pathologia et Mycologia Applicata 11: 170-171.
1959m Notes on Boletes. XI. Mycologia 51: 564-577 1962b New genera of fungi. VIII. Persoonia 2: 407-
(with W. H. Snell and E. A. Dick), [published in 1960] 415.
1959n New and interesting species of Basidiomycetes. 1962c The Agaricales in Modern Taxonomy. 2nd ed.
Mycologia 51: 578-594. [published in 1960]
VII. Verlag J. Cramer, Weinheim. 915 pp.
1960a New genera of fungi. XI. A marasmioid agaric 1962d Estudio sobre los Basidiomicetes Antarcticos.
with bilateral trama. Lilloa 30: 375-379. Contribution del Institute Antarctico Argentine (Bue-
1960b Dos especies interesantes de Agaricales en Punta nos Aires) 71: 3-45 (with A. Corte).
Lara. Boletin de la Sociedad Argentina de Botanica 1962e. A study of Antarctic Basidiomycetes. Pp. 161-
8: 216-218. 163. In: Biologic Antarctique, Hermann, Paris (with
1960c Pluteus lilacinus. Mycologia 52: 337-338. A. Corte).
1 960d Three new species of Secotiaceae from Patagon- 1962f Review of: Flora Czechoslovak Republic-Gas-
ia. Persoonia 1: 385-391. teromycetes, Houby-Bfichatky (Gasteromycetes-puff-
1960e Studies on secotiaceous fungi. VII. Secotium and balls), A. Pilat., ed. Mycologia 52: 968-970.
Neosecotium. Madrono 15: 152-158 (with A. H. 1962g Review of: The Biology of Mycorrhiza, by J. L.
Smith). Harley. Mycologia 52: 970-972.
106 Chapter Four

1962h Review of: Champignons Comestibles et Vene- 1964f Marasmius congolais recueillis par Mme. Goos-
neux, by M. Locquin and B. Cortin. Mycologia 53: sens-Fontana et d'autres collecteurs Beiges. Bulletin
314-316. du Jardin Botanique de 1'etat Bruxelles 34: 317-388.
1962i Keys for the determination of the Agaricales. Ver- 1964g Areal und okologie des Ektotrophs in Siidamer-
lag J. Cramer, Weinheim. 64 pp. ika. Zeitschrift fUr Pilzkunde 30: 8-14.
1963a New genera of fungi. Xl-Endolepiotula. Sydowia 1964h A monograph of the genus Galerina Earle. Haf-
16: 260-262. ner Publishing Company, New York. 384 pp. (with A.
H. Smith).
1963b Oak mycorrhiza fungi in Colombia. Mycopath-
1964J Las Setas y Las Trufas. Leonard Hill (Books)
ologia et Mycologia Applicata 20: 239-252.
Ltd., Mexico D. F. 467 pp. [Spanish translation of
1963c Notes on secotiaceous fungi: Galeropsis and
196 Ig]
Brauniella. Koninklijke Nederlandse Akademie van
Wetenschappen-Amsterdam, Proceedings, Series C, 1964J Studies on secotiaceous fungi. X. Additional data
66: 106-117.
on Gastroboletus. Mycologia 56: 310-313 (with A.
H. Smith).
1963d Monographs of South American Basidiomycetes
1965a Die Rohrlinge. I. Die Boletaceae (Die Pilze Mit-
especially those of the east slope of the Andes. VI.
teleuropas, Band V). Verlag Klinkhardt, Bad Heil-
"Mesophelliaceae" and "Cribbeaceae" of Argentina
brunn. 131 pp.
and Brazil. Darwiniana 12: 598-611 (with J. E.
Wright and E. Horak). 1965b Marasmius. Flore iconographique des champi-
1963e Skepperiella populi y la filogenia de las "Cy- gnons du Congo 14: 253-278, 3 plates. Jardin Bota-
nique de l'etat, Bruxelles.
phellaceae." Boletin de la Sociedad Argentina Botan-
ica 10: 209-214. 1 965c Monographs of South American Basidiomycetes,
1963f Four interesting European Russulae of subsec-
Brazil. II. The genus Marasmius in South America.
tions Sardoninae and Urentinae, sect. Russula. Sy-
Sydowia 18: 106-358.
dowia 16: 289-301.
1965d Monographs of South American Basidiomycetes,
1963g The delimitation of the genus Pseudobaeospora.
Mycologia 55: 13-17. Brazil. X. Xeromphalina. Boletin de la Sociedad Ar-
1963h Un hongo nuevo comestible de Sudamerica. Bol- gentina Botanica 10: 302-310.
etin de la Sociedad Argentina de Botanica 10: 207-
1965e Interesting and new Agaricales from Brazil. Atlas
208. do Institute de Micologia (Universidade Federal de
1963i Der "Ektotroph", seine Definition, geographische Pernambuco) 2: 15-59.
Verbreitung und Bedeutung in der Forstokologie, pp. 1965f Two interesting Basidiomycetes from the State of
223-231. In: International Mykorrhiza-Symposium. Sao Paulo. Rickia 2: 11-16 (with O. Fidalgo).
Weimar, 1960.
1965g Typification of revalidated pre-starting point
1963J Beitrag zur Frage der Stropharia imaina Benedix. taxa. Taxon 14: 180-184.
1965h Schliissel zur Bestimmung der Familien und Gat-
1963k Paraphyses. Taxon 12: 147-150 (with I. J. Ga- tungen der Basidiomyzetenordnung Agaricales. Se-
mundi). paratdruck aus der Schweizerische Zeitschrift fur Pilz-
19631 A revision of the genus Thaxterogaster Singer. kunde 1965/66: 1-44.
Madrono 17: 22-26 (with A. H. Smith). 1965i Forest mycology and forest communities in South
1963m The genus Fuscoboletinus. Mycologia 55: 352- America. I. The early fall aspect of the mycoflora of
the Cordillera Pelada (Chile), with a mycogeographic
357 (with W. H. Snell and E. A. Dick).
analysis and conclusions regarding the heterogeneity
1963n The Herpell method of preparing fleshy fungi.
of the Valdivian Floral district. Mycopathologia et
Mycologia 55: 683-684. Mycologia Applicata 26: 129-191 (with M. Moser
1964a New genera of fungi XII: Hybogaster. Sydowia and the collaboration of I. Gamundi, E. R. de la Sola
17: 12-16. and G. Sarmiento).
1964b New genera of fungi XIII Rhodoarhenia. Sy- 1965J Review of: A contribution toward a monograph
dowia 17: 142-145. of North American species of Suillus, by A. H. Smith
1964c Monographs of South American Basidiomycetes, and H. Thiers. Mycologia 57: 669-674.
especially those of the east slope of the Andes and 1965k Una nueva especie de Hohenbuehelia sobre stipa.
Brazil. VII. Boletes and Related Groups in South Revista de la Facultad de Ciencias Agrarias, Univer-
America-The families Paxillaceae, Gomphidiaceae, sidad Nacional de Cuyo (Mendoza) 12: 165-167.
Boletaceae and Strobilomycetaceae. Nova Hedwigia 1966a Notes on cyphellaceous fungi. Darwiniana 14:
7: 93-132. 9-18.
1964d Monographs of South American Basidiomycetes, 1966b Macromycetes del Suelo de Bosques incendia-
especially those of the east slope of the Andes and dos, pp. 589-601. In: Piterbarg, R. O., ed., Impacto
Brazil. VIII. Oudemansiellinae, Macrocystidinae, de los incendios naturales sobre los hongos rizopob-
Pseudohiatulinae in South America. Darwiniana 13: ladores en plantulas de Nothofagus dombeyi. Simposio
145-190. de Microbiologia del Suelo, Bahia Blanca, 1966.
1964e Die Gattung Gerronema. Nova Hedwigia 7: 53- 1966c The type of Boletus amabilis. Mycologia 58:
92. 157-159.

1966d Monographs of South American Basidiomycetes, skaylo, E., ed., Mycorrhizae, Proceedings of the First
especially those of the east slope of the Andes and North American Conference on Mycorrhizae. Govern-
Brazil. IX. Tricholoma in Brazil and Argentina. Dar- ment Printing Office, Washington, D. C.
winiana 14: 19-35. 197 Id A
revision of the genus Melanomphalia as a ba-
1967a La Micologia y los recursos naturales renovables. sis for thephylogeny of the Crepidotaceae, pp. 441-
Noticiario mensual Museo Nacional de Historia Nat- 474. In: Petersen, R. H., ed., Evolution of the Higher
ural 11: 7-9. Basidiomycetes. University of Tennessee Press, Knox-
ville.
1967b Nuevos hongos descubiertos en Chile. Biologica
(Santiago) 41: 69-71. 1971e Bayesian analysis of generic relations in Agari-
Some cales. Nova Hedwigia 21: 753-787 (with R. E. Mach-
1967c Agaricales from the Congo. Persoonia 4:
355-377 (with K. Grinling). ol).
1967d Notes sur genre Laccaria. Bulletin de

le la So- 1972a New Basidiomycete from the Antarctic, pp. 179-
180. In: Llano, G. A., ed., Antarctic Terrestrial Biol-
ci6t6 Mycologique de France 83: 104-123.
1967e Die Rb'hrlinge. II. Die Boletoideae und Strobi- ogy. Antarctic Research Series 20.
1972b Notes on some leucosporous and rhodosporous
lomycetaceae (Die Pilze Mitteleuropas, Band VI).
Verlag Klinkhardt, Bad Heilbrunn. 151 pp. European agarics. Nova Hedwigia 23: 305-351 (with
H. Clemengon).
1967f Fungi distribution. Antarctic Map Folio series 5,
1972c Cyanophilous spore walls in the Agaricales and
pp. 16-17. In: Greene, S. W., J. L. Gressis, D. Koob,
G. A. Llano, E. D. Rudolph, R. Singer, W. C. Steere, agaricoid Basidiomycetes. Mycologia 64: 822-829.
and F. C. Ugolini, eds., Terrestrial life of Antarctica. 1973a The genera Marasmiellus, Crepidotus and Sim-
American Geographical Society of New York, 24 pp. ocybe in the Neotropics. Beihefte zur Nova Hedwigia
44: 1-517.
1967g Proposal to conserve the generic name Rhodo-
phyllus Quel [Fungi]. Taxon 16: 242-243. 1973b Notes on bolete taxonomy. Persoonia 7: 313-
1967h Review of: Die unterirdischen Pilze des Karpath- 320.
enbeckens, by L. Szemere. Mycologia 58: 985-986. 1973c The Marasmius-blight fungus. Mycologia 65:
1968a Sand-dune inhabiting fungi of the South Atlantic 468-473 (with L. T. Lucas and T. D. Warren).
coast from Uruguay to Bahfa Blanca. Mycopathologia 1973d Diagnoses fungorum novorum Agaricalium. III.
et Mycologia Applicata 34: 129-143. Beihefte zur Sydowia 7: 1-106.

1968b Review of: Les Russules d'Europe et d'Afrique 1973e Nomenclatorial status of Gomphidius Fries. Tax-
du Nord, by H. Romagnesi. Mycologia 60: 1127- on 22: 445-446.
1130. 1973f Marinus Anton Donk (1908-1972). Mycologia
1969a Mycoflora Australis. Beihefte zur Nova Hedwigia 65: 503-506.
29: 1-405.
1973g Review of: The North American Species of Psa-
1969b Augusto Chaves Batista. Sydowia 22: 343-359. thyrella, by A. H. Smith. Mycologia 65: 975-976.
1969c Alberto Castellanos 1896-1968. Taxon 18: 308- 1973h Using Micheli as a field book. Mcllvainea 1(2):
309. 14-18 (with R. E. Machol).
1970a Flora Neotropica Monograph No. 3. Omphalinae 1973i Review of: Die Grundziige eines natiirlichen Sys-
(C7/r0cy6e-Tricholomataceae, Basidiomycetes). Haf- tems der Pilze, by H. Kreisel. Mycologia 65: 1378-
ner Publishing Co., New York &
London. 84 pp. 1380.
1970b Flora Neotropica Monograph No. 4. Phaeocol- 1974a Notes on Galerina. Bulletin mensuel de la So-
lybia (Cortinariaceae-Basidiomycetes). Hafner Pub- ciete Linneenne de Lyon, numero special 43: 389-
lishing Co., New York &

London. 13 pp. 405.
1970c Monograph No. 5. Strobilo-
Flora Neotropica 1974b A monograph of Favolaschia. Beihefte zur Nova
mycetaceae (Basidiomycetes). Hafner Publishing Co., Hedwigia 50: 1-108.
New York & London. 34 pp. 1975a The Agaricales in Modern Taxonomy. 3rd ed.,
1970d New agarics from South America. Nova Hed- Verlag J. Cramer, Vaduz. 912 pp.
wigia 20: 785-792. 1975b Interesting and new species of Basidiomycetes
1970e The topotype of Boletus amabilis Peck. Mycol- from Ecuador. Beiheft zur Nova Hedwigia 51: 239-
ogia 62: 590-596. 246.
1970f Armillariella mellea. Schweizerische Zeitschrift 1975c The neotropical species of Campanella and
fur Pilzkunde 48: 25-29, 65-69. Aphyllotus with notes on some species of Marasmiel-
lus. Nova Hedwigia 26: 847-896.
197 la Two new ectotroph-forming boletes from India.
Mycopathologia et Mycologia Applicata 43: 25-33 1975d Supplement to "Agaricales in Modern Taxono-
(with B. Singh). my." Nova Hedwigia 26: 435-436.
1971b Neue Arten von Agaricales. Schweizerische Zeit- 1975e Albert Pilat (1903-1974). Mycologia 67: 445-
schrift fur Pilzkunde 49: 118-128 (with H. Clemen- 447.
con). 1975f Tropical Russulaceae: Lactarius sect. Polysphae-
1971c Forest mycology and forest communities in rophori in the Gulf area. Nova Hedwigia 26: 897-
South America. II. Mycorrhiza sociology and fungus 901.
succession in the Nothofagus dombeyi-Austrocedrus 1975g Review of: Hribovite Huby, by A. Pilat and A.
chilensis woods of Patagonia, pp. 204-215. In: Hac- Dermek. Mycologia 67: 440-441.
108 Chapter Four

1975h Review of:
Agarikovye Shlyapochnye Griby 1978a Notes on bolete taxonomy. II. Persoonia 9: 421-
(For. Agaricales)Primorskovo Kraya [Die Blatterpilze 438.
und Rohrlinge (Agaricales) von Primorsky Region], 1978b Omphalia alutacea. Schweizerische Zeitschrift
by L. N. Vassilieva. Mycologia 67: 1076. fur Pilzkunde 56: 154-156.
1976a Marasmieae (Basidiomycetes-Tricholomataceae). 1978c On Calocybe names. Mycotaxon 6: 500-502
Flora Neotropica Monograph No. 17. New York Bo- (with S. A. Redhead).
tanical Garden, Bronx, New York. 347 pp.
1978d Review of: A Preliminary Agaric Flora of East
1976b Le genre Lepista et la nomenclature correcte. Africa, by D. N. Pegler. Mycologia 70: 476-477.
Bulletin de la Societe Mycologique de France 92: 1978e Origins of the deficiency of Amazonian soils-a
127-128. new approach. Acta Amazonica 8: 315-316.
1976c An interesting subspecies of Boletus erythropus 1978f Keys for the identification of the species of Agar-
(Fr. ex Fr.) Krombh. eska Mykologie 30: 1-2 (with icales. II. Sydowia 31: 193-237.
A. Dermek and J. Kuthan).
1978g Hallucinogenic mushrooms, pp. 201-214. In:
1976d LouisSecretan (1758-1839). Schweizerische Rumack, B. H., and E. Salzman, eds., Mushroom Poi-
Zeitschrift fur Pilzkunde 54: 81-86 (with C. Rege).
soning: Diagnosis and Treatment. CRC Press, West
1976e Notes on Chroogomphus (Gomphidiaceae). eska Palm Beach, Florida.
Mykologie 30: 81-89 (with J. Kuthan). 1978h Interesting and new species of Basidiomycetes
1976f An early-fruiting Armillariella. Kew Bulletin 31: from Ecuador. II. Nova Hedwigia 29: 1-98.
433-435. 1979a A new psilocybian species of Copelandia. Jour-
nal of Natural Products 42: 469-474 (with R. A.
1976g Einige interessante europaische Hygrophoraceae.
Zeitschrift fur Pilzkunde 42: 5-13 (with J. Kuthan).
Weeks and W. L. Hearn).
1976h Notes on Boletes. eska Mykologie 30: 143-155 1979b decomposition and ectomycorrhiza in Am-
Litter
azonian forests. Acta Amazonica 9: 25-41 (with I.
(with J. Kuthan).
Araujo).
1976i Agaricales nouvelles des Antilles Fran^aises. Bul-
1979c Hohlenbewohnende Agaricales. Sydowia 32:
letin de la Societe Mycologique de France 92: 445-
299-304 (with U. Passauer).
447 (with J. P. Fiard).
1980a Comparison of some lignicolous white-spored
1976J Amparoinaceae and Montagneaceae. Revue de American agarics with European species. eska My-
Mycologie 40: 57-64.
kologie 34: 57-73 (with J. Kuthan).
1977a Praktische Anwendung der Erkenntnisse iiber die
1980b Review of: North American Species of Lactarius,
ektotrophe Mykorrhiza bei Waldbaumen. Allgemeine
by L. R. Hesler and A. H. Smith. Mycologia 72: 649.
Forstzeitung (Wien) 88: 53-56.
1980c New taxa of Russula from the Lesser Antilles.
1977b A new species of Gastroboletus and its phylo- 12: 92-96 (with D. N.
Mycotaxon Pegler).
genetic significance. Mycologia 69: 59-72 (with E. E.
198 la New genera of Agaricales. Mycologia 73: 500-
Both).
510.
1977c Amerikanische und Asiatische Agaricales, die in
1981b Resinomycena gen. nov. (Agaricales), an ally of
Europa und Nordafrika vorkommen. Zeitschrift fur
Pilzkunde 43: 119-130. Hydropus, Mycena and Baeospora. Mycotaxon 13:
150-170 (with S. A. Redhead).
1977d Die Gruppe der Laccaria laccata (Agaricales).
1981c Illustrations of tropical Agaricales. Nova Hed-
Plant Systematics and Evolution 126: 347-370.
wigia 35: 151-156.
1977e An interpretation of Palaeosderotium. Mycolo- 198 Id Notes on Bolete Taxonomy. HI. Persoonia 2:
gia 69: 850-854. 269-302.
1977f Are Secretan 's fungus names valid? Taxon 26: 1982a Galeropsidaceae west of the Rocky Mountains.
251-255 (with R. E. Machol).
Mycotaxon 14: 82-90 (with P. Ponce de Leon).
1977g Keys for the identification of the species of Agar- 1982b Notes on Russula taxonomy. I. The Russulae of
icales. I. Sydowia 30: 192-279. Catalonia. Collectanea Botanica 13: 669-700.
1977h Controle de vassoura-de-bruxa. Acta Amazonica 1982c North Indian Agaricales. II. Sydowia 35: 236-
7: 570. 241 (with S. S. Saini and N. S. Atri).
1977i The
species concept in Agaricales and its adap- 1982d Basidiomycetes of Costa Rica. I. Brenesia 19/20:
tation totaxonomy, pp. 381-392. In: Clemenson, H., 31-47 (with L. D. Gomez).
ed., Species Concept in Hymenomycetes. Herbette 1982e Hydropus (Basidiomycetes-Tricholomataceae-
Symposium. Bibliotheca Mycologica 61. Myceneae). Flora Neotropica Monograph No. 32.
1977J Review of: A Monographic Study of the Genus New York Botanical Garden, Bronx, New York. 153
Pouzarella, a New Genus in the Rhodophyllaceae, pp.
Agaricales, Basidiomycetes, by S. J. Mazzer. Mycol- 1982f A Correction by Rolf Singer. Botanical Museum
ogia 69: 215-216. of Harvard University 8: 1-5.
1977k Review of: Memoirs of the New York Botanical 1983a Weiss-und rosasporige Agaricales (Tricholoma-
Garden Vol. 28, No. 1. Mycologia 69: 216-217. taceae und Pluteaceae) aus osterreich. Sydowia 36:
19771 Taxonomic position of Hydropus floccipes and 277-287. [published in 1984]
allied species-a quantitative approach. Mycologia 69: 1983b Problemas sistematicos en los Agaricales. Boletin
1162-1172 (with R. E. Machol). de la Sociedad Mexicana de Micologia 18: 13-19.

1983c Acanthocytes in Amparoina and Mycena. Cryp- 1988a Agaricales mil Volva. Zeitschrift fur Mykologie
togamie Mycologie 4: 111-115. 54: 41-44.
1983d The Sydney Rules and the Nomenclature of Rus- 1988b (iber einige Crepidotaceae. Zeitschrift fur My-
sula species. Mycotaxon 18: 191-200 (with R. E. kologie 54: 69-72.
Machol). 1988c The role of fungi in periodically inundated Am-
1983e Sister Mary Cecilia Bodman, 1905-1982. My- azonian forests. Vegetatio 78: 27-30.
cologia 75: 932-933 (with P. Ponce de Le6n). 1988d Notes on Conocybe (Bolbitiaceae). Plant System-
1983f The ectotrophically mycorrhizal fungi of the neo- atics and Evolution 159: 107-121 (with A. Haus-
tropical lowlands, especially central Amazonia. Beih- knecht).
efte zur Nova Hedwigia 77: 1-352 (with I. Araujo and
1988e Notes on Neotropical Austroboletus species. My-
M. H. Ivory).
cologia 80: 46-53 (with C. B. Wolfe and R. Walsh).
1984a Tropical Russulaceae. II. Lactarius section Pan-
1988f The phytogeography of the Boletineae (Basidio-
uoidei. Nova Hedwigia 40: 435-447.
mycetes, Agaricales) based on Mexican species. Re-
1984b The Amazonian forests and in
role of fungi in vista Mexicana de Micologia 4: 267-274.
603-614. In: Sioli, H., ed., The Am-
reforestation, pp.
azon. Limnology and Landscape Ecology of a Mighty 1988g Laccaria gomezii, a new agaric species from the
querceta of Colombia and Costa Rica. Mycotaxon 33:
Tropical River and Its Basin. Dr. W. Junk Publishers, 223-227 (with G. M. Mueller).
Dordrecht, Boston, Lancaster.
1984c Adaptation of higher fungi to Varzea conditions.
1988h How to become a mycologist. III.Mutualistic
Amazoniana 8: 31 1-319.
interactions. Mcllvainea 8(2): 10-11 (with G. M.
Mueller).
1984d Agaricus brunnescens Peck and Agaricus bis-
1988i New tropical species in the Paxillaceae. Mycol-
porus (Lange) Imbach. Mycotaxon 20: 479-482.
ogia Helvetica 3: 207-213 (with D. J. Lodge).
1984e Three proposals to amend the Code. Taxon 33:
745-747. 1989a New taxa and new combinations of Agaricales
1984f Veloporphyrellus, a new genus of Boletaceae (Diagnoses Fungorum Novorum Agaricalium IV).
Fieldiana: Botany, n.s., 21: 1-133.
from Costa Rica. Brenesia 22: 293-298 (with L. D.
Gomez). 1989b Agaricales from Henderson Islands (South Pacif-
ic). Mycologia 81: 793-796.
1984g The Basidiomycetes of Costa Rica. HI. The ge-
nus Phylloporus (Boletaceae). Brenesia 22: 163-181 1989c Conocybe hornana-eine neue Art mit Volva aus
(with L. D. Gomez). osterreich (mit einer Farbtafel, siehe bei Seite 80).
1985 Studies on Secotiaceous Fungi. I. A new species Beitrage zur Kenntnis der Pilze Mitteleuropas 5: 87-
of Cystangium. Mycologia Helvetica 1: 417-425. 91 (with A. Hausknecht).
1986a The Agaricales in Modern Taxonomy, 4th ed. 1989d Reflections on Mycorrhizal Research, pp. 625-
Koeltz Scientific Books, Koenigstein, Federal Repub- 627. In: Stirton, C. H., and J. L. Zarucchi, eds., Ad-
lic of Germany. 981 pp.
vances in Legume Biology. Monographs in System-
atic Botany from the Missouri Botanical Garden 29.
1986b The nomenclature of Russula species and the Syd-
ney Rules. Persoonia 13: 189-192 (with R. Machol).
1989e Two interesting fungi of the Western Hemisphere.
The Mycologist 3: 70-71 (with J. Garcia).
1986c Litter decomposing and ectomycorrhizal basidi-
omycetes in an Igapo Forest. Plant Systematics and 1989f Review of: Pareys Buch der Pilze, by M. Bon.
Evolution 153: 107-117 (with I. A. Aguiar). Mycologia 81: 826.
1986d How to become a mycologist. I. The conserva- 1990a The Boletineae of Mexico and Central America.
tion of material and notes. Mcllvainea I and II. Beihefte zur Nova Hedwigia 98. 70 pp. (with
7(2): 18-23.
J. Garcia and L. D. Gomez).
1986e A new species of Psilocybe (Agaricales). Plant
Systematics and Evolution 151: 295-301 (with A. 1990b Clitocybe obsoleta. Zeitschrift fur Mykologie 56:
Hausknecht). 5-7.
1986f Basidiomyceten-Nomenklatur im kommenden 1990c New species of Phylloporus and Tricholomopsis
Internationalen Botanischen Kongress, Berlin, Juli from Colombia, with notes on Phylloporus boletino-
1987. Zeitschrift fur Mykologie 52: 439-443. ides. Mycologia 82: 452-459 (with C. L. Ovrebo and
1987a Mushrooms and Truffles, 2nd ed. Koeltz Scien- R. E. Hailing).

tific Books, Koenigstein, Federal
Republic of Ger- 1990d Some interesting agarics from eastern Austria.
many. 389 pp (with B. Harris). Plant Systematics and Evolution 170: 133-150 (with
1987b Phaeocollybia in the oak woods of Costa Rica, A. Hausknecht).
with notes on extralimital taxa. Mycologia Helvetica 1990e Agaricales new for Mexico or Central America. An-
2: 247-266. ales del Instituto de Biologia de la Universidad Nacional
1987c Mycena theobromicola, a fungus associated with de Autonoma de Mexico, Sen Bot. 60(1): 27-36.
cacao pod rot. Mycologia 79: 463-464. 1990f Upper Eocene gilled mushroom from the Domin-
1987d The starting point rules and stability of nomencla- ican Republic. Science 248: 1099-1101 (with G. O.
ture. Plant Systematics and Evolution 156: 177-181. Poinar, Jr.)
1987e How to become a mycologist. II. The teaching 1990g A new species of Melanomphalia (Crepidota-
of mycology, especially Agaricales. Mcllvainea 8(1): ceae). Mycotaxon 37: 47-52 (with C. L. Ovrebo and
22-26. D. P. Lewis).
110 Chapter Four

199 la The Boletineae of Mexico and Central America. 19925 The group of Conocybe mesospora in Europe
III. Beihefte zur Nova Hedwigia 102. 97 pp. (with J. (Bolbitiaceae). Plant Systematics and Evolution 180:
Garcia and L. D. Gomez). 77-104 (with A. Hausknecht).
1992c Some boletes from Florida. Mycologia 84: 724-
1 99 1 b A newspecies of Marasmius from northwestern 728 (with R. Williams).
India. Nova Hedwigia 52(1-2): 227-230 (with A. Ku-
1994 Toward a definition of the genus in mycological
mar, T. N. Lakhanpal and S. L. Stephenson).
taxonomy, pp. 92-94. In: Clemen^on, H., ed., Generic
1992a The Boletineae of Mexico and Central America. Concepts in Mycology. Proceedings of the Second
IV. Beihefte zur Nova Hedwigia 105. 62 pp. (with J. Herbette Symposium in Lausanne, Switzerland, 1991.
Garcia and L. D. Gomez). Mycologia Helvetica 6.
The Publications of Rolf Singer 1 1 1

Index to Genera
Acurtis 13 Cystoderma 28 Lachnella 48

Aeruginospora 13 Cystolepiota 5, 28-29, 52 Lachnocladium 48
Agaricus 5, 6, 13, 18-19, 40, 52, Lactarius 49-50
75, 84, 87-88, 94 Daedalea 5, 29 Lactocollybia 5, 49, 50
Agrocybe 13-14 Deconica 29 Lampteromyces 5, 49
Alnicola 14 29 Leccinum 49
Dennisiomyces 5,
Amanita 14 Dermoloma 29 Lentinellus 50
Amparoina 5, 14 Descolea 5, 29 Lentinus 6, 50, 98
Amyloflagellula 14 Diacanthodes 5, 29
5,
Lepiota 5, 41, 50, 52
Anthracophyllum 14 Dictyopanus 29 Lepista 50
Aphyllotus 14
5,
Leptoglossum 50
Arcangeliella 14 Eccilia 30 Leptonia 50
Armillariella 5, 14-15, 75
Elasmomyces 30 Leucoagaricus 5, 50-51
Arthrosporella 5, 15 Endolepiotula 5, 30 Leucocoprinus 5, 50, 51
Asterotus 5, 15
Endoptychum 30 Leucoinocybe 5, 51
Austroboletus 15 Entoloma 30 94
Leuocomyces 6,
Austrogaster 5, 15 30
Epicnaphus 5, Leucopaxillus 51
Limacella 5 1
Baeospora 5, 15 Favolaschia 30-31 Linderomyces 5, 51
Bolbitius 16 Fayodia 31 Lulesia 5, 51
Boletellus 16 Fistulinella 31 Lyophyllum 5 1
Boletinus 16 Flammula 31-32
Boletochaete 5, 16 Fulvidula 32 Macowanites 51-52
Boletus 5, 16-17, 77 Macrocystidia 52
Bondarzewia 5, 17 Galerina 3, 32-37 Macrolepiota 5, 52
Brauniellula 5, 17 Galeropsis 37 Macrometrula 5, 52
Gastroboletus 37 Manuripia 5, 52
Calathinus 17 Gerronema 5, 37-38 Marasmiellus 3, 29, 52-57, 61, 72,
Callistodermatium 5, 17 Gloeocantharellus 5, 38 74
Callistosporium 5, 17 Gloiocephala 38-39 Marasmius 3, 5, 6, 14, 57-67, 76,
Calocybe 17-18 Gomphidius 39 90
Calyptella 18 Gomphus 39 Martellia 67
Camarophyllus 18 Grifola 39 Melanoleuca 67-68
Campanella 18 Gymnomyces 39 Melanomphalia 68
Cantharellula 5, 18 Gymnopilus 5, 17, 39-40 Melanotus 68-69
Cantharellus 5, 18, 38 Gyrodon 40 Merulinus 6, 88
Chaetocalathus 5, 18 Gyroporus 6, 40, 98 Micromphale 69
Chaetotyphula 19 Micropsalliota 69
Chamaemyces 19 Hebeloma 40 Mniopetalum 69
Chamonixia 19 Heimiomyces 5, 40 Mycena 3, 5, 51, 54, 69-73
Cheimonophyllum 5, 19 Hemimycena 40-41 Mycenella 5, 73-74
Chlorophyllum 19 Hericium 41 Mycoalvimia 5, 74
Chroogomphus 19 Hiatula 41 Myxocollybia 74
Clavaria 19 Hiatulopsis 5, 41
Clavulina 19 Hohenbuehelia 41 Naematoloma 74
Clitocybe 3, 19-21 Hybogaster 5, 41 Naucoria 74
Clitocybula 21 Hydropus 42-44
3, Neoclitocybe 5, 53, 74-75
Clitopilus 21 Hygroaster 5, 44 Neohygrophorus 5, 75
Collybia 5, 6, 15, 21-22, 40, 49, 76, Hygrocybe 45 Neopaxillus 5, 75
80 Hygrophoropsis 45 Neosecotium 5, 75
Conocybe 3, 22-23 Hygrophorus 5, 44, 45, 75 Nivatogastrium 5, 75
Copelandia 23 Hygrotrama 5, 45-46 Nolanea 5, 28, 75
Coprinites 5, 23 Hymenogaster 46 Nothoclavulina 5, 75
Coprinus 23-24 Hypsizygus 5, 46 Nothopanus 5, 75
Coriolus 24
Cortinarius 24 Inocybe 46-47 Octavianina 75
Craterellus 24 Ixocomus 47 Omphalia 5, 74, 75
Crepidotus 24-26, 80, 87
3, 6, Omphalina 19, 20, 75-76
26-28
Crinipellis 3, Krombholzia 47 Oudemansiella 76
Cyptotrama 5, 28 Kuehneromyces 5, 47
Cystangium 5, 28 Pachylepyrium 5, 76
Cystoagaricus 5, 28 Laccaria 47-48 76
Palaeocephala 5,
Index to Genera 113

Panaeolopsis 5, 76 Poromycena 84 Setchelliogaster 93
Panaeolus 76 Porphyrellus 84 Simocybe 94
Panellus 76 Porpoloma 6, 84 Skepperiella 94
Panus 5, 15, 76 Psathyra 84 Smithiomyces 6, 94
PaxillusS, 51, 76-77, 80 Psathyrella 85-86 Stigmatolemma 94
Pegleromyces 5, 77 Pseudobaeospora 86 Strobilomyces 94
Pseudoclitocybe 86 Strobilurus 6, 94
Phaeocollybia 77
Pseudohiatula 86 Stropharia 94
Phaeodepas 77
Phaeogyroporus 5, 77 Pseudoomphalina 86 Suillus 95
Phaeomarasmius 5, 78 Psiloboletinus 6, 86
Psilocybe 86-87 Tectella 95
Phaeomycena 78
Phellinites 5, 78 Pterula 6, 93 Thaxterogaster6, 95
Pulveroboletus 87 Tricholoma 95-96
Phlebopus 78
Pholiota 5, 78-79 Pyrrhoglossum 6, 87-! Tricholomopsis 96
Tubaria 96
Pholiotina 79
Radiigera 88 Tylopilus 96-97
Phylloboletellus 5, 79
80 Rajapa 6, 88
Phyllobolites 5, Ramaria 88 Veloporphyrellus 6, 97
Phylloporus 6, 80, 86 88 Volvariella 97
Resinomycena 6,
Physocystidium 5, 80
Resupinatus 88 Volvolepiota 97
Pistillaria 80
Rhodoarrhenia 6, 88
Pleurocollybia 80 6,
Rhodocybe 88-89 Weraroa 6, 97
Pleurocybella 6, 80
Rhodophyllus 89-90
Pleuroflammula 6, 80 Rimbachia 90 Xanthochrous 6, 98
Pleuromycenula 6, 80 Ripartitella 6, 90 Xanthoconium 98
Pleurotellus 80 Xerocomus 98
Ripartites 90
Pleurotus 5, 6, 80, 81 Russula 3, 90-93 Xeromphalina 98
Pluteus 3, 81-84 Xerula 98
Podabrella 6, 84 Scutiger 93 Xerulina 6, 98
Polyplocium 84 Scytinopogon 6, 93
Polyporus 5, 17 Secotium 5, 6, 28, 75, Zelleromyces 6, 98-99
1 14 Index to Genera
Index to Species
aberrans 18, 24, 32, 48, 61, 74, 92 allocystis 13, 57 apora 85
aberrantissima 19 allomorpha 38 aporpohyphes 57
abieticola 69 allospora 32 aporposeptus 52
abietina 41 alnea 78 aporpus 42, 57, 81, 89
acaciarum 50 alneti 17, 94 aposcenum 29
acadiensis 88 alnetorum 18, 29, 50 appalachiensis 90
acanthosyrinus 24 alneus 52 applanatus 24
acicola 19, 21, 32 alni 48 aprilis 19
aconquijensis 19, 69 alpestris 32, 86 aquilus 52
acris 51 alpina 32 aquosus 44, 81
acutesquamosus 96 alpinus 57 araguense 46
acutissima 85 alsophilae 30 araucana 45, 53, 55, 62, 86
acutiuscula 29 altaica 47, 79, 91 araucariae 57, 96
adisianus 40, 57 altaicum 95 araucariicola 81
adonis 69 altoandina 79 araujae 69
aegerita 13 altus 42 arcticus 24
aequatoriae 86 alutacea 88, 90 arcyospora 90
aequatorialis 18, 49, 57, 69 alutaceum 37 arenaria 32
aeruginea 18 aluteacea 90 arenarius 96
aeruginosa 94 alvaradoi 38, 52 argentea 46
aestivalis 22, 94-95 alveolatus 39 argentinensis 81
affinis 14, 19, 47 amara 5, 41, 47 argentinus 15
africana 17, 52 amarescens 75 argillospora 85
africanus 42 amazonaicus 81 argipoda 68
aggericola 86 amazonensis 21, 48 aristoteliae 24, 73
agloea 32, 69 amazonica 22, 29, 40-41, 52, 69, arsitophylli 86
agriensis 81 77, 94 asema 19
agrocybeoides 96 amazonicum 17, 37, 74 asemiformis 57
agrocybiformis 78 amazonicus 76, 98 asemus 57
aimara 32, 57 amazoniensis 29, 45-46, 52 asiaticus 16
alba 22 ambiguus 52 aspilocephalus 57
albella 67 americanus 51, 59 asteliae 32
albelloavellanea 21 amianthinum 28 asterophora 69
albellum 49 ammophila 45 ater 57
albellus 44 amnicola 90 atlanticus 57
albertianus 57 amparae 90 atriavellaneus 81
albida 84, 97 amphicystis 52, 81 atrofibrillosus 50, 84
albidigleba 5 1 ampullicystis 33 atroincrustata 69
albidobisa 19 amylochaete 19 atroincrustatus 57
albidomurinus 57 amyloidea 50 atropapillata 17
albidus 24 ananas 16 atropurpurea 90
albifolius52 anastomosans 38, 84 atrorubens 57
albipes 26, 45 andina 30, 32, 37, 94 attenuata 77
albissimus 51 androsaceiformis 52 aucarum 90
albobrunnescens 52 anelligera 32 augustinensis 19
albocinerea 22 angelesianus 5, 75 aurantiaca 29-30, 45, 49
albocremea 88 angiospermarum 49 aurantiacus 19
albogilva 28 angusta 40 aurantioalbida 78
albogriseolum 37 angustifolia 32 aurantiomfescens 52
alboniger 88 angustifolium 37 aureoflava 41
albonigroides 90 angustisperma 19 australe 28
albopubescens 42 angustispermus 42, 52 australis 24, 29, 48, 76, 79,
albostipitatus 81 angustispora 88 austrinus 16

albovelutina 94 angustisporus 81 austroalcalina 69
albus 42, 62 annulifer 48 austroanatinus 89
alcalivirens26 annulosa 85 austroandina 32, 98
alexandri 37 antarctica 5, 29, 32, 35, 38, 75 austroandinus 25
alfaroae 31 antarcticus 13, 52 austroavenacea 69
algeriensis 69 aosma 69, 90 austrocalyptrata 32
alkalivirens 21 apatelius 57 austrocapillaris 69
alliodora 19 apoda 80 austrochilense 69
allochrous 48 apogonus 57 austrochilensis 18
Index to Species 1 1 5
austrodelica 90 brunneomarginatus 53 caucasicum 28, 41
austrofallax 28 brunneosetosa 16 caucasicus 58
austrofilaris 79 brunneosperma 50 cavipes 42
austromontana 90 brunneotingens 29 cecropiae 48
austrororida 69 brunnescens 67, 88 cedretorum 33
austrorotula 58 brunneum 37 cellulosus 96
austrorubida 26 bryogeton 37 cenospilus 58
aztecus 58 bullulifer 25 centroamericanus 58, 80
azurea 21, 50 bullulifera 32, 41, 47 cerasphora 46
burserae 81 cerina 33
bactrosporus 58 buzungolo 58 cernohorskyi 90

baeocephalus 58 byssiseda 5, 74 cervinus 81
baeocystis 86 chalybaeum 49
baeosperma 78 caballeroi 80 chamaeleon 90
baeospermum 15 caerulipes 86 chapinii 98
baeospora 68, 75 caesioannulata 86 cheilocystidiatum 5, 37
baeosporoides 52 caesiocana 70 chiapasensis 58, 95
baeosporus 52 caespitosus 42 chilensis 16, 21, 39, 45-46, 48, 71,
bakshii 98 calendulina 78 78-79, 86, 88
bambusiniformis 58 calhouniae 58 chiloensis 85
barlae 90 caliensis 58 chiriquiensis 42
bathomphalus 58 californica 32, 67 chloephorus 79
batistae 21, 45 californicus 39, 42, 53 chlorinodorus 42
bauhiniae 53 callistosporoides 86 chlorinosma 70
beniensis 42, 58, 77, 81 calocera 30 chlorocystis 23
berkeleyi 53 calolepis 25 chloroxantha 70
berteroana 32 calongei 37, 41 chocoruensis 80
beschidica 86 calospora 47 chrysoblepharis 58
bethlehemicum 37 calyciformis 50 chrysocephalus 58
bezerrae 58 camaragibensis 42 chrysocorypha 70
bibulum 95 campanelliformis 5, 14 chrysocystidiata 86
bifida 5, 52 camphoratus 30 chrysopeplus 6, 98
bingaensis 58 campinaranae 24, 31, 42, 45, 58, 98 chrysophlebius 81
biornata 69 campinensis 48 chrysophorum 17
bipigmentata 17 Candida 18 chusqueophila 70
bisphaerigera 31 candidipes 86 cibaria 80
bisporiger 44, 53 candidissimus 5, 19 cibarius 58
bisporum 69 candidum 37 ciliata 74
bisporus 24, 42, 88 cantharelluloides 89 ciliatus 77

blechni 78 capillata 38 cinerellus 81
blechnophila 70 caracassensis 53 cinereofusca 29
blestiana 45 carbonaria 86 cinereus 53
boletinoides 76-77 carbonicola 23, 48 cingulata 33
bolivarianus 53 carcharus 58 cinnabarinus 6, 98, 99
boliviae 24, 58 carecomoeis 26 cinnamomea 5, 33, 38, 80
boliviana 31-32 carelica 46 cinnamomeopallida 69
bolivianus 42, 82 caricicola 79 cinnamomeus 39
bonairensis 18 carminis 58 circularis 80
borealis 32 carnelioruber 19 circumscissus 81
boyacensis 24, 90 carneotinctus 58 citricolor 25
brachypus 96 carpenterianus 58 citrina 90
brachysporus 58 carpophilus 58 citrinopileatus 81
brasiliensis 54, 76-78, 90, 93 cascadensis 32 citrinus 42, 51
braunii 5, 77 castaneiceps 47 cladophyllus 59
bresadoliano-affinis 19 castaneipes 18, 32 clavuligera 15
brevibasidiata 19 castaneomarginata 70 clitocybe 53, 74
brevicolumna 19 castaneostipitata 70 cnacopolius 53
brevipes 95 castanescens 32 cnaphocystis 70
brevispora 67 castellanoi 58 coccineus 16
brevisporus 68, 95 castilloi 74 cocosensis 53
bridgesiana 46 catalaunica 20, 46, 50, 67 codinae 84
bromeliacearum 58 catamarcae 25 cohaerens 59
brunnea 46 catamarcensis 26, 56 cohortalis 59
brunneipes 40 caucae 48 collybiiformis 50
brunneobasis 42 caucaensis 90 collybioides 5, 77, 86, 88
brunneocinctus 58 caucasica 17, 20, 46, 93 collybiomorphum 37
1 1 6 Index to Species
coluiana 20 cystidiatus 76 dusenii 31, 42
columbiana 19, 33, 68, 75, 77, 90 cystidiosa 20 dussii 6, 80
columbianus 53 cystidiosus 42, 47, 53 dwyeri 53
commixta 26-27 dysodes 59
compactus 39 dactylocystis 85
confertifolius 42, 53 dadmunii 91 echinata 30
confragosuliformis 77 daguae 53 echinocephalus 53
confragosulus 77 daguense 37 echinosperma 76
confusa 38 dauliporpus 59 echinosphaerus 59
confusus 94 dealbata 30 echinosporus 5, 75
conica 31, 88 dealbatus 5, 15 edite 53
coniceps 17 debiliformis 70 eggersii 27
coniferarum 94 decapitatus 68 eitenianus 42
conigenoides 94 6, decipiens 25, 33, 42, 92 elaeocephalus 59
conquistensis 59, 83 defibulata 75, 86 elaeophylla 33, 77
conspicua 33 defibulatus 25, 53, 77, 82, 89 elasticum 37
constricta 5, 28-29 degiustoi 5, 78 elatipes 96
convexulus 76 delasotae 41 elegans 96
convoluticeps 59 delica 70, 91 eliae 82
coprophila 14 dendroegrus 53 ellipsoidea 6, 80

coquimbensis 33, 81 dendrophora 18 ellipsoidosporus 59
cordobensis 86 dennisianum 5, 45-46 elongatispora 18
corner! 42, 51, 96 dennisii 28, 37, 59, 82 elongatisporus 53
coroicae 27 densifolia 5, 15, 51 elvirae 95
coroicensis 94 depauperata 40, 76 emarginata 68
corrugatiformis 59 depauperatum 28, 37 embrapaensis 21
corrugatus 59 depauperatus 42, 47 emetica 91
corticalis 94 depigmentata 91 emmetensis 33-34
corticum 53 depilata 70 endoglobulosa 70
cortinarioides 68 depressum 30 enodis 53
cortinatellum 95 desfontaineae 70 eorotula 59
cortinatum 95 deusta 31 epelaeus 59
coryleti 29 devenulatus 53 epibiotica 40
corynophloeus 53 devia 14 epibrya 18, 40
costaricense 37 dicotyledonum 27 epigloeus 25
costaricensis 49, 59, 70, 91, 96 dicranorum 33 epipolia 84
cotapatae 37 diemii 14, 22, 50, 95 episemus 59
cothurnatus 95 59
digilioi epitheliosa 91
couleu 53 dilutecinnamomea 46 epitrichialis 53
crassa 14 dimorphocystis 33, 41 equicrinius 60
crassotunicata 67, 91 dinae 70 erythrobasis 46
craterellus 5 diplocystis 14, 18, 40 esculentus 94
cremeo-avellanea 91 dipterocarpi 27 espeletiae 41, 82
cremeospora 67 diptychocystis 82 espinosae 20
crepidotoides 88 discernibilis 33 etenianus 42
crinipellis 85 discogena 70 eucalypticola 25
crinipelloides 53, 89 dissiliens 42 eucladopus 60
crinis-equi 59 dissimilis 89 eucryphiae 82
crispuliformis 40 dissocystis 46 eucryphiarum 70
cristulatus 25 distoriformis 53 eugeniae 54
cristulispora 91 ditopa 5, 15, 75 eugrammus 5, 75
cryptochrous 89 ditopotrama 59 euosmus 60
cryptotrichus 59 dodecaphyllus 59 eurhizus 6, 88
cuatrecasasii 59 dombeyi 95 eurysporus 19
cuimisedus 59 dominicana 5, 23 eutheles 46
culmicola 38, 41 dominicanus 82 evelata 34
cundinamarcae 54, 59 dominici 33 exalbicans 91
curvipes 74 dragonosporus 89 exannulatus 50
cuticularis 98 drepanocladi 40 excedens 22
cyanea 17 dryinus 59 excentriciformis 39
cyanella 17, 70 dryogeton 53 excentricus 42
cyanocephala 70 dryopteridis 70 exiguus 16, 40, 60
cyanosyringea 70 dubius 42, 97 exilis 20, 82
cyanoxantha 91 duckeana 20
cyatheae 59, 73 duckeanus 87 fagnani 95
cymatelloides 5, 76 dumontii 30, 57, 68, 70, 76 fallax 16, 28, 34, 67, 82

farlowii 89 fuscomycelinus 43 guzmanii 25, 54
fascicularis 85 fuscoviolaceus 52 gymnocystis 80
fascifer 37 fusicystis 60
fastigiatus 82 fusisporum 95 haediniformis 60
favoloides 81 haedinopsis 86
favrei 17 galeriniformis 24 haematocephalus 60-61
februaria 70, 86 galerinoides 17, 47, 89 haematopoda 71
fendleri 30 galeropsidoides 27 halophila 22
fernandeziana 70 gamundiae 34 handelii 91
fernandezianus 24, 82 gargal 39 harperi 25
ferreri 91 geesterani 81 hawaiiensis 89
ferrotincta 91 gelatinascens 91 haywardii 82
ferruginascens 39 geophyllomorpha 46 heimii 21
ferrugineogranulatus 89 geotropoides 50 helbergeri 98
ferrugineus 60 gerronematoides 43 heliconiae 30
festivus 84 gigacystis 46 helminthobasis 71
fibrillosa 17 giganteus 5, 41 helminthocystis 54
fibrosissima 50 gigantospora 18 helveticus 39
fibulatus 80, 82 gigantosporus 54 helvoloides 61
fibuliger 16 gigasporus 24, 49 hemimycena 61
filaris 79 gilkeyae 99 hemitrichialis 71
filiformis 33-34 gilvoides 54, 89 hemixanthus 98
filopes 30 gilvus 54, 78 hendecaphyllus 61
fimbriata 20 glabrescentipes 5, 29 hendersoniensis 82
fimbriatus 42 glabripes 17, 43 herbarum 71
firma 14 glacialis 34 herbivorus 23
flammeum 37 glaucoalbum 95 hericiiforme 48
flammuliformis 87 glaucus 82 heroica 71
flava 18, 77, 80 globiger 82 herrerae 27, 40
flavescens 86 globulifera 68 hesleriaffinis85
flavomycelina 21 globuliger 98 heteracantha 14
flavornatus 24 gloeoloma 70 heterocheilus 61
flavus 60, 98 glutinigera 78 heterochroa 91
flocculosipes 94 glutinocothurnata 70 heterocystis 25, 43
floridana 18-19 glutinosa 76 heterodermus 16
floridanus 16, 81 glutinosus 82 heteroloma 68
flotowiophilus 60 gomezii 40, 43, 47, 50, 60, 70 heterospora 17
fluminensis 80, 82 gomezpompae 54 heterotrama 71
fluvialis 42-43 graciliaffinis 15 hexaphyllus 61
foetens 14, 67, 91 gracilis 27, 43, 91 hiemalis 13, 82, 95
foliicola 27, 54, 60, 72 gracillimus 51 hieronymi 18
fontqueri 91 graminicola 49 hiorami 61
fragile 95 graminum 15, 60 hirtellus 95
fragilior 43 grandisetulosus 60 hobsonii 21
fraterniger 43 granulatus 95 hoehnelii 91
fraternus 89 granulosiceps 16 hololeuca 45
freindlingiae 31 granulosum 28 hololeucus 82, 89
frostii 16 gregarius 77 homomorpha 46
fuegiana 21, 34, 91 grinlingii 18 hondurensis 96
fuligineorotula 60 grisea 31, 51 hongoi 82, 91
fuligineotomentosus 16 griseobrunnea 88 hornana 22
fuliginosa 73 griseofuscescens 60 horridulus 61
fulvescens 5 1 griseolobrunneolus 54 horridus 82
fulvidula 5, 77 griseomellea 15 humboldtii 14, 91, 96
funariae 34 griseoradiata 70 humicola 39
funebris 73 griseoradiatella 7 1 humilior 43
furfuracea 96 griseorete 71 hyacinthina 20
furfurella 30 griseum 69 hyalinotricha 71
fusca 91 griseus 16 hyalinotrichus 61
fuscata 46, 9 1 guadelupae 16 hyalinum 94
fuscimarginata 22 guanacastensis 40, 49, 60 hydrogeton 41
fuscoalbus 29, 43 guayarensis 91 hydrophoroides 43
fuscocinnamomea 46 guttatus 77 hydropodoides 61, 71
fuscocystidiata 70 guyanensis 60 hygrocyboides 28
fuscogriseus 43 guzmaniana 86 hygrophilus 43
fuscomarginatus 77 guzmanianus 60 hygrophoroides 76
1 1 8 Index to Species
hylaeae 23, 61 kabulensis 20 leucozonitiformis 62
hylaeicola 61, 82 karelica 14 levis 43
hymenocephala 68 karstenii 34 levisporus 25, 43
hyperythrus 43 katangensis 61 liberiensis 91
hypnorum 34 keissleri 82 lifotama 74

hypocarycinus 16 kermesina 71 lignatilis 85
hypolepargum 38 kerrii 24 lignicola 14, 89
hypolissus 54 kisangensis 61 lilaceipes 88
hypopolius 43 kivaczensis 24 lilaceus 51
hypoxanthus 98 krieglsteineri25 lilacinoalbus 62

hypsizyga 34, 71 kubickae 78 lilloi 14
hypsizygus 24, 61 kuhneri 71 limulatellus 78
kuhneriana 22, 71 linderi 16, 62, 98
ianthinocystis 89 kuthanii 71, 82 lindquistii 21, 81
icterinoides 71 linicolor 76
icterinus 25, 54 laccarioides 18, 38 lipocystis 43
idahoensis 67 laccata 47-48 liquidambari 62
idroboi 54, 61, 71, 91 lacrimans 71 lithocarpi-sequoiae 95
igapoensis 25, 49, 71 lacrimosa 5, 49 littoralis 78
iguazuensis 57, 61, 63, 82 lactariiformis 89 lividobrunneus 96
ilicis 61 lactarioides 45 lobauensis 22
illinorum 54 lacteoniger 89 lodgeae 21
illudens 98 lacuster 43 lohwagii 71
immutabilis 43 laeta 34, 89 lomatiae 62
incarnata 52 laetifrons 82 longicrinita 38
incarnatipallens 54 laetissima 87 longicystis 40
inconspicua 21, 34 laetus 82 longifimbriata 38
incrustatus 21, 54 laevissima 85 longinqua 34, 87
indica 52, 71 lamellosa 38 longisperma 39
indicus 43, 98 langei 51 longispora 67
inedulis 88 lariceti 6, 80 louisii 62
infernalis34 laricicola 20 lowyi 89
infumata 75 laricinus 37, 49 luridus 25
infundibuliforme 38 larigna 34 lutea 39
infundibuliformis 18 lateritiorosella 45 29
luteifolia
inobasis 38 lateritius 5, 51 luteofuscum 17
inocyboides 68 latifolius 25 luteolosperma 34, 68
inodermatoides 54 latispermus 54 lutescens 46
insignis 14, 27 latispora 34, 53, 74 lutescentipes 43
intermedia 30, 34 latisporum 69 lyophylliformis 89
intervenosa 71 latiusculospermus 61 lysochlorus 54
inundabilis 98 laurifoliae 54
iodactylus 61 lazoi 14 maackiae 32, 78
ionocephala 71 lecythidacearum 54, 71 maasii 62
iopus 27 leguminosarum 61, 91 machangaraensis 34
irazuensis 78 lenta 5, 51 mackleri 68
isauri 87 lenticeps 32 macquariensis 34, 85
ivoryi 16 lentispora 22 macracantha 76
ixoleuca 71 leoninus 61-62 macrobasidium 5, 28
ixotrama 21 lepiotospora 97 macrocystidiata 7 1
ixoxantha 71 leporina 22 macrolobieti 62

izonetae 22, 50 leptocystis39 macropoda 92
izonetas 61 leptosarcum 38 macrorhizus 5, 50
leptus 62 macrosphaerigera 27
jaapii 34 leucobryicola 34 macrospora 74, 97
jacuticus 47 leucocephala 68 macrosporum 5, 69, 75,
jalapensis 96 leucocyanescens 82 macrosporus 21, 47

janauariensis 61, 85 leucomodesta 91 maculata 67, 92
janthinosarx 32 leucomycelinus 80, 87 maculatipes 43
japonicus 5, 49 leucophyllus 54 maculosipes 82
javanica 22 leucoporphyria 19 magellanicum 6, 95
josserandii 38 leucopus 46 magna 94
juaniicola 71 leucorotalis 62 magnisetulosus 62
jujuyensis 64 leucosarx 17 magnispora 22
juniperinus 54 leucoxantha 71 magnoliae 62
junquillea 94 leucozonites 64 mainsii 30, 34

mairei 92 microleucus 81 nanella 92
maireiaffinis 79 microloxus 40 nanoclavula 72
majalis 34 micromelaena 72 nanorotalis 63
major 17-18, 46, 48, 83-84, 92 micromphale 72 nanus 44

majus 38 micropolia 72 napoensis 44, 55, 63
majuscula 80 microrotalis 62 naucoria 89
mallophora 92 microsperma 22 naucorioides 35, 79
malvacearum 78 microspora 14, 41, 49, 74, 79, 87, nebularum 13, 33, 63, 78
mamorensis 39, 49 89, 92 neglectus 63
manausensis 80 microsporus 43, 98 neoantipus 23
manuripiensis 62 microstena 72 neodiscopoda 92
marasmioides 43, 89 microtephra 72 neophlebophorus 83
marchantiae 38 microtrichialis 72 neorotula 63
margaritispora 5, 73 microxantha 72 neosessilis 57, 61, 63
mariae 38, 92 miniatus 5, 80 neospeirea 72
martensii 43 minima 23, 74 neotropica 21
marthae 5, 15, 29, 31, 34, 62, 79, minor 16, 25, 35, 40, 44, 81, 83, 85, neotropicalis 15, 46, 54, 72
85, 87, 90 91, 93, 96-97 neotropicus 49
martini 25 minusculus 54, 62 nephelogenes 63
martipanis 86 minuta 15 nephrodes 25
matrisdei 46, 62, 68 minutalis 54 neuquenensis 46, 50, 94
mazatecorum 22, 68, 87 minutissimum 38 nicaraguensis 97
mazatecus 62 minutula 92 nigra 41
megalobasidium 43 minutuliaffinis 72 nigripes 35
megalocystis 34 minutulus 51 nigromarginatus 44
megalosperma 79 minutus 25, 54 nigropallescens 83
megalospermus 62 mirabilis 27, 76 nitens 89
megalospora 22, 27, 77 misionensis 54, 63 92
nitida
megalosporum 69 missionensis 27, 83 niveum 28

megasporus 24 missionum 23 nobilior 21
megistospora 45 mitissima 92 nobilis 88
megistosporus 62 mitissimus 97 nodulisporus 5, 44

melandeta 71 mixta 23, 34 nodulosporum 51
melanoloma 41 mixtiliformis 46 nogalesii 63
melanomphax 5, 37-38 modesta 49 nondorbingi 92
melanopotamicus 83 moellerianus 76 nordmaniana 35
melanosarx 68 molliformis 25 nothofageti 20
melinocephala 71 mollis 35, 52 nothofagetorum 72
melinocephalus 62 molybdites 19 nothofagi 40, 85
mellea 21 montagnei 15 nothofaginea 92
melleum 38 montana 19, 30, 34, 48, 79, 88, 91 nothofagineus 55
melliolens 20 montanus 5, 17, 44, 56, 97 nothofagorum 39
membranacea 74 monticola 20, 23, 40, 51 nothomyrciae 63, 85
mercedis 78 montivaga 20, 35, 92 nothomyriciae 72
meridae 30 montoyae 97 notodryina 46
meridionalis 34 moseri 16, 38, 85, 95 novaezelandiae 6, 97
mesites 27, 34, 43, 66, 89 mostnyae 72, 75, 96 novembris 79, 96
mesocephalus 62 mucronosporus 78 nubigena 35
mesopora 98 muliercula 87 nubigenum 5, 75
mesospora 22, 87, 92 multicaudata 72 nubigenus 55, 76, 89
mesosporus 16, 54, 62, 83 mundula 15 nuda 50
metachromatica 92 mundulus 63 nudum 74
metuloidigera 31 munyozii 63, 72 nudus 47
metuloidophora 85 muricellata 84 nueschii 95
mexicana 27, 34, 45, 77 musacearum 55, 68 nutans 77
mexicanus 6, 25, 83, 94 mutabilis 5, 44-45, 47
michoacanus 16 myceliosa 74 oaxacana 5 1
microcarpa 79 myceniformis 68 oaxacanus 63
microcarpus 6, 84 mycenoides 35, 39, 44, 89 oblongisporus 55
microcephalus 43 myocephalus 63 obscura 47
microdendron 62 myosura 5, 15, 23 obscuratipes 20, 47
microhaedinus 62 myrceugenellae 78 obtecta 92
microjonia 71 myxacioides 79 occidentale 69
microlepis 94 occidentalis 27, 33, 44, 52, 67, 93
microleuca 72 nana 14, 35, 45 ochraceorobustum 96
microleucoides 72 nancyae 5, 17 ochraceous 39
120 Index to Species

ochropus 63 papyracea 75 pilgerodendri 64
octayensis 45 paraboliciformis 72 pilosiceps 26
odoratus 30 paradoxa 14 pineti 28
oedipus 79 paradoxum 28 pinetorum 44
oleiger 63 paraensis 19, 55, 83 piterbargii 72
oligocalyptrata 35 paraibensis 45 platense 28
oligocystis 83 paraiboensis 18 platensis 44
oligogloea 31 paranaguae 21 platycystis 44, 74
oligophylla 15, 20 pararotula 64 platyhyphes 55
oligopora 31 parksii 39, 67 platyphylla 68
oligoporpa 77 parlatorei 26, 55 platyspermus 64
oligosporus 78 pascuus 78 pleuracanthus 64
olivaceiceps 74 paspali 87 pleurotelloides 64
olivaceoalcalina 72 patagonica 35, 40-41, 50, 72, 85, 97 pleurotiformis 41
olivaceonana 74 patagonicum 84, 96 pleurotus 20, 46, 49, 75
olivaceus 15 patagonicus 18, 51, 68, 89 plocamophora 47
olivascenticeps 62-63 pataguae 29 plumbeus 75
omniumsanctorum 72 patouillardii 16, 27, 92 pluteoides 92
omphalina 55 paucigibba 47 pluvialis 73
omphaliniformis 44 paulensis 49, 64, 88 podocarpi 18, 26-27, 64
omphalioides 88 pauper 94 podocarporum 39
omphaliopsis 96 pauperum 28, 94 poecila 73
omphalophora 72 paurosporus 55 poliocephala 73
omphalophorus 55 paxilloides 14 poliocephalus 68
onoticus 63 payettensis 35 poliomphax 84
oreadeoides 72 pearsoniana 72 polychaetopus 64
oreades 75 peckii 92 polychrous 77
oregonensis 99 peladae 21, 35, 48, 87 polycladoides 64
oreina 35, 92 pentstemonis 96 polycystidiosa 85
orinocensis 55 peralbidus 97 polycystis 25, 64, 83, 92
osornensis 55 perangusta 35 polylepidicola 20
ostreatoroseus 81 perangustispermus 55 polylepidis 26, 64, 69, 74
ovalis 94 perbrevisporus 90 polyphyllus 55
oxydabilis 47, 92 pergelatinosa 41 polytrichorum 35
pericystis 87 polytropa 74
pachycystis 40, 92 perlongispermus 64 polytropoides 94
pachytrichus 19 pernambucensis 55 ponderosum 28
pacifica 68 perniciosa 17 popperianum 74
pacificus 63 perrara 35 populi 94
pallenticeps 63, 66 perrarus 64 poromycenoides 64
pallescens 6, 93 perreductus 64 porphyrocephala 73
pallida 15, 91 persobria 92 porpoarachnoides 90
pallidipes 49 peruviana 87 porrigens 6, 80
pallidissima 74 petiolicola 55 portentosa 75
pallidocinctus 63 peullensis 18, 49, 55, 94 portentosum 84
palmarum 5, 17, 25, 39 pezizoides 6, 88 potamogeton 55, 97
palmigena 90 phaeocyanescens 40 potassiovirens 29, 55
palmivora 72 phaeocystidiata 85 pouzarianus 83
panamensis 45, 63, 86, 94 phaeocystis 64 praeandina 20, 74
pandoanus 63 phaeoxanthus 80 praeandinus 64
pandoensis 55 phalaropus 5, 30 praecox 64
panerythrus 63 phellodendri 95 praegrandispermus 64
pantelaeodes 74 philippinensis 5, 29 praemultifolia 6, 80
pantherina 31 phlegmacoides 35 praetervisa 35, 87
pantholocystis 55 phoebophila 21 prancei 50
pantoleuca 92 phylladophila 21 pratensis 40, 73
pantoleucus 6, 97 phylladophiloides 21 procera 5, 14, 52, 79
pantopolia 72 phylladophilus 24 proteus 45
pantoxanthum 38 phyllogenus 44 proxima 23, 35
panuoides 49 phyllophila 27 proximella 48
panziensis 50 physospora 35 proximus 40
papillata 35 piceae 27 pruinatipes 40
papillatomarginatus 55 piceetorum 92 pruinosipes 90
papillatus 44 pichinchensis 64 psammophila 92, 97
papillifer55 pictiformis 16 psathyrelloides 79
papilligera 72 piguicola 72 pseudoalnicola 73

pseudoamylospora 45 rhodopoda 93 sayanus 50
pseudocamerina 35 rhodotrama 21 scandens 20, 64
pseudocerina 35 riberaltensis 56, 83 schiffneri 93
pseudoconidiophora 41 ribesina 73 schini 27, 60
pseudocorrugatus 64 ricardii 83 schultesii 65
pseudocupressiformis 64-65 rickenii 50 schusteri 26, 38
pseudohiulca 47 rimosellus 83 sclerobasidium 24

pseudolepida 92 rimosoaffinis 83 scleronematis 65
pseudominutus 65 riograndensis 44, 83 scotophysinus 65
pseudoniveus 65 rionegrensis 22, 29, 44, 84 scototephrodes 65
pseudoparaphysatus 55 riparia 35, 85 scrobiculatus 98
pseudoripartites 96 riparius 43, 65 segregabilis 56
pseudosordidum 96 roberti 23 segregis 85
pseudostipitaria 27 robertsonii 65 sejunctum 6, 84
pseudostylotabes 73 rogersii 75 sejunctus 65
psilocybe 17 rondoniensis 97 semiglobata 36
psychotriophilus 65 rosacea 93 semiimbricata 32
pteridophytorum 18, 76, 87 rosaceifolia 73 semilanceata 87
pterigena 31 rosaemariae 87 semililacea 93
pterosporus 94 rosascentifolius 56 semimarginellus 44
puberula 3 1 rosea 29 semipilosa 73
pubescentipes 35 roseipes 30, 93 semirudis 76
pucarensis 35 roseogrisea 31 semiustum 5 1
puellaria 93 roseolus 26 semiustus 56

puiggarii 15 roseomaculatus 39 separatum 69
pulcher 46 roseoscabrum 49 septembris 79
pulcherrima 80 roseus 26 septentrionalis 39, 51, 79, 93
pulchra 5, 14, 35 rostrata 35 septicoides 56
pulverulentus 83 rostratus 47, 78 septotricha 27
pumanquensis 29, 90 rotaliformis 65 sepuchralis 85
purgatorii 49 rotundifoliae 47 sequoiae 36
purpurascens 5, 38 rubellus 16 sergii 83
purpurea 27, 31, 69 ruber 65, 95 sericeus 13
purpurellus 80 rubra 18, 29 serotina 93
pusillus 29, 44 rubriceps 5, 26, 52 serotinus 16, 26
pusio 65 rubricitrinus 16 sessile 5, 28
puttemansii 65 rubrimontana 73 sessiliaffinis 65
pycnophylli 41 rubroflava 45 sessilis 39
pyrenaica 30, 93 rubromarginatus 65 setchellianus 52
pyrrhys 6, 87 rubrotomentosus 83 setulosifolius 65
pyxidata 75 rufomarginatus 65 shideiae 66
pyxidatoides 44 ruforotula 65 shoreae 87
rugososulcatus 83 sibiricus 40, 47, 66
quercetorum 77 ruizlealii 5, 30, 79 sierrae 87
quercinus 55 russularia 89 silvicola 66
quercusoleoidis 93 rustica 27 simocyboides 36
quercuum 45, 49, 83, 94 rutilus 19, 39 sinensis 28
quinteroensis 35 sipamnae 28
quitensis 26, 39 sabalensis 31 sleumeri 22
sabalis 44 smaragdina 93
radicata 23 sacchari 56 smithii 39, 93
radicellicola 35, 65 saccosporus 24 solitaria 22
radicicola 40, 54, 98 sachaensis 28 sororiicolor 93
ramealis 55-56 sahleri 36 sosarum 73
ramorum 56, 66 sainii 51 sotae 73
recedens 41, 44, 79, 88 salicinus 83 spadicea 97
reductoaffinis 94 saltensis 36 spadiceospinulosus 78
reductus 56 sanctaemarthae 56 sparrei 15, 68
religiosa 39 sanctaerosae 75, 97 spathularia 39
reticulatorugosa 23 sanctixaverii 36, 65, 83 spegazzinianus 83
retipes 47 sanguirotalis 65 spegazzinii 41
rheicolor 5, 40 sanmartiniana 5, 76 speireoides 78
rhizophorae 85 sapiicola 83 sphaerodermatoides 66
rhododendri 6, 65, 88 sapindacearum 27 sphaerosporus 26, 56
rhodoleuca 45 saviczii 15 sphagneti 24
rhodophyllus 56 saxegothaeae 73, 80 sphagnophila 93

spiculosus 66 sublateralis 75 terrigenum 17
spinifer 5, 16 subleptoloma 20 terrigenus 78
spinosissimus 5, 14, 66 sublevisporus 26 tessulatus 5, 46
splendidus 16 submarasmioides 40 tetrachroa 22
spodoxutha 20 submarginatus 66 tetrachrous 66
spumosa 79 subminutula 93 tetrasphaerigera 3 1
squamosidisca 6, 90 subminutus 84 tetrasphaerophora 73

squamulosa 73 subniger 97 tetraspora 22-23, 30, 34, 48, 77
squarrosiceps 85 subnigra 76 tetrasporum 46, 93
squarrosipes 87 subobscura 76 thaxteri 67
squarrosoamthystina 47 subolivaceomelleus 56 theobromicola 5, 74
squarrulosa 80 subpallidipes 49 theophili 38
squarrulosum 76 subpallidum 38 thermophila 93, 96
steinbachii 93 subpapillata 36 thujina 36
stenocystis 44, 56, 76 subprona 86 thwaitesii 31
stenophylloides 56 subpurpureum 28 tigrensis 15, 25-26
stenosporus 56 subramealis 56 tilcariensis 86
stephanobasis 83 subrenispermus 24 tilieti 3 1
steppicola 20 subreticulatus 49 tomentosum 69

stercorarius 26 subrhacodium 88 tournefortiae 20
stictospora 23 subroseus 39 transformabilis 45
stipitatum 88 subruforotula 66 trichocephala 73
stramineus 6, 98 subsolitarius 17 trichodermialis 56
stratosus 56 subsquamulosa 20 tricholomatoides 96
streptopus 45 substigmaticus 84 trichorhizus 66
striatella 22, 45 substriatulus 17 tricolor 51,56
striatipes 23 substrictior 90 trinitatis 66
strictipes 87 subtenerrima 73 triplocystis 84

strobilomyces 5 36
subtibiicystis triplotricha 73
strophosa 96 subtorulosa 93 triscopa 36
36
stylifera subtropicalis 41, 56, 74 tristis 86
subalbidum 95 subtruncata 36 tropicalis41, 75, 98
subamyloidea 31 subulifera 20, 73 truncatus 26, 98
subandina 85 subumbratilis 76 truncicola 41
subarborescens 66 subvelata 23, 93 truncorum 66
subarctica 36 subvelutina 74 tsugae 36
subarduennensis 77 subvelutipes 17 tucumana 14, 23, 79, 94
subargentea 47 subxerophytica 23 tucumanense 51
subattenuata 77 sucrense 38 tucumanensis 16, 28, 50, 56, 68, 81
subbadia 36 sudorella 73 tucumanus 44, 84
subbryophilum 50 sulcatus 84 tundrae 36
subbullulifera 36 sutherlandii 81 tungurahuae 20
subcaelestina 75 swanetica 45, 75 turficola73
subcartilagineus 44 syringea 22 turfosa 36
subcellulosus 97 tuxtlae 26
subceracea 36 tabacinus 97 tuxtlaensis 23, 26
subcerina 36 taimbesinhoensis 36 tylostomatoides 95
subconocyboides 85 tanyspermus 66
subcoracinus 56 tapinia 45 uchumachiensis 36
subcortinaria 50 tasmaniensis 66 umbilicatus 24, 56
subfibrillosus 83 tatrensis 48 umbonatus 5, 18
subfibrosoides 47 teapae 31 umbonifer 57
subfulva 67 tener 56 umbratilis 76
subfuscocinnamomea 47 tenera 23, 56 umbrinoalbidus 84
subgeophyllomorpha 47 tenerrima 23 undata 30
subgigacystis 47 tennesseensis 93 universitaria 68
subglabripes 36 tenuiparietalis 66 urosphaera 94
subheteromorpha 45 tenuipilosa 28 urubambae 22
subhirtellus 56 tenuisperma 31 uspallatae 24
subhybrida 22 tenuitunicata 18 ustilago 66
subhygrophana 20 tephrocybe 90 uyedae 49
subhygrophanoides 20 tephromelanus 66
subhyperella 87 tequendamae 47 valdiviae 13
subinamyloidea 73 tereticeps 66 valdiviana 73, 86, 88
subincarnatus 78 terraefirmae 44, 66 valdivianus 66
subinodermatoides 56 terreum 84 valdiviensis 26, 48, 87

varariotecta 3 1 victoriae 36 xanthodictyon 38
variabiliceps 66-67 vigintifolius 67 xanthophaeus 26
variabilispora 67 vinaceobrunnea 37 xanthophylla 50
variegata 69, 93 vinaceogriseus 97 xanthoporphyria 19
variipes 84 vinicolor 39 xanthopus 84
variisporus 26 vinosa 93 xanthosarx 44
varzeae 57, 79, 86 vinosus 57, 61, 63 xanthydrorheus 49
varzeicola 84 violaceum 95 xavierii 68
vasilievae 20, 73, 79 violascens 5, 17, 31 xerampelina 93
velutinellus 24 violeorotalis 67 xerampelinus 67
velutinoaffinis26 viperinus 78 xerophilus 44
velutipes 22, 36
viridiflava 15 xerophytica 14, 23
venezuelae 97 viscidulus 44, 84 xerophyticus 57, 67
venezueliana 93 viscosa 14, 68 xestocephalus 67
venososulcatus 21 vittaeformis 37 xuthophyllus 44
ventalloi 67 volvatus 57 xylodendron 67
ventallonii 67 vulgaris 21 xylophila 23
veraecrucis 25, 44, 49 xylophilus 84
vergeliensis 67 wahgiensis 93
weberi 17 yalae 38, 57, 67
verna 93
wettsteinii 19 yalensis 73
vernalis 47, 93
vernifera 68 williamsii 97 yangambiensis 67
witteana 18 yarizae 67
verrucispora 79
witteanus 67 yungense 29
verruculosa 98
wolfeana 31 yungensis 15, 33, 84, 87, 98
veselskyi 45
yungicola 26, 37
vesiculosa 67
vexans 36 xanthocephala 73 zaragozae 84
viatica 36 xanthocephalus 26 zellerianus 30

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Mycological Contributions of Rolf Singer

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Mycological Contributions of Rolf Singer

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UNIVERSITY OF

Vlycological Contributions of Rolf Singer:

Gregory M. Mueller and Qiuxin Wu, Editors

February 28, 1997

PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY

'. t II. Polyp

IHust e text (not

This paper meets the requirements of ANSI/NISO Z39.48-1992 (Permanence of Paper).

Mycological Contributions of Rolf Singer:

Gregory M. Mueller and Qiuxin Wu, Editors

T>iE LIBRARY OF THE

Accepted December 6, 1995

PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY

Introduction and Acknowledgments v

Gregory M. Mueller and Qiuxin Wu

Index to Species 115

2. Books and bound publications of Rolf Singer 3

3. Number of publications of Rolf Singer by age 3

4. Genera in which Singer and associates described at least 40

5. Geographic distribution of Singer's type specimens 4

6. Herbaria with large holdings of Singer's type specimens 4

1. List of new genera published by Rolf Singer 5

2. Type localities and numbers of Singer's new taxa 6

3. Herbaria with holdings of Singer's type specimens 7

4. Rolf Singer's fieldwork, by country and year 1 1

5. Itinerary of Rolf Singer's fieldwork since 1941 11

Singer, 1906-1994, was one of the most influential figures in the

Gregory M. Mueller Department of Botany, Field Museum of Natural History,

Qiuxin Wu Department of Botany, Field Museum of Natural History, Roosevelt

Betty A. Strack Department of Botany, Field Museum of Natural History, Roose-

Clark L.Ovrebo Department of Botany, University of Central Oklahoma, Ed-

Singer had on our knowledge of fungi. Two in-

teresting works by Martha Singer, Rolf's wife and

Singer's Contributions to Mycology 1

Ecology Ethnomycology and Mushroom

lished 439 scientific papers or books. taxa.

Singer's Contributions to Mycology

Figure 5. Geographic distribution of Singer's type 100

specimens. Type localities for Singer's new taxa occur

The Field Itinerary of Rolf Singer

Argentina 1948-1952, 1955-1966, 1969,

Index to Genera, Species, and Infraspecific Taxa 13

coprophila Singer, 1941e: 99 diemii Singer, 1954b: 120, 121

firma var. tucumana Singer, 1959b: 398 Singer M 562 (LIL)

umana (1959b: 398), except for the herbarium ANTHRACOPHYLLUM

Lara, 9 June 1949, Singer Sill (LIL)

identical to that of Nothoclavulina ditopa AUSTROBOLETUS

1-4 May 1967, Singer M 6574 (SCO)

var. umbrinobrunnea Singer, 1956f:

puiggarii ssp. tigrensis Singer, 1970a: 8 AUSTROGASTER Singer, 1962a: 57

Index to Genera, Species, and Infraspecific Taxa 15

BOLBITIUS chilensis Singer, 1967b: 69

Caracaraf, km 115, 4 April 1978, Singer B hypocarycinus Singer, 1945i: 798

subsolitarius Singer, 19451: 798 luteofuscum Singer, 1953d: 115

Type information: USA. Florida: Alachua Coun-

substriatulus Singer, 1943a: 156 bipigmentata Singer, 1977g: 267

cyanea Singer ex Redhead & Singer, 1978c:

Index to Genera, Species, and Infraspecific Taxa 17

ionym: Calocybe rubra Rick in Singer, 1948. austrochilensis Singer, 1969a: 86

CALYPTELLA HOLOTYPE: Argentina. Catamarca, Santa

floridana Singer, 1950e: 249

CAMPANELLA CANTHARELLULA Singer, 1936e: 281

aberrant Singer, 1975c: 855 Type species: Cantharellus umbonatus Gmelin